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Diplopoda (millipede)

Myriapoda

Taxonomy
Diplopoda was named by Gervais (1844) [also attributed to de Blainville in Gervais 1844]. It is extant.

It was reranked as the order Diplopoda by Scudder (1890); it was reranked as the class Diplopoda by Hoffman (1969), Wilson and Anderson (2004) and Riquelme et al. (2014).

It was assigned to Myriapoda by Scudder (1890) and Dzik (1975); to Dignatha by Grimaldi and Engel (2005); and to Myriapoda by Hoffman (1969), Wilson and Anderson (2004), Štamberg and Zajíc (2008) and Riquelme et al. (2014).

Sister taxa
Arthropleuridea, Chilognatha, Penicillata

Synonymy list
YearName and author
1844Diplopoda Gervais
1890Diplopoda Scudder p. 43
1969Diplopoda Hoffman p. 578
1975Diplopoda Dzik p. 17
2004Diplopoda Wilson and Anderson p. 171
2005Diplopoda Grimaldi and Engel
2008Diplopoda Štamberg and Zajíc p. 81
2014Diplopoda Riquelme et al. p. 5

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
RankNameAuthor
phylumArthropodaLatreille 1829
classMyriapoda()
DignathaTiegs 1947
subclassDiplopodaGervais 1844
subclassDiplopodaGervais 1844

