PBDB Taxon

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Cycadales

Cycadopsida - Cycadales

Taxonomy

Cycadales was named by Dumortier (1829). It is extant.

It was assigned to Gymnospermae by Knowlton (1919); to Gymnospermae by MacGinitie (1941); to Cycadopsida by Meyen (1984) and Doweld (2001); to Cycadophyta by Taylor et al. (2008); and to Cycadidae by Christenhusz et al. (2011).

Synonymy list

Year	Name and author
1829	Cycadales Dumortier p. 65
1919	Cycadales Knowlton
1941	Cycadales MacGinitie
1984	Cycadales Meyen p. 89
2001	Cycadales Doweld p. iii
2008	Cycadales Taylor et al. p. 1028
2011	Cycadales Christenhusz et al.

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Rank	Name	Author
kingdom	Plantae
subkingdom	Embryophyta
phylum	Cycadophyta	Bessey 1907

Rank	Name	Author
class	Cycadopsida	Brongniart 1843
order	Cycadales	Dumortier 1829
order	Cycadales	Dumortier 1829

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Cycadales Dumortier 1829

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G. †Amuriella Prinada 1956

†Amuriella extra Prinada 1956

G. †Anthrophyopsis Nathorst 1878

†Anthrophyopsis crassinervis Nathorst 1878

†Anthrophyopsis decurrens Kimura and Kim 1989

†Anthrophyopsis miassica Vladimirovich 1958

†Anthrophyopsis naruensis Doludenko and Svanidze 1970

†Anthrophyopsis nilssoni Nathorst 1878

G. †Apoldia Wesley 1958

†Apoldia tenera Compter 1883

G. †Archaeocycas Mamay 1973

†Archaeocycas whitei Mamay 1973

G. †Aricycas Ash 1991

†Aricycas paulae Ash 1991

Fm. †Beaniaceae

G. †Androstrobus Schimper 1870

†Androstrobus major Van Konijnenburg-Van Cittert 1968

†Androstrobus manis Harris 1941

†Androstrobus munku Archangelsky and Villar de Seoane 2004

†Androstrobus patagonicus Archangelsky and Villar de Seoane 2004

†Androstrobus rayen Archangelsky and Villar de Seoane 2004

†Androstrobus wonnacotti Harris 1941

†Androstrobus zamioides Schimper 1872

G. †Beania Carruthers 1869

†Beania carruthersi Nathorst 1902

†Beania gracilis Carruthers 1869

†Beania kochi Harris 1937

G. †Behuninia Chandler 1966

†Behuninia joannei Chandler 1966

G. †Bucklandia Sternberg 1825

†Bucklandia anomala Sternberg 1825

Invalid names: Clathraria anomala Stokes and Webb 1824 [objective synonym]

†Bucklandia dichotoma Sharma 1969

†Bucklandia guptae Sharma 1967

†Bucklandia sahnii Bose 1953

G. †Bureja Prinada 1956

†Bureja rigida Prinada 1956

G. †Cavamonocolpites de Lima 1989

†Cavamonocolpites punctatus de Lima 1989

G. †Ctenozamites Nathorst 1886

†Ctenozamites cycadea Brongniart 1886

†Ctenozamites kachchhensis Bose and Banerji 1984

†Ctenozamites surangei Sukh-Dev and Zeba-Bano 1981

Fm. Cycadaceae Persoon 1807

G. †Beaniopsis Ganju 1947

†Beaniopsis rajmahalensis Ganju 1944

G. †Ctenophyllum Schimper 1870

†Ctenophyllum angustifolium Fontaine 1896

†Ctenophyllum braunianum Göppert 1844

Invalid names: Pterophyllum decussatum Emmons 1856 [synonym], Pterophyllum spatulatum Emmons 1857 [synonym], Pterozamites abbreviatus Braun 1843 [synonym], Pterozamites obtusifolius Rogers 1843 [synonym]

†Ctenophyllum densifolium Fontaine 1896

†Ctenophyllum giganteum Fontaine 1883

†Ctenophyllum grandifolium Fontaine 1883

†Ctenophyllum latifolium Fontaine 1889

†Ctenophyllum lineare Emmons 1857

†Ctenophyllum pachynerve Fontaine 1905

†Ctenophyllum robustum Emmons 1857

Invalid names: Ctenophyllum emmonsi Fontaine 1883 [synonym]

