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Colepiocephale lambei

Reptilia - Pachycephalosauridae

Taxonomy
Stegoceras lambei was named by Sternberg (1945). Its type specimen is NMC 8818, a skull, and it is not a trace fossil. Its type locality is Bow Island Ferry (NMC), which is in a Campanian terrestrial horizon in the Foremost Formation of Canada. It is the type species of Colepiocephale.

It was synonymized subjectively with Stegoceras validum by Sues and Galton (1987), Maryanska (1990), Maryanska et al. (2004) and Currie (2005); it was recombined as Colepiocephale lambei by Sullivan (2003), Sullivan (2006), Schott et al. (2009), Longrich et al. (2010), Longrich (2013), Mallon et al. (2015) and Goodwin and Evans (2016).

Synonymy list
YearName and author
1945Stegoceras lambei Sternberg p. 537
1964Stegoceras lambei Kuhn p. 32
1964Stegoceras lambei Russell p. 13
2003Colepiocephale lambei Sullivan p. 189 fig. 3
2003Stegoceras lambei Williamson and Carr fig. 11
2006Colepiocephale lambei Sullivan p. 352
2009Colepiocephale lambei Schott et al. pp. 772-774
2010Stegoceras lambei Lehman p. 788
2010Colepiocephale lambei Longrich et al. p. 280
2013Colepiocephale lambei Longrich p. 52
2015Colepiocephale lambei Mallon et al. p. 429
2016Colepiocephale lambei Goodwin and Evans p. 2

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
RankNameAuthor
Eosuchia()
Neodiapsida
SauriaGauthier 1984
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Ornithischia()
Neornithischia()
Clypeodonta
Cerapoda
Marginocephalia
Pachycephalosauria()
familyPachycephalosauridae
genusColepiocephale
specieslambei()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
R. M. Sullivan 2003Nearly fully-domed pachycephalosaur; differs from all other pachycephalosaurs in: having no lateral (posterior supraorbital and postorbital) and no posterior squamosal shelf; squamosals restricted to lateral sides and much reduced, with only a hint of their margins lying ventrally under the parietal; parietal strongly down-turned, forming a steeply inclined border across entire posterior breadth of skull; occiput reduced; frontoparietal mass thick and broad posteriorly, medial frontal lobe high and relatively narrow; lateral frontal lobes recessed posteriorly; frontoparietal dome with pronounced constriction at contact of postorbital and posterior supraorbital (viewed dorsally); peripheal elements (postorbital, posterior supraorbital, anterior supraorbital and prefrontal) reduced and situated low along the frontoparietal; differs further from Stegoceras validum in lacking supratemporal fossae; differs further from Prenocephale edmontonense in having incipient anterior supraorbitals, which are not incorporated into the anterolateral (frontal) part of the dome.imens of this taxon and was unable to refer them to this species.
R. K. Schott et al. 2009Pachycephalosaurid dinosaur characterized by the following autapomorphies: a frontoparietal dome that reaches its greatest width near posterior margin of sutural surface for postorbital; inflated parietal region of dome overhangs anterior margin of supratemporal fenestrae; sutural surface for postorbital highly arched such that there is a diastema between sutural surfaces for postorbital and posterior supraorbital; diastema accompanied by an indentation in lateral wall of dome; narrow and high frontonasal boss flanked by supraorbital lobes positioned at posterior end of boss; supraorbital lobes highly distinct and columnar. Differs from Stegoceras validum and Hanssuesia sternbergi in the prominence of the supraorbital lobes and the posterodorsal inflation of the parietal. Differs from Prenocephale, Tylocephale, and Pachycephalosaurus in having open supratemporal fenestrae and a more prominent frontonasal boss.