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subcl. Diplopoda Gervais 1844 [millipede]
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Subcl. †Arthropleuridea Waterlot 1934
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Or. †Arthropleurida Waterlot 1934
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Fm. †Arthropleuridae Scudder 1885
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G. †Arthropleura Jordan 1854
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Arthropleura armata Jordan 1854
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Invalid names: Arthropleura affinis Goldenberg 1873 [synonym], Arthropleura zeilleri Boule 1893 [synonym], Halonia irregularis Geinitz 1855 [synonym]
Arthropleura britannica Andree 1913
Arthropleura cristata Richardson 1959
Arthropleura enodis Guthörl 1934
Arthropleura fayoli Boule 1893
Arthropleura maillieuxi Pruvost 1930
Arthropleura mammata Salter 1863
Arthropleura moyseyi Calman 1914
Arthropleura punctata Goldenberg 1873
Invalid names: Carcinochelus Goldenberg 1877 [synonym], Carcinochelus anthracophilus Goldenberg 1877 [nomen dubium], Halonia Geinitz 1855 [synonym]
G. †Arthropleurion Goldenberg 1877
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Arthropleurion inermis Goldenberg 1877
G. †Branchiopus Dumeril 1806
G. †Branchipusites Goldenberg 1873
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Branchipusites anthracinus Goldenberg 1873
G. †Leptozoa Goldenberg 1877
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Leptozoa rugosa Goldenberg 1877
G. †Propater Troschel 1863
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Propater astacorum Troschel 1863
Invalid names: Tiphoscorpionidae Kjellesvig-Waering 1986 [synonym], Troxites Goldenberg 1854 [nomen dubium], Troxites germari Goldenberg 1854 [nomen dubium]
Subcl. Chilognatha Latreille 1802
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Infracl. Helminthomorpha Pocock 1887
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G. †Archicambala Cook 1895
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Archicambala dawsoni Scudder 1868
Superor. †Archipolypoda Scudder 1882
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G. †Albadesmus Wilson and Anderson 2004
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Albadesmus almondi Wilson and Anderson 2004
G. †Anaxeodesmus Wilson 2005
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Anaxeodesmus diambonotus Wilson 2005
G. †Anthracodesmus Peach 1899
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Or. †Archidesmida Wilson and Anderson 2004
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Fm. †Archidesmidae Scudder 1885
Fm. †Zanclodesmidae Wilson et al. 2005
Or. †Cowiedesmida Wilson and Anderson 2004
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Fm. †Cowiedesmidae Wilson and Anderson 2004
Or. †Euphoberiida Hoffman 1969
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Fm. †Euphoberiidae Scudder 1882
G. †Kampecaris Page 1856
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Kampecaris dinmorensis Clarke 1951
Kampecaris forfarensis Peach 1882
G. †Lauravolsella Haug et al. 2023
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Lauravolsella willemeni Haug et al. 2023
G. †Palaeodesmus Wilson and Anderson 2004
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Palaeodesmus tuberculata Brade-Birks 1923
Or. †Palaeosomatida Hannibal and Krzemiński 2005
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Fm. †Palaeosomatidae Hannibal and Krzemiński 2005
Or. †Pleurojulida Wilson and Hannibal 2005
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Fm. †Pleurojulidae Schneider and Werneburg 1998
G. †Pneumodesmus Wilson and Anderson 2004
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Pneumodesmus newmani Wilson and Anderson 2004
Fm. †Woodesmidae Ross et al. 2018
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G. †Woodesmus Ross et al. 2018
Invalid names: Euphoberidae Scudder 1882 [empty]
G. †Archiulus Scudder 1868
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Archiulus brassi Dohrn 1868
Archiulus euphoberioides Scudder 1895
Archiulus glomeratus Scudder 1890
Archiulus lyelli Scudder 1895
Archiulus xylobioides Scudder 1868
Unr. Colobognatha Brandt 1834
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Or. Platydesmida de Saussure 1860
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Fm. Andrognathidae Cope 1869
Or. Polyzoniida Cook 1895
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Fm. Polyzoniidae Gervais 1844
Fm. Siphonotidae Cook 1895
Or. Siphonophorida Newport 1844
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Fm. Siphonophoridae Newport 1844
Fm. Siphonorhinidae Cook 1895
G. †Dolesea Hannibal and May 2020
