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Caenagnathus collinsi

Reptilia - Caenagnathidae

Taxonomy
Caenagnathus collinsi was named by Sternberg (1940). Its type specimen is CMN 9776, a mandible, and it is a 3D body fossil. Its type locality is RTMP Quarry Q112, Sand Creek (111 ft level) [NMC], which is in a Campanian terrestrial horizon in the Dinosaur Park Formation of Canada. It is the type species of Caenagnathus.

It was synonymized subjectively with Chirostenotes pergracilis by Paul (1988), Sues (1997), Osmólska et al. (2004) and Currie (2005).

Synonymy list
YearName and author
1940Caenagnathus collinsi Sternberg p. 81
1956Caenagnathus collinsi Wetmore p. 6
1960Caenagnathus collinsi Wetmore p. 4
1964Caenognathus collinsi Russell p. 13
1970Caenagnathus collinsi Steel p. 20
1970Caenagnathus collinsi Swinton pp. 136-137
1971Caenagnathus collinsi Brodkorb p. 39
1971Caenagnathus collinsi Cracraft p. 805
1978Caenagnathus collinsi Brodkorb p. 223
1981Caenagnathus collinsi Barsbold
1982Caenagnathus collinsi Morris p. 488
1983Caenagnathus collinsi Barsbold p. 94
1986Caenagnathus collinsi Gauthier p. 9
1987Caenagnathus collinsi Currie p. 52
1988Caenagnathus collinsi Currie and Russell p. 984
1992Caenagnathus collinsi Currie p. 239
1994Caenagnathus collinsi Currie et al.
2008Caenagnathus collinsi Longrich p. 994
2009Caenagnathus collinsi Zanno et al. p. S15
2013Caenagnathus collinsi Longrich et al. p. 28
2014Caenagnathus collinsi Lamanna et al. p. 10 fig. 6
2015Caenagnathus collinsi Currie and Koppelhus p. 626
2015Caenagnathus collinsi Funston et al. p. 180
2015Caenagnathus collinsi Sues and Averianov p. 53
2015Caenagnathus collinsi Tsuihiji et al. p. 61
2016Caenagnathus collinsi Funston and Currie p. 1
2019Caenagnathus collinsi Lee et al. p. 2
2020Caenagnathus collinsi Funston p. 113
2020Caenagnathus collinsi Rhodes et al. p. 1

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
RankNameAuthor
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Theropoda()
Neotheropoda
AverostraPaul 2002
Tetanurae
Coelurosauria()
Maniraptora
infraorderOviraptorosauria
familyCaenagnathidae
subfamilyCaenagnathinae
genusCaenagnathus
speciescollinsi

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Caenagnathus collinsi Sternberg 1940
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Diagnosis
ReferenceDiagnosis
N. R. Longrich et al. 2013Caenagnathid characterized by large size (symphyseal length approximately 60 mm, dentary length approximately 160 mm; mandible length greater than 200 mm). Dentary long and low. Dentary narrow in dorsal view, with posterior rami weakly bowed outward, diverging at an angle of approximately 15°. Tip of beak flattened, shovel shaped and projecting anteriorly. Symphysis not downturned. Tip of beak with three symphyseal ridges, including a large median ridge. Symphyseal shelf extended caudally between lingual shelves. Dentary excluded from dorsal margin of external mandibular fenestra by anterior process of surangular. Posterodorsal process of dentary elongate, extending posteriorly below the coronoid eminence of the surangular. Surangular slender and straplike in lateral view. Ventral flange of surangular weakly developed. Lateral cotyle of articular extremely reduced.
G. F. Funston 2020Large caenagnathid oviraptorosaur diagnosed by the following combination of features and autapomorphies (indicated by asterisks): elongate dentary symphysis* (shared with Anzu wyliei); low quadrate articular ridge of mandible; large posterior protuberance on proximal end of metatarsal II*; groove between proximal articular surface and flexor tubercle present in manual ungual II-3 but not I-2*; rounded ventral edge of preacetabular blade; low ilium above the acetabulum; inclined ventral edge of pubic peduncle of ilium; pubic peduncle of ilium with anteroposterior ridge on ventral surface for contact with pubis, forming concavo-convex contact*; pubes relatively straight in anterior view, producing transversely narrow proximal end; iliac contact of pubis with anteroposterior concavity for ridge on the pubic peduncle of the ilium, forming concavo-convex contact*.