†Ctenophyllum taxinum Lindley and Hutton 1835

†Ctenophyllum truncatum Fontaine 1883

†Ctenophyllum wannerianum Fontaine 1900

†Ctenophyllum wardii Fontaine 1896

G. †Ctenopsis Berry 1911

†Ctenopsis latifolia Fontaine 1889

G. †Cycadinocarpus Schimper 1870

†Cycadinocarpus chapini Newberry 1888

†Cycadinocarpus circularis Newberry 1895

†Cycadinocarpus keuperianus Schenk 1870

†Cycadinocarpus rajamahalensis Feistmantel 1877

G. †Cycadites Buckland 1836

†Cycadites acutus Emmons 1856

†Cycadites cutchensis Feistmantel 1876

†Cycadites megalophyllus Buckland 1836

†Cycadites pungens Lesquereux 1891

†Cycadites rajmahalensis Oldham 1863

Invalid names: Cycadites blanfordianus Oldham 1863 [synonym], Cycadites conferta Morris 1863 [synonym]

†Cycadites tenuinervis Fontaine 1883

†Cycadites unjiga Dawson 1882

†Cycadites wadianus Bose et al. 1983

G. †Cycadospadix Schimper 1870

†Cycadospadix hennocquei Pomel 1870

G. Cycas Linnaeus 1753

†Cycas fujiiana Yokoyama 1911

G. †Dichotozamites Berry 1911

†Dichotozamites cycadopsis Fontaine 1889

G. †Dioonites Miquel 1851

†Dioonites borealis Dawson 1882

†Dioonites buchianus Ettingshausen 1852

Invalid names: Pterophyllum abietinum Göppert 1816 [synonym]

†Dioonites carnallianus Göppert 1843

†Dioonites columbianus Dawson 1873

†Dioonites dunkerianus Göppert 1843

†Dioonites feneonis Brongniart 1828

G. †Encephalartopsis Fontaine 1889

†Encephalartopsis nervosa Fontaine 1889

†Encephalartopsis oregonensis Fontaine 1905

G. Encephalartos Lehmann 1834 [bread tree]

†Encephalartos cretaceus Lesquereux 1891

G. †Glossozamites Schimper 1870

†Glossozamites distans Fontaine 1889

†Glossozamites fontaineanus Ward 1899

†Glossozamites oblongifolius Kurr 1870

G. †Leprospermum Heer 1877

†Leprospermum thurmanni Heer 1877

G. †Leptocycas Delevoryas and Hope 1971

†Leptocycas gracilis Delevoryas and Hope 1971

G. †Menucoa Petriella 1969

†Menucoa cazaui Petriella 1969

G. †Microzamia Corda 1846

†Microzamia dubia Berry 1905

†Microzamia gibba Reuss 1844

G. †Platypterygium Feistmantel 1886

†Platypterygium densinerve Fontaine 1889

†Platypterygium rogersianum Fontaine 1889

G. †Sphenozamites Brongniart 1849

†Sphenozamites beani Lindley and Hutton 1832

†Sphenozamites oblanceolatus Penhallow 1902

†Sphenozamites rogersianus Fontaine 1883

Invalid names: Calamites punctatus Emmons 1857 [synonym]

G. †Zamiopsis Fontaine 1889

†Zamiopsis dentata Fontaine 1889

Invalid names: Zamiopsis longipennis Fontaine 1889 [synonym], Zamiopsis pinnatifida Fontaine 1889 [synonym]

†Zamiopsis laciniata Fontaine 1889

†Zamiopsis petiolata Fontaine 1889

G. †Zamiostrobus Endlicher 1836

†Zamiostrobus macrocephalus Endlicher 1836

†Zamiostrobus virginiensis Fontaine 1883

G. †Cycadenia Bock 1969

†Cycadenia elongata Bock 1969

G. †Cycadinorachis Sharma 1969

†Cycadinorachis omegoides Sharma 1969

Fm. †Dirhopalostachyaceae

G. †Dirhopalostachys Krassilov 1975

G. †Eucommiidites Erdtman 1948

†Eucommiidites minor Groot and Penny 1961

†Eucommiidites stuartii Kemp 1970

†Eucommiidites troedsonii Potonié 1958

†Eucommiidites troedssonii Potonié 1958

G. †Fascisvarioxylon Jain 1964

†Fascisvarioxylon mehtae Jain 1964

†Fascisvarioxylon sahnii Gupta 1971

Invalid names: Sewardioxylon Gupta 1971 [synonym]

G. †Gymnovulites Shukla 1948

†Gymnovulites deccanii Paradkar 1976

G. †Haitingeria Krasser 1916

†Haitingeria krasseri Schuster 1911

G. †Heilungia Prinada 1956

†Heilungia amureansis Novopokrovsky 1956

G. †Mesosingeria Archangelsky 1963

†Mesosingeria coriacea Archangelsky 1963

†Mesosingeria oblonga Villar de Seoane 2005

G. †Michelilloa Archangelsky and Brett 1963

†Michelilloa waltoni Archangelsky and Brett 1963

G. †Nikania Prinada 1956

†Nikania pectinata Prinada 1956

G. †Nilsonia Brongniart 1825

†Nilsonia californica Fontaine 1894

†Nilsonia compta Phillips 1905

†Nilsonia gibbsii Newberry 1863

Invalid names: Nilsonia johnstrupi Heer 1882 [synonym]