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Dolesea subtila Hannibal and May 2020
Unr. Eugnatha Attems 1898
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Superor. Juliformia Attems 1926
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Or. Julida Leach 1814
Fm. †Proglomeridae Fritsch 1899
Or. Spirobolida Cook 1895
Or. Spirostreptida Cook 1895
Superfm. †Xyloiuloidea Cook 1895
Superor. Merochaeta Cook 1895
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Or. Polydesmida Pocock 1887
Superor. Nematophora Verhoeff 1913
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Or. Callipodida Pocock 1894
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Invalid names: Lysiopetaloidea Verh 1900 [synonym]
Or. Stemmiulida Cook 1895
Fm. †Hexecontasomatidae Hannibal 2000
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G. †Hexecontasoma Hannibal 2000
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Hexecontasoma carinatum Hannibal 2000
Fm. †Oklahomasomatidae Hannibal and May 2020
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G. †Oklahomasoma Hannibal and May 2020
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Oklahomasoma richardsspurense Hannibal and May 2020
G. †Sinosoma Huang and Hannibal 2018
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Sinosoma luopingense Huang and Hannibal 2018
Or. Siphoniulida Pocock 1894
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Fm. Siphoniulidae Pocock 1894
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G. Siphoniulus Pocock 1894
Invalid names: Archiulidae Scudder 1873 [invalid subgroup]
Infracl. Pentazonia Brandt 1833
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Or. Glomeridesmida Cook 1895
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Fm. Glomeridesmidae Latzel 1884
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G. Glomeridesmus Gervais 1844
Superor. Oniscomorpha Pocock 1887
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Or. †Amynilyspedida Hoffman 1969
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Fm. †Amynilyspedidae Hoffman 1969
Fm. †Sphaerherpestidae Fritsch 1899
Or. Glomerida Cook 1895
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Fm. Glomerididae Leach 1815
Invalid names: Sphaerotheriida Brandt 1833 [empty]
Or. †Zosterogrammida Wilson 2005
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Fm. †Zosterogrammidae Wilson 2005
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G. †Casiogrammus Wilson 2005
G. †Purkynia Fritsch 1899
G. †Zosterogrammus Wilson 2005
Or. Chordeumatida Cook 1895
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Fm. Craspedosomatidae Cook 1895
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G. Atractosoma Fanzago 1876
G. Craspedosoma Leach 1814
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Craspedosoma aculeatum Menge 1854
Craspedosoma affine Koch and Berendt 1854
Craspedosoma angulatum Koch and Berendt 1854
Craspedosoma armatum Menge 1854
Craspedosoma setosum Menge 1854
G. †Euzonus Menge 1854
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Euzonus collulum Menge 1854
Invalid names: Heterochordeumatidae Pocock 1894 [empty]
G. †Decorotergum Jell 1983
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Decorotergum warrenae Jell 1983
Fm. Iulidae Leach 1814
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G. Iulus Linnaeus 1758
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Iulus florissantellus Cockerell 1907
Subcl. Penicillata Latreille 1825
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Or. Polyxenida Lucas 1840
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Fm. Lophoproctidae Silvestri 1897
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G. Lophoproctus Pocock 1894
Superfm. Polyxenoidea Lucas 1840
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Fm. Polyxenidae Lucas 1840
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G. †Electroxenus Nguyen Duy-Jacquemin and Azar 2004
G. †Libanoxenus Nguyen Duy-Jacquemin and Azar 2004
G. Pauropsxenus Silvestri 1948
G. Polyxenus Latreille 1802
Superfm. Synxenoidea Silvestri 1923
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Fm. Synxenidae Silvestri 1923
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G. Phryssonotus Scudder 1885
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Invalid names: Lophonotus Menge 1854 [replaced]
Fm. †Proiulidae Fritsch 1899
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G. †Tomiulus Martynov 1936
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Tomiulus angulatus Martynov 1936
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Locomotion: actively mobilep
Created: 2004-02-29 09:38:49
Modified: 2004-02-29 11:38:49
Source: p = phylum
Reference: Kiessling 2004

