PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Claosaurus agilis

Reptilia

Taxonomy

Hadrosaurus agilis was named by Marsh (1872). Its type specimen is YPM 1190, a partial skeleton, and it is a 3D body fossil. Its type locality is Smoky Hill River (YPM 1190), which is in a Coniacian/Santonian marine shale in the Niobrara Formation of Kansas. It is the type species of Claosaurus.

It was recombined as Thespesius agilis by Hay (1902); it was recombined as Claosaurus agilis by Zittel (1890), Marsh (1890), Marsh (1896), Williston (1898), Nopcsa (1900), Nopcsa (1901), Williston (1902), Hatcher (1902), Beecher (1902), Brown (1910), Brown (1913), Perrier (1928), Hay (1930), Osborn (1930), Russell (1930), Gilmore (1933), Lull and Wright (1942), Gregory (1948), Tatarinov (1964), Kuhn (1964), Weishampel and Weishampel (1983), Coombs (1988), Weishampel and Horner (1990), Horner (1992), Carpenter et al. (1995), Breithaupt (1999), Horner et al. (2004), Lund and Gates (2006), Godefroit et al. (2008), Prieto-Márquez (2010), Prieto-Márquez and Norell (2010), Prieto-Márquez (2010), McDonald et al. (2010), Prieto-Marquez and Salinas (2010), Prieto-Márquez and Wagner (2010), Prieto-Márquez (2011), Prieto-Márquez (2011), Prieto-Márquez (2012), Ramírez-Velasco et al. (2012), Zheng et al. (2013), Prieto-Márquez et al. (2016), Prieto-Márquez et al. (2016), Cruzado-Caballero and Powell (2017) and Dalla Vecchia (2020).

Synonymy list

Year	Name and author
1872	Hadrosaurus agilis Marsh
1875	Hadrosaurus agilis Cope p. 247
1877	Hadrosaurus agilis Marsh p. 349
1890	Claosaurus agilis Marsh
1890	Claosaurus agilis Zittel p. 763
1896	Claosaurus agilis Marsh p. 223 fig. 55
1898	Claosaurus agilis Williston p. 69
1900	Claosaurus agilis Nopcsa p. 557
1901	Claosaurus agilis Nopcsa p. 211
1902	Claosaurus agilis Beecher p. 313
1902	Claosaurus agilis Hatcher p. 385
1902	Thespesius agilis Hay p. 502
1902	Claosaurus agilis Williston p. 247
1910	Claosaurus agilis Brown p. 273
1913	Claosaurus agilis Brown p. 389
1928	Claosaurus agilis Perrier p. 3088
1930	Claosaurus agilis Hay p. 215
1930	Claosaurus agilis Osborn p. 534
1930	Claosaurus agilis Russell p. 138
1933	Claosaurus agilis Gilmore p. 48
1942	Claosaurus agilis Lull and Wright p. 134
1948	Claosaurus agilis Gregory p. 30
1964	Claosaurus agilis Kuhn p. 17
1964	Claosaurus agilis Tatarinov p. 561
1983	Claosaurus agilis Weishampel and Weishampel p. 44
1988	Claosaurus agilis Coombs, Jr. p. 817
1990	Claosaurus agilis Weishampel and Horner p. 556
1992	Claosaurus agilis Horner p. 15
1995	Claosaurus agilis Carpenter et al. p. 289
1999	Claosaurus agilis Breithaupt p. 62
2004	Claosaurus agilis Horner et al. p. 439
2006	Claosaurus agilis Lund and Gates p. 271
2008	Claosaurus agilis Godefroit et al. p. 66
2010	Claosaurus agilis McDonald et al. p. 33 fig. 39
2010	Claosaurus agilis Prieto-Marquez and Salinas p. 835 fig. 20
2010	Claosaurus agilis Prieto-Márquez p. 1
2010	Claosaurus agilis Prieto-Márquez and Norell p. 19
2010	Claosaurus agilis Prieto-Márquez and Wagner p. 1243 fig. 3
2011	Claosaurus agilis Prieto-Márquez p. 62
2012	Claosaurus agilis Prieto-Márquez p. 526
2012	Claosaurus agilis Ramírez-Velasco et al. p. 380
2013	Claosaurus agilis Zheng et al.
2016	Claosaurus agilis Prieto-Márquez et al. p. 1
2017	Claosaurus agilis Cruzado-Caballero and Powell p. 11
2020	Claosaurus agilis Dalla Vecchia p. 585

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()

Rank	Name	Author
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Ornithischia	()
—	Genasauria
—	Cerapoda
—	Ornithopoda	()
—	Clypeodonta
—	Iguanodontia	()
—	Dryomorpha
—	Styracosterna
—	Hadrosauriformes
—	Hadrosauroidea	()
genus	Claosaurus
species	agilis	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Claosaurus agilis Marsh 1872

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Diagnosis

Reference	Diagnosis
A. Prieto-Márquez 2011	Hadrosauroid dinosaur characterized by the unique combination of the following humeral and iliac characters: deltopectoral crest length less than 48% of humerus length and with wide arcuate laterodistal corner; anteroposterior width of supraacetabular crest being approximately 75 per cent of length of central plate of ilium; and ventral apex of supraacetabular crest positioned anterior to the posteroventral corner of ischial peduncle of ilium.
A. Prieto-Márquez et al. 2016	Claosaurus agilis was diagnosed by a unique combination of characters consisting of deltopectoral crest length less than 48% of the humeral length and with wide arcuate laterodistal corner, and ilium with supraacetabular crest as wide as 75% of the length of the central plate, with ventral apex of the crest positioned cranial to the caudal tuberosity of the ischiadic process.