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Crossochelys corniger

Reptilia - Meiolaniidae

Taxonomy
Niolamia argentina was named by Ameghino (1899). Its type specimen is MLP 26-40, a skull (almost complete skull and lower jaw), and it is a 3D body fossil. Its type locality is Cañadón Blanco (Niolamia locality), which is in an Eocene terrestrial tuff in Argentina. It is the type species of Niolamia.

Synonymy list
YearName and author
1899Niolamia argentina Ameghino
1901Niolamia argentina Woodward p. 176
1937Niolamia argentina Simpson
1938Niolamia argentina Simpson
2011Niolamia argentina Sterli and de la Fuente
2013Niolamia argentina Pérez-García and Murelaga
2014Niolamia argentina de la Fuente et al.
2015Niolamia argentina Sterli

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
OlfactoresJefferies 1991
subphylumVertebrata
Gnathostomata()
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
RankNameAuthor
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
Mesochelydia
Perichelydia
Meiolaniformes
familyMeiolaniidaeLydekker 1887
genusCrossochelys
speciescorniger

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
J. Sterli 2015Niolamia argentina is differentiated from other meiolaniids by the significantly large cranial scute A, contact of scute A with scute X, separation of the raised D scutes by a large, rectangular scute X, scute X does not separate G scutes in adults, the absence of scute H, contact between scutes D and F, straight, posterolaterally projecting scute B, presence of a broad shelf at the back of the skull formed by scutes A–C, protruding nasal bones, absence of pterygoid covering the foramen caroticum basisphenoidale, participation of the quadrate in the foramen stapedio-temporale, presence of a triturating surface with only two ridges (labial and lingual ridges), an undivided apertura narium externa, absence of a nasomaxillary sinus, presence of spines on the posterolateral border of the carapace, the presence of supracaudal scute and a broad pygal embayment and the presence of ventrally closed tail rings.