Age range

Maximum range based only on fossils: base of the Homerian to the top of the Rancholabrean or 430.50000 to 0.01200 Ma
Minimum age of oldest fossil (stem group age): 427.4 Ma

Collections (188 total)


Time interval Ma Country or state Original ID and collection number
Wenlock433.4 - 427.4United Kingdom (South Lanarkshire) Casiogrammus ichthyeros (155868)
Wenlock433.4 - 427.4United Kingdom (Scotland) Cowiedesmus eroticopodus, Albadesmus almondi, Pneumodesmus newmani (26907)
Homerian430.5 - 427.4United Kingdom (Scotland) Albadesmus almondi (155916) Cowiedesmus eroticopodus, Albadesmus almondi (155913) Diplopoda indet. (155915) Pneumodesmus newmani (84333)
Pridoli423.0 - 419.2United Kingdom (England) Kampecaris dimnorensis (11128)
Lochkovian419.2 - 410.8United Kingdom (Scotland) Archidesmus macnicoli (155912 155918) Archidesmus macnicoli, Sigmastria dilata (155933) Kampecaris forfarensis (195121 195122 195123 195127 195128)
Lochkovian - Pragian419.2 - 407.6United Kingdom (England) Kampecaris dinmorensis (233872)
Early/Lower Emsian409.1 - 402.5Germany (Alken) Eoarthropleura devonica (27049)
Emsian407.6 - 393.3United Kingdom (Scotland) Kampecaris tuberculata (120607)
Late/Upper Emsian402.5 - 391.9Canada (New Brunswick) Gaspestria genselorum (155932)
Late/Upper Emsian402.5 - 391.9Canada (Quebec) Gaspestria genselorum (155931)
Frasnian382.7 - 372.2USA (New York) Tiphoscorpio hueberi (14295)
Frasnian382.7 - 372.2Canada (Quebec) Zanclodesmus willetti (155925)
Late/Upper Famennian364.7 - 360.7USA (Pennsylvania) Orsadesmus rubecollus (35044)
Tournaisian358.9 - 346.7United Kingdom (Scotland) Anthracodesmus macconochiei (127550) Archidesmida indet. 1 (195789) Archipolypoda indet. 3 (194781) Chilognatha indet. 5, Archipolypoda indet. 2 (195401) Juliformia indet. 4, Woodesmus sheari (183472) Woodesmus sheari (195790)
Visean346.7 - 330.9United Kingdom (Wales) Halonia sp. (34478)
Brigantian336.0 - 330.9Germany Euphoberia sp., Arthropleura sp. (153594)
Pendleian330.9 - 327.0United Kingdom (Scotland) Diplopoda indet. (134654)
Pendleian330.9 - 327.0United Kingdom (England) Arthropleura sp. (224886)
Serpukhovian330.9 - 323.2Poland Palaeosoma robustum (195184)
Pennsylvanian323.2 - 298.9USA (New Mexico) Diplopoda indet. (217076)
Westphalian A318.7 - 316.9Belgium Acantherpestes sp., Xylobius renieri (155630) Arthropleura maillieuxi (155633 155634) Arthropleura sp. (155640)
Westphalian A318.7 - 316.9France Arthropleura armata (155422)
Westphalian A318.7 - 316.9United Kingdom (England) Acantherpestes giganteus, Euphoberia robusta, Euphoberia armigera (127287) Euphoberia woodwardi (96873) Eurypterus ferox (155939) Xylobius platti (34905) Xyloiulus sp. (124367)
Westphalian A318.7 - 316.9France (Nord-Pas-de-Calais) Arthropleura armata (155425 155426) Arthropleura sp. (155427 155428)
Westphalian A318.7 - 316.9Canada (Nova Scotia) Amynilyspes springhillensis (155664) Xylobius sigillariae, Amynilyspes sp., Archiulus xylobioides, Archiulus lyelli, Xylobius similis, Xylobius fractus, Xylobius dawsoni, Archiulus euphoberioides (28069) Xylobius sigillariae, Archiulus xylobioides (155665)
Westphalian A318.7 - 316.9Netherlands (Limburg) Lauravolsella willemeni (110966)
Westphalian A318.7 - 316.9United Kingdom (Lancashire) Arthropleurida indet. (172765)
Westphalian318.7 - 307.5Germany (Saarbrücken) Arthropleura armata, Arthropleura punctata (124243)
Westphalian318.7 - 307.5Germany Arthropleura armata (137875)
Westphalian318.7 - 307.5United Kingdom Eurypterus mammatus (155940)
Westphalian B316.9 - 314.6France (Nord-Pas-de-Calais) Arthropleura armata (155424) Arthropleura mammata (155423)
Westphalian B316.9 - 314.6United Kingdom (England) Amynilyspes sp. (105060) Arthropleura moyseyi (127218) Euphoberia ferox, Myriacantherpestes sp. (125238) Lepidoptera caterpillar indet. (155938)
Westphalian B316.9 - 314.6Belgium Arthropleura mammata (155628 155632) Arthropleura mammata, Euphoberia ferox (155631) Arthropleura sp. (155641 155642 155643)
Westphalian B316.9 - 314.6United Kingdom Euphoberia brownii (195182) Euphoberia ferox (195183)
Westphalian B316.9 - 314.6United Kingdom (Staffordshire) Anaxeodesmus diambonotus (153546)