†Nilsonia lata Dawson 1882

†Nilsonia nigracollensis Wieland 1905

†Nilsonia nipponensis Yokoyama 1889

†Nilsonia oregonensis Fontaine 1905

†Nilsonia orientalis Heer 1878

†Nilsonia parvula Heer 1876

†Nilsonia pasaytensis Penhallow 1907

†Nilsonia polymorpha Schenk 1867

†Nilsonia pterophylloides Nathorst 1878

†Nilsonia sambucensis Ward 1905

†Nilsonia schaumbergensis Dunker 1889

†Nilsonia stantoni Ward 1905

†Nilssonia brevis Brongniart 1825

†Nilssonia fissa Feistmantel 1879

†Nilssonia princeps Oldham and Morris 1862

Fm. †Nilssoniaceae Kimura and Sekido 1975

G. †Nilssoniocladus Kimura and Sekido 1975

†Nilssoniocladus yukonensis Peppe et al. 2007

Invalid names: Phylloteuthis subovata Meek and Hayden 1860 [synonym]

Invalid names: Phylloteuthis Meek and Hayden 1860 [synonym]

G. †Pseudoctenis Seward 1911

†Pseudoctenis azcaratei Herbst and Troncoso 2000

†Pseudoctenis ballii Feistmantel 1886

†Pseudoctenis eathiensis Richards 1911

†Pseudoctenis fissa Du Toit 1927

†Pseudoctenis fragilis Bose and Banerji 1984

†Pseudoctenis grossa Holmes 1982

G. †Palaeozamia Endlicher 1836

G. †Paracycas Harris 1964

†Paracycas cteis Harris 1964

G. †Phasmatocycas Mamay 1973

†Phasmatocycas kansana Mamay 1973

G. †Pleiotrichium Weyland and Greifeld 1953

†Pleiotrichium commelinoides Weyland and Greifeld 1953

G. †Sarmatiella Prynada 1956

†Sarmatiella brevifolia Prynada 1956

G. †Stangerites Oldham and Morris 1863

G. †Sueria Menéndez 1965

†Sueria rectinervis Menéndez 1965

G. †Taeniopteris Brongniart 1828

†Taeniopteris acuminata McClelland 1850

†Taeniopteris auriculata Fontaine 1889

†Taeniopteris cheondaeriensis Kimura and Kim 1989

†Taeniopteris crassinervis Feistmantel 1877

Invalid names: Macrotaeniopteris misrai Agarwal 1972 [synonym], Taeniopteris buskoghatensis Bose and Banerji 1981 [synonym]

†Taeniopteris densinervis Feistmantel 1876

†Taeniopteris ensis Oldham 1863

†Taeniopteris feddeni Feistmantel 1881

†Taeniopteris glandulata Srivastava 1971

†Taeniopteris haburensis Bose and Banerji 1981

†Taeniopteris kashmirensis Hazra and Prasad 1957

†Taeniopteris kelberi Kustatscher and Van Konijnenburg-van Cittert 2010

†Taeniopteris kutchensis Bose and Banerji 1981

†Taeniopteris lanceolata Oishi 1932

†Taeniopteris major Lindley and Hutton 1833

†Taeniopteris minensis Oishi 1932

†Taeniopteris mungyeongensis Kimura and Kim 1989

†Taeniopteris nabaensis Oishi 1932

†Taeniopteris nervosa Fontaine 1889

Invalid names: Angiopteridium densinerve Fontaine 1889 [synonym], Angiopteridium pachyphyllum Fontaine 1889 [synonym], Angiopteridium strictinerve Fontaine 1889 [synonym], Anomozamites angustifolius Fontaine 1889 [synonym], Anomozamites virginicus Fontaine 1889 [synonym]

†Taeniopteris oldhamii Bose and Banerji 1981

†Taeniopteris oregonensis Fontaine 1905

†Taeniopteris orovillensis Fontaine 1896

†Taeniopteris plumosa Dawson 1882

†Taeniopteris shastensis Knowlton 1919

†Taeniopteris shitakensis Oishi 1932

†Taeniopteris spatulata McClelland 1850

†Taeniopteris stankevichii Sixtel 1962

†Taeniopteris stenoneura Schenk 1867

†Taeniopteris stenophylla Kryshtofovich 1910

†Taeniopteris vittata Brongniart 1831

†Taeniopteris yorkensis Fontaine 1900