PBDB Taxon

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Prohegetotherium schiaffinoi

Mammalia - Notoungulata

Taxonomy

Propachyrucos schiaffinoi was named by Kraglievich (1932). Its type specimen is MNHN-DP-186, a maxilla (partial left maxilla with P2-M2), and it is a 3D body fossil. Its type locality is Santa Lucía Basin, which is in a Deseadan fluvial sandstone in the Fray Bentos Formation of Uruguay.

It was recombined as Prohegetotherium schiaffinoi by Reguero and Cerdeño (2005), Seoane et al. (2017) and Seoane et al. (2019).

Synonyms

Prohegetotherium carettei was named by Minoprio (1947). Its type specimen is MACN 16609, a skull (skull and jaws), and it is a 3D body fossil. Its type locality is Divisadero Largo (Mariño), which is in a Santacrucian terrestrial sandstone in the Mariño Formation of Argentina.

It was recombined as Ethegotherium carettei by Simpson et al. (1962) and López (2002); it was synonymized subjectively with Prohegetotherium sculptum by Reguero (1999); it was synonymized subjectively with Prohegetotherium schiaffinoi by Reguero and Cerdeño (2005).

Synonymy list

Year	Name and author
1932	Propachyrucos schiaffinoi Kraglievich
1947	Prohegetotherium carettei Minoprio pp. 371-374 figs. 3-5
1949	Prohegetotherium carettei Simpson and Minoprio
1962	Ethegotherium carettei Simpson et al.
2002	Ethegotherium carettei López pp. 298-303 figs. 1-3
2005	Prohegetotherium schiaffinoi Reguero and Cerdeño
2017	Prohegetotherium schiaffinoi Seoane et al.
2019	Prohegetotherium schiaffinoi Seoane et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha

Rank	Name	Author
—	Mammaliaformes
class	Mammalia
—	Theriamorpha	(Rowe 1993)
—	Theriiformes	()
—	Trechnotheria
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Afrotheria	Stanhope et al. 1998
—	Paenungulatomorpha
—	Paenungulata	Simpson 1945
—	Sudamericungulata
order	Notoungulata
suborder	Typotheria	Zittel 1892
—	Hegetotheriidae	(Ameghino 1894)
—	Hegetotheriinae	(Ameghino 1894)
genus	Prohegetotherium
species	schiaffinoi	(Kraglievich 1932)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Prohegetotherium schiaffinoi Kraglievich 1932

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Invalid names: Prohegetotherium carettei Minoprio 1947 [synonym]

Diagnosis

Reference	Diagnosis
G.M. López 2002 (Ethegotherium carettei)	Hegetoterine similar to Prohegetotherium, but 20% smaller. Upper and lower incisors and canines directed mesially, originating with 10s P1/1, which are oriented backwards, a false diastema. Upper canine with arched crown; upper jugals poorly imbricated, without development of parastylar areas and with discontinuous enamel, with a characteristic distribution on each tooth; P1/1 slightly reduced and similar in size to 10s P2/2. Unlike Prohegetotherium, P1 lacks labial groove and parastyle, P2 has a deep lingual groove and M3 is very large. i1-2 enlarged and procumbent; lower jugals not imbricate; p2 with a median lingual groove; p4 much smaller than ml. Trigonid of lower molars rounded in outline and not cordate as in Prohegetotherium. The m3 does not hint at the formation of a third lobe as in Prohegetotherium and has a band of dentin in a postlingual position.
M. A. Reguero and E. Cerdeño 2005	Smaller than Prohegetotherium sculptum and S. altiplanense, with shorter and wider nasals. I3, c, and p1 more reduced than in P. sculptum. Lower cheek teeth not im- bricated. Lobes of lower cheek teeth rounded rather than angu- lar, as in P. sculptum and Hegetotherium. Anterior lobe of p3 relatively more elongated than in S. altiplanense. Talonid of m3 without labial groove and with lingual groove fainter than in P. sculptum.