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Nanhsiungchelyidae

Reptilia - Testudines - Nanhsiungchelyidae

Taxonomy

Nanhsiungchelyidae was named by Yeh (1966). Its type is Nanhsiungchelys.

It was assigned to Trionychoidea by Hirayama et al. (2001); to Cryptodira by Yeh (1966) and Tong et al. (2012); to Adocusia by Syromyatnikova and Danilov (2009), Danilov et al. (2013) and Syromyatnikova and Danilov (2013); to Eucryptodira by Peng et al. (2001), Hutchison and Holroyd (2003), Hutchison et al. (2013) and Mallon and Brinkman (2018); to Trionychoidae by Ke et al. (2021); and to Cryptodira by Joyce and Norell (2005) and Ke et al. (2023).

Subtaxa

Anomalochelys, Basilemys, Bulganemys, Bulganemys, Charitonyx, Hanbogdemys, Hanbogdemysa, Jiangxichelys, Kharakhutulia, Nanhsiungchelys (type genus), Yuchelys, Zangerlia

Synonyms

Charitonyx was named by Chkhikvadze (1980). It is not extant.

It was considered a nomen dubium by Sukhanov et al. (2008).
Hanbogdemysa was named by Sukhanov (2000). It is not extant. Its type is Hanbogdemysa orientalis.

It was considered a nomen nudum by Sukhanov and Narmandakh (2006) and Sukhanov et al. (2008).

It was assigned to Nanhsiungchelyidae by Sukhanov (2000).
Bulganemys was named by Sukhanov and Narmandakh (2000). Its type is Bulganemys jaganchobili.

It was considered a nomen nudum by Sukhanov and Narmandakh (2006).

It was assigned to Nanhsiungchelyidae by Sukhanov (2000).

Synonymy list

Year	Name and author
1966	Nanhsiungchelyidae Yeh p. 191
1980	Charitonyx Chkhikvadze
2000	Hanbogdemysa Sukhanov
2000	Bulganemys Sukhanov p. 342 fig. 17.24
2000	Hanbogdemysa Sukhanov pp. 3340-342 fig. 17.23
2000	Bulganemys Sukhanov and Narmandakh p. 342
2001	Nanhsiungchelyidae Hirayama et al. p. 129
2001	Nanhsiungchelyidae Peng et al. p. 26
2003	Nanhsiungchelyidae Hutchison and Holroyd
2005	Nanhsiungchelyidae Joyce and Norell p. 4
2009	Nanhsiungchelyidae Syromyatnikova and Danilov p. 75
2012	Nanhsiungchelyidae Tong et al.
2013	Nanhsiungchelyidae Danilov et al.
2013	Nanhsiungchelyidae Hutchison et al.
2013	Nanhsiungchelyidae Syromyatnikova and Danilov
2018	Nanhsiungchelyidae Mallon and Brinkman
2021	Nanhsiungchelyidae Ke et al.
2023	Nanhsiungchelyidae Ke et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda

Rank	Name	Author
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
—	Testudinata	(Oppel 1811)
order	Testudines	Batsch 1788
suborder	Cryptodira
family	Nanhsiungchelyidae
family	Nanhsiungchelyidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Nanhsiungchelyidae Yeh 1966

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G. †Anomalochelys Hirayama et al. 2001

†Anomalochelys angulata Hirayama et al. 2001

G. †Basilemys Hay 1902

†Basilemys gaffneyi Sullivan et al. 2012

†Basilemys imbricarius Cope 1876

†Basilemys morrinensis Mallon and Brinkman 2018

†Basilemys nobilis Hay 1911

†Basilemys ogmius Cope 1875

†Basilemys praeclara Hay 1911

†Basilemys sinuosa Riggs 1906

†Basilemys variolosa Cope 1876

G. †Bulganemys Sukhanov and Narmandakh 2006

Invalid names: Bulganemys jaganchobili Sukhanov and Narmandakh 2000 [nomen nudum]

G. †Hanbogdemys Sukhanov and Narmandakh 2006

†Hanbogdemys jaganchobili Sukhanov and Narmandakh 2006

†Hanbogdemys orientalis Sukhanov and Narmandakh 1975

G. †Jiangxichelys Tong and Mo 2010

†Jiangxichelys ganzhouensis Tong and Mo 2010

†Jiangxichelys neimongolensis Brinkman and Peng 1996

G. †Kharakhutulia Sukhanov et al. 2008

†Kharakhutulia kalandadzei Sukhanov et al. 2008

G. †Nanhsiungchelys Yeh 1966

†Nanhsiungchelys wuchingensis Yeh 1966

†Nanhsiungchelys yangi Ke et al. 2023

G. †Yuchelys Tong et al. 2012

†Yuchelys nanyangensis Tong et al. 2012

G. †Zangerlia Mlynarski 1972

†Zangerlia dzamynchondi Sukhanov and Narmandakh 2006

†Zangerlia testudinimorpha Mlynarski 1972

†Zangerlia ukhaachelys Joyce and Norell 2005

Invalid names: Zangerlia dzamynchondi Sukhanov and Narmandakh 2000 [nomen nudum]

Invalid names: Bulganemys Sukhanov and Narmandakh 2000 [nomen nudum], Charitonyx Chkhikvadze 1980 [nomen dubium], Hanbogdemysa Sukhanov 2000 [nomen nudum]

Diagnosis

Reference	Diagnosis
H.-k. Yeh 1966	With the diagnosis of the type genus Nanhsiungchelys.
R. Hirayama et al. 2001	Large trionychoids with shell length well exceeding over 50 cm, and up to 90 cm; shell surface deeply sculptured by the linearly arranged ridges and vallies; marginal region of carapace and plastron thickened; neural series complete to suprapygals except for Basilemys nobilis; anterior portion of epiplastron projected and enormously thickened; humero-pectoral sulcus deeply intersecting the entoplastron; incisura columellae auris closed posterior; coracoid fan-shaped; thelial process of ilium absent; two or fewer short and stout phalanges in manus and pes; limbs covered by epidermal bones with pitted sculpture.