Basic info | Taxonomic history | Classification | Included Taxa |
Morphology | Ecology and taphonomy | External Literature Search | Age range and collections |
Heterocoenia
Taxonomy
Heterocoenia was named by Milne-Edwards and Haime (1848) [Sepkoski's age data: J u K Maes Sepkoski's reference number: 796,920,1270]. It is not extant.
It was assigned to Cyathophorinae by Wells (1956); to Scleractinia by Sepkoski (2002); to Stylinidae by Squires and Demetrion (1992) and López-Pérez (2005); and to Heterocoeniidae by Baron-Szabo (2006) and Baron-Szabo (2021).
It was assigned to Cyathophorinae by Wells (1956); to Scleractinia by Sepkoski (2002); to Stylinidae by Squires and Demetrion (1992) and López-Pérez (2005); and to Heterocoeniidae by Baron-Szabo (2006) and Baron-Szabo (2021).
Species
H. bacellaris, H. balkhanensis, H. decipiens, H. dendroides (syn. H. garumnica), H. densa, H. distans, H. erecta, H. exigua, H. fuchsi, H. gracilis, H. grandis, H. hilli, H. inflexa, H. minima, H. minuta, H. minutisima, H. pusilla, H. tetraseptata, H. verrucosa, H. washitaensis, Hexasmilia elmari, Hexasmilia ferryi
Species lacking formal opinion data
Synonyms
|
Synonymy list
Year | Name and author |
---|---|
1848 | Heterocoenia Milne-Edwards and Haime |
1857 | Baryhelia Milne-Edwards and Haime |
1870 | Hexasmilia de Fromentel |
1936 | Miyakosmilia Eguchi |
1952 | Bacillastraea Alloiteau |
1956 | Heterocoenia Wells |
1956 | Miyakosmilia Wells |
1977 | Paronastraea Beauvais |
1992 | Heterocoenia Squires and Demetrion p. 18 |
2002 | Baryhelia Sepkoski |
2002 | Heterocoenia Sepkoski |
2002 | Miyakosimilia Sepkoski |
2002 | Pachycoenia Sepkoski |
2002 | Paronastrara Sepkoski |
2005 | Heterocoenia López-Pérez p. 81 |
2006 | Heterocoenia Baron-Szabo p. 89 |
2008 | Hexasmilia Löser |
2021 | Heterocoenia Baron-Szabo |
Is something missing? Join the Paleobiology Database and enter the data
|
|
If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Heterocoenia Milne-Edwards and Haime 1848
show all | hide all
†Heterocoenia bacellaris Goldfuss 1826
†Heterocoenia balkhanensis Kuzmicheva 1987
†Heterocoenia decipiens Hackemesser 1936
†Heterocoenia decipiens ramosa Hackemesser 1936
†Heterocoenia densa Eguchi 1936
†Heterocoenia distans Milne et al. 1848
†Heterocoenia erecta Felix 1903
†Heterocoenia exigua Michelin 1846
†Heterocoenia fuchsi Felix 1903
†Heterocoenia gracilis Quenstedt 1881
†Heterocoenia grandis Reuss 1854
†Heterocoenia hilli Wells 1932
†Heterocoenia inflexa Eichwald 1865
†Heterocoenia minima d'Orbigny 1850
†Heterocoenia minuta Morycowa 1971
†Heterocoenia minutisima Reig 1997
†Heterocoenia pusilla Reig Oriol 1992
†Heterocoenia tetraseptata Eguchi 1951
†Heterocoenia verrucosa Reuss 1854
†Heterocoenia washitaensis Wells 1933
Invalid names: Bacillastraea [synonym], Baryhelia Milne-Edwards and Haime 1857 [synonym], Hexasmilia de Fromentel 1870 [synonym], Miyakosmilia Eguchi 1936 [synonym], Pachycoenia [synonym], Paronastraea Beauvais 1977 [synonym]
Diagnosis
Reference | Diagnosis | |
---|---|---|
R.C. Baron-Szabo 2021 | Colonial massive, hemispherical, foliose, encrusting, ramose, plocoid, cerio-plocoid, (sub-) fasciculate; subphaceloid to reptoid when developments of coenosteum reduced. Budding extracalicular, extracalicular-marginal, and by septal division often dividing the corallite in three new ones (“trinity-arrangement” sensu Baron-Szabo,
2014). Corallites circular to elongate or irregularly polygonal in outline. They are directly united by their walls, or separated by extensive vesicular to dense coenosteum, or loosely connected by fragments of exotheca in form of traverses. Septa compact, arranged in various symmetries (e.g., trimerally, hexamerally, bilaterally, indistinct). One main septum, with remaining septa sometimes reduced to rudimentary spines. Costate zone present, weakly developed, or absent. Colony surface granulated or smooth. Columella, synapticulae, and paliform structures absent. Intertrabecular distance ranging between 60 and 120 μm in septa. Endothecal dissepiments thin, mainly tabulate to subtabulate, vesicular (often in peripheral and thecal areas). Exothecal dissepiments large vesicular. Wall often thick, possibly septothecal and paraseptothecal. Corallite wall tends to become more flaky, “bubbly” (cf. sclerenchymal deposits sensu Kołodziej, 1995, p. 4), and disintegrated during the process of budding. |
Measurements
No measurements are available
|
|
||||
|
|
||||
|
|
||||
|
|||||
|
|
||||
|
|
||||
|
|||||
|
|
||||
|
|
||||
Source: f = family, o = order, c = class | |||||
References: Kiessling 2004, Kiessling 2008 |
Age range: base of the Late/Upper Kimmeridgian to the top of the Lutetian or 155.70000 to 41.30000 Ma
Collections (47 total)
Time interval | Ma | Country or state | Original ID and collection number |
---|---|---|---|
Late/Upper Kimmeridgian | France | Baryhelia crassa (68507) | |
Early/Lower Tithonian | Lebanon | Baryhelia hexaconena (34869) | |
Early/Lower Tithonian | Italy | H. minutisima (154046) | |
Late/Upper Berriasian | Austria | H. inflexa (192211) | |
Valanginian | Switzerland (Alpstein ) | H. inflexa (233521) | |
Early/Lower Barremian | Turkmenistan | H. balkhanensis (91685) | |
Late/Upper Barremian | Bulgaria | Hexasmilia sp. (151443) | |
Late/Upper Barremian - Early/Lower Aptian | Austria | H. minima, H. inflexa (221476) | |
Barremian | Mexico (Puebla) | H. pusilla (89865) | |
Late/Upper Aptian | Spain (Valencia) | Hexasmilia elmari (84708) | |
Late/