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Gopherus (gopher tortoise)
Taxonomy
Gopherus was named by Rafinesque (1832). It is extant. Its type is Testudo polyphemus. It is the type genus of Gopherinae, Gopherinii.
It was assigned to Gopherinii by Swingland and Klemens (1989); to Testudines by Anquetin et al. (2009); and to Testudinidae by Hay (1930), Galbreath (1953), Auffenberg (1971), Carroll (1988), Hutchison (1996), Frank and Ramus (1996), Wall and Maddox (1998), Reynoso and Montellano-Ballesteros (2004), Joyce et al. (2004), de la Fuente et al. (2014), van Dijk et al. (2014), Vlachos and Rabi (2018), Vlachos (2018) and May (2019).
It was assigned to Gopherinii by Swingland and Klemens (1989); to Testudines by Anquetin et al. (2009); and to Testudinidae by Hay (1930), Galbreath (1953), Auffenberg (1971), Carroll (1988), Hutchison (1996), Frank and Ramus (1996), Wall and Maddox (1998), Reynoso and Montellano-Ballesteros (2004), Joyce et al. (2004), de la Fuente et al. (2014), van Dijk et al. (2014), Vlachos and Rabi (2018), Vlachos (2018) and May (2019).
Species
G. (Gopherus), G. (Oligopherus), G. agassizii, G. auffenbergi (syn. G. pargensis), G. berlandieri, G. brevisternus, G. depressus (syn. G. dehiscus), G. donlaloi, G. edae (syn. Testudo pansa, Testudo vaga, Testudo hollandi), G. flavomarginatus, G. hexagonatus (syn. G. huecoensis, Testudo campester, Bysmachelys canyonensis), G. laticuneus, G. mohavensis, G. morafkai, G. polyphemus, Testudo milleri
Synonyms
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Synonymy list
Year | Name and author |
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1832 | Gopherus Rafinesque |
1857 | Xerobates Agassiz |
1930 | Gopherus Hay p. 101 |
1937 | Bysmachelys Johnston p. 447 |
1953 | Gopherus Galbreath p. 38 |
1961 | Testudo milleri Brattstrom p. 546 |
1971 | Gopherus Auffenberg p. 116 |
1982 | Scaptochelys Bramble p. 853 figs. 1-11 |
1988 | Gopherus Carroll |
1989 | Gopherus Swingland and Klemens p. 124 |
1992 | Xerobates Hutchison |
1996 | Gopherus Frank and Ramus |
1996 | Gopherus Hutchison p. 348 |
1996 | Gopherus (Oligopherus) Hutchison p. 348 |
1998 | Gopherus Wall and Maddox p. 11 |
2004 | Gopherus Joyce et al. |
2004 | Gopherus Reynoso and Montellano-Ballesteros p. 823 |
2009 | Gopherus Anquetin et al. |
2014 | Gopherus de la Fuente et al. |
2014 | Gopherus van Dijk et al. |
2018 | Gopherus Vlachos |
2018 | Oligopherus Vlachos |
2018 | Gopherus Vlachos and Rabi p. 663 |
2019 | Gopherus May |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. Gopherus Rafinesque 1832 [gopher tortoise]
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Subg. †Gopherus (Gopherus) Rafinesque 1832
Gopherus agassizii Cooper 1863 [Mojave desert tortoise]
Gopherus berlandieri Agassiz 1857
†Gopherus brevisternus Loomis 1909
†Gopherus depressus Brattstrom 1961
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Invalid names: Gopherus dehiscus Des Lauriers 1965 [synonym]
†Gopherus donlaloi Reynoso and Montellano-Ballesteros 2004
†Gopherus edae Hay 1906
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Invalid names: Testudo hollandi Hay 1906 [synonym], Testudo pansa Hay 1908 [synonym], Testudo vaga Hay 1908 [synonym]
Gopherus flavomarginatus Legler 1959
†Gopherus hexagonatus Cope 1893
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Invalid names: Bysmachelys canyonensis Johnston 1937 [synonym], Gopherus huecoensis Strain 1966 [synonym], Testudo campester Hay 1908 [synonym]
†Gopherus laticuneus Cope 1873
†Gopherus mohavensis Merriam 1919
Gopherus morafkai Murphy et al. 2011 [Sonoran desert tortoise]
Gopherus polyphemus Daudin 1802 [Florida gopher tortoise]
Invalid names: Bysmachelys Johnston 1937 [synonym], Gopherus (Oligopherus) Hutchison 1996 [invalid subgroup], Scaptochelys Bramble 1982 [synonym], Testudo milleri Brattstrom 1961 [nomen dubium], Xerobates Agassiz 1857 [synonym]
Diagnosis
No diagnoses are available