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Yangchuanosaurus shangyouensis

Reptilia - Avetheropoda - Metriacanthosauridae

Taxonomy
Yangchuanosaurus shangyouensis was named by Dong et al. (1978). Its type specimen is CV 00215, a skeleton, and it is a 3D body fossil. Its type locality is Shangyou Reservoir, which is in an Oxfordian/Kimmeridgian terrestrial mudstone in the Shangshaximiao Formation of China.

It was recombined as Metriacanthosaurus shangyouensis by Paul (1988).

Synonyms
Synonymy list
YearName and author
1978Yangchuanosaurus shangyouensis Dong et al. pp. 302-304 figs. 1-2
1981Yangchuanosaurus shangyouensis Molnar et al. p. 145
1981Yangchuanosaurus shangyouensis Olshevsky p. 36
1983Yangchuanosaurus shangyouensis Dong et al. p. 68
1983Yangchuanosaurus magnus Dong et al. pp. 83-93 figs. 54-63, Pl. 27-31
1985Yangchuanosaurus shangyouensis Zhao p. 288
1988Metriacanthosaurus shangyouensis Paul p. 290
1990Yangchuanosaurus magnus Molnar et al. p. 192
1990Yangchuanosaurus shangyouensis Molnar et al. p. 192
1991Yangchuanosaurus shangyouensis Molnar p. 165
1992Yangchuanosaurus shangyuensis Bakker et al. p. 9 fig. 3E
1992Yangchuanosaurus magnus Bakker et al. p. 22
1992Yangchuanosaurus magnus Dong p. 85
1992Yangchuanosaurus shangyouensis Dong p. 85
1992Yangchuanosaurus magnus Gao p. 323
1992Yangchuanosaurus shangyouensis Gao p. 323
1994Yangchuanosaurus magnus Currie and Zhao p. 2038
1994Yangchuanosaurus shangyuensis Currie and Zhao p. 2038
1997Yangchuanosaurus magnus Dong p. 122
1997Yangchuanosaurus shangyouensis Dong p. 122
1999Yangchuanosaurus magnus Gao
1999Yangchuanosaurus shangyouensis Gao
2003Yangchuanosaurus magnus Rauhut p. 27
2003Yangchuanosaurus shangyouensis Rauhut p. 27
2004Yangchuanosaurus magnus Holtz, Jr. et al. p. 74
2004Yangchuanosaurus shangyouensis Holtz, Jr. et al. p. 74
2007Yangchuanosaurus shangyouensis Sampson and Witmer p. 38
2008Yangchuanosaurus shangyouensis Watabe et al. p. 33
2008Yangchuanosaurus magnus Watabe et al. p. 35
2009Yangchuanosaurus shangyuensis Wu et al. p. 13
2009Yangchuanosaurus magnus Wu et al. p. 14
2009Yangchuanosaurus magnus Xing et al. p. 1394
2009Yangchuanosaurus shangyouensis Xing et al. p. 1394
2012Yangchuanosaurus shangyouensis Carrano et al. p. 246

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
OlfactoresJefferies 1991
subphylumVertebrata
Gnathostomata()
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
RankNameAuthor
Eosuchia()
Neodiapsida
SauriaGauthier 1984
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Saurischia()
Theropoda()
Neotheropoda
AverostraPaul 2002
Tetanurae
orderAvetheropoda
superfamilyAllosauroidea
familyMetriacanthosauridae
genusYangchuanosaurus
speciesshangyouensis

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
Z. Dong et al. 1983 (Yangchuanosaurus magnus)An extremely large megalosaur with a massive skull that displays two maxillary depressions, one of which penetrates the maxilla. Four of the five sacral spines are fused with the anterior sacral centrum more robust than the other centra.
Z. Dong et al. 1983Body length is approximately seven meters, cranial height and length are of 1:1.6, maxilla is deeply concave and not perforated, five sacral centra are fused but lack any additional buttressing, ilium is low with a preacetabular process that is broad, flatly expanded anteriorly, and lacks ventral curvature. Ventral keels on the cervical vertebrae are absent.
M. T. Carrano et al. 2012The original diagnoses of Yangchuanosaurus and its two constituent species (Dong et al. 1978, 1983) include features that now characterize ‘sinraptorids’, allosauroids and tetanurans more generally. More recent work (Currie & Zhao 1994) has distinguished Yangchuanosaurus from Sinraptor primarily on the presence of sinraptorid characters together with the absence of Sinraptor autapomorphies (i.e. a unique character combination but no autapomorphies). Yangchuanosaurus has a slightly higher ratio of skull height to length (0.5) than Sinraptor (0.4) and correspondingly has a proportionally taller maxilla. The dorsal vertebral neural spines are also lower (about 1.8 times centrum height compared to 2.0 in S. dongi) and the centra are relatively longer. Yangchuanosaurus may also exhibit a more pronounced margin of the antorbital fossa on the jugal, although this is difficult to ascertain in some sinraptorid specimens. We concur that Yangchuanosaurus can be distinguished from other sinraptorids, but in the absence of a detailed restudy of the type materials we can offer no new autapomorphies to define this genus.