Moscovian315.2 - 307.0Czech Republic (Pilsen) Euphoberia histrix, Euphoberia varians, Euphoberia absens, Acantherpestes gigas, Acantherpestes vicinus, Acantherpestes ornatus, Acantherpestes foveolatus, Julus constans, Isojulus setipes, Isojulus marginatus, Pleurojulus biornatus, Pleurojulus levis, Pleurojulus aculeatus, Pleurojulus longipes, Pleurojulus falcifer, Pleurojulus pinguis, Xylobius pstrossi, Julus costulatus, Xylobius sellatus, Xylobius tabulatus, Amynilyspes typicus, Amynilyspes crescens, Archiscudderia paupera, Archiscudderia tapeta, Archiscudderia coronata, Archiscudderia regularis, Archiscudderia problematica, Glomeropsis ovalis, Glomeropsis crassa, Glomeropsis magna, Purkynia lata (128574)
Westphalian C314.6 - 309.8Germany Arthropleura affinis (138349) Arthropleura armata (137874 138002 138345 138350 138354 138362 138364) Arthropleura armata, Branchipusites anthracinus (138365)
Westphalian C314.6 - 309.8Belgium Arthropleura britannica (123823)
Westphalian C314.6 - 309.8France (Nord-Pas-de-Calais) Arthropleura sp. (155429)
Westphalian C314.6 - 309.8Germany (Saarland) Arthropleura armata (124244) Arthropleura armata, Arthropleura enodis (122601) Arthropleura armata, Troxites germari (123956) Chonionotus lithanthracis (122623)
Westphalian C - Westphalian D314.6 - 307.5Germany Arthropleura armata (123825)
Westphalian C - Westphalian D314.6 - 307.5Canada (Nova Scotia) Xylobius sp. (80111)
Westphalian D309.8 - 307.5USA (Illinois) Euphoberia armigera (155944) Euphoberia armigera, Acantherpestes major, Euphoberia carri (122528) Euphoberia armigera, Acantherpestes major, Euphoberia flabellata (154202) Euphoberia armigera, Euphoberia anguilla (155943) Euphoberia armigera, Euphoberia horrida, Euphoberia granosa, Euphoberia carri, Amynilyspes wortheni (155941) Euphoberia armigera, Euphoberia major, Euphoberia granosa (152106) Euphoberia armigera, Xylobius frustulentus, Acantherpestes major, Euphoberia granosa, Euphoberia spinulosa, Euphoberia simplex, Euphoberia tracta (122521) Euphoberia armigera, Xylobius frustulentus, Acantherpestes major, Euphoberia horrida, Euphoberia granosa, Euphoberia carri, Euphoberia anguilla, Archiulus glomeratus (122527) Euphoberia sp., Pleurojulus biornatus, Euphoberia armigera, Xylobius mazonus, Xylobius frustulentus, Acantherpestes major, Euphoberia granosa, Euphoberia anguilla, Acantherpestes inequalis, Euphoberia hystricosa, Euphoberia cuspidata, Euphoberia spinulosa, Euphoberia simplex, Euphoberia tracta, Archiulus glomeratus (21981) Hexecontasoma carinatum (122493 132455) Pleurojulus biornatus (133734) Pleurojulus biornatus, Amynilyspes wortheni, Sphaerotheriida indet., Oniscomorpha indet. (112638) Pleurojulus biornatus, Zosterogrammus stichostethus (112630)
Westphalian D309.8 - 307.5United Kingdom (Somerset) Arthropleura sp., Arthropleura armata (153596)
Westphalian D309.8 - 307.5Germany Halonia irregularis (169136)
Westphalian D309.8 - 307.5Germany (Saarland) Arthropleura armata (132751)
Westphalian D309.8 - 307.5USA (Ohio) Plagiascetus lateralis, Xylobius sp. (85292)
Westphalian D309.8 - 307.5United Kingdom (Wales) Euphoberia sp., Pleurojulus levis (127543)
Westphalian D309.8 - 307.5USA (Pennsylvania) Arthropleura cristata (225177)
Westphalian D309.8 - 307.5United Kingdom (England) Arthropleura sp. (126868)
Stephanian307.5 - 300.4France Arthropleura zeilleri (233867)
Stephanian307.5 - 300.4Italy (Sardinia) Arthropleura armata (153483)
Stephanian307.5 - 300.4France (Saône-et-Loire) Arthropleura sp. (236920)
Stephanian A305.1 - 301.5USA (Ohio) Arthropleura cristata (11188)
Gzhelian303.7 - 298.9France Arthropleura fayoli (103635)
Stephanian B301.5 - 301.0Spain Arthropleura armata (169144) Arthropleura sp. (233868)
Stephanian B301.5 - 301.0France Amynilyspes fatimae, Blanziulus parriati (161212) Arthropleura sp., Amynilyspes fatimae, Blanziulus parriati (122490)
Stephanian B301.5 - 301.0Czech Republic Anthracojulus pictus (155305)
Stephanian B301.5 - 301.0Czech Republic (Rakovník) Julus pictus (154179)
Stephanian C301.0 - 300.4Spain Arthropleura sp. (233869 233870 233871)
Stephanian C301.0 - 300.4Germany Arthropleura sp. (216757 216758)
Asselian298.9 - 295.5USA (New Mexico) Spirobolida indet. (124386)
Asselian - Sakmarian298.9 - 290.1Germany (Rhineland-Palatinate) Julus brassii, Carcinochelus anthracophilus (122606)
Artinskian290.1 - 279.3USA (Oklahoma) Helminthomorpha indet., Karstiulus fortsillensis, Oklahomasoma richardsspurense, Dolesea subtila (67862)
Artinskian290.1 - 279.3USA (Texas) Xylobius sp. (11149)
Kungurian279.3 - 272.3USA (Texas) Xylobius sp. (195185)
Griesbachian251.902 - 250.5Russian Federation (Kemerovo) Tomiulus angulatus (105040)
Aegean247.2 - 242.0France (Moselle) Hannibaliulus wilsonae (161211)
Pelsonian247.2 - 242.0China (Yunnan) Diplopoda indet., Sinosoma luopingense (127488)
Pliensbachian - Toarcian190.8 - 174.1Australia (Queensland) Decorotergum warrenae (77179)
Early/Lower Barremian130.0 - 125.45Lebanon Electroxenus jezzinensis (123000) Libanoxenus hammanaensis (122471)
Cenomanian100.5 - 93.9France (Alpes-de-Haute- Provence) Phryssonotus sp. (123441)
Early/Lower Cenomanian99.6 - 93.5Myanmar Glomerida indet. (172900) Phryssonotus burmiticus, Polyxenidae indet., Polydesmida indet., Siphonophoridae indet., Polyzoniida indet., Heterochordeumatidae indet., Andrognathidae indet. (144466) Phryssonotus burmiticus, Siphonophora hui, Monstrodesmus grimaldii, Pauropsxenus ordinatus, Pauropsxenus extraneus, Propolydesmus cretaceus, Siphonorhinus globosus, Siphonorhinus peculiaris (161726) Polyxenida indet., Phryssonotus burmiticus, Polydesmida indet., Siphonophoridae indet., Polyzoniida indet. (196547) Polyxenida indet., Stemmiulida indet., Cambalidea indet., Polyxenidae indet., Polydesmida indet., Siphonophorida indet., Siphonophoridae indet., Glomerida indet., Polyzoniida indet., Glomeridesmidae indet., Siphonotidae indet., Siphonorhinidae indet., Chordeumatida indet., Heterochordeumatidae indet., Andrognathidae indet. (184692) Polyxenus burmiticus (123020) Siphonophora hui (199763) Stemmiulida indet., Cambalidea indet., Cambalidae indet., Polydesmida indet., Glomerida indet., Glomeridesmidae indet., Siphonotidae indet., Siphoniulus muelleri, Siphoniulus preciosus, Heterochordeumatidae indet., Andrognathus burmiticus, Callipodida indet. (184444)
Middle Campanian - Late/Upper Campanian83.5 - 70.6Mongolia (Omnogov) Diplopoda indet. (13776 28243 37022)
Bridgerian50.3 - 46.2USA (Wyoming) Iulus telluster (121306)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Craspedosoma angulatum (138659) Hyleoglomeris groehni (110735 206315)
Priabonian38.0 - 33.9Poland Craspedosoma angulatum, Lophonotus hystrix, Polyxenus colurus, Polyxenus lophurus, Polyxenus caudatus, Glomeris denticulata, Julus politus, Julus badius, Julus rubens, Craspedosoma obtusangulum, Craspedosoma aculeatum, Craspedosoma armatum, Craspedosoma cylindricum, Craspedosoma setosum, Euzonus collulum (120811) Polyxenus conformis, Polyxenus ovalis, Julus laevigatus, Craspedosoma affine (109625)
Chadronian37.2 - 33.9USA (Colorado) Iulus florissantellus (106381) Parajulus cockerelli (123119)
MP 3028.4 - 23.03Germany (Nordrhein-Westfalen) Julus antiquus (127628)
Chattian28.1 - 23.03France Protosilvestria sculpta (160861 160862)
Aquitanian - Burdigalian23.03 - 15.97Australia (Queensland) Julida indet. (106497)
Early/Lower Miocene23.03 - 15.97Mexico (Chiapas) Anbarrhacus adamantis (217222) Maatidesmus paachtun (217221) Myrmecodesmus sp. (217216) Parastemmiulus elektron (157206) Polydesmida indet. (198592 198593 217215) Siphonophora sp. (217217) Spirobolida indet., Diplopoda indet., Juliformia indet., Stemmiulidae indet., Polydesmida indet., Xystodesmidae indet. (172774) Stemmiulidae indet., Polydesmida indet., Platyrhacidae indet., Siphonophora sp., Xystodesmidae indet. (217220)
Burdigalian20.44 - 15.97India (Kerala) Polyxenus miocenica (184505)
Burdigalian20.44 - 15.97Kenya Diplopoda indet. (27856)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Docodesmus sp., Lophoproctus sp., Siphonophora sp., Lophodesmus sp., Inodesmus sp., Siphonophora hoffmani, Siphonophora velezi (159268) Siphonocybe sp., Docodesmus brodzinskyi (122473) Stemmiulidae indet., Chelodesmidae indet., Docodesmus sp., Lophoproctus sp., Glomeridesmus sp., Epinannolene sp., Siphonophora sp., Siphonotus sp., Iomus sp., Psochodesmus sp., Dasyodontus sp., Inodesmus sp., Siphonophora hoffmani (122472)
Sarmatian12.7 - 11.608Germany (Wurttemberg) Julidae indet., Julus antiquus (113752)
Pliocene5.333 - 2.588USA (Arizona) Parajulus onychis (152091)
Ruscinian - Villanyian5.333 - 2.588Hungary (Barany) Diplopoda indet. (35354)
Early/Lower Pleistocene2.588 - 0.781Hungary (Barany) Julus sp. (35459)
Calabrian1.8 - 0.774Spain (Murcia) Diplopoda indet. (34128)
Rancholabrean0.3 - 0.012USA (California) Spirobolus australis (150336)
Late/Upper Pleistocene (high glacial)0.129 - 0.0117USA (Texas) Diplopoda indet. (93103)
Late/Upper Pleistocene0.129 - 0.0117Jamaica Rhinocricus spp., Chondrotropis sp., Cyclodesmus porcellanus, Caraibodesmus verrucosus (224573)
Late/Upper Pleistocene (glacial)0.129 - 0.0117USA (California) Julus cavicola (155393)
Late/Upper Pleistocene0.129 - 0.0117USA (California) Diplopoda indet. (225801) Julus occidentalis (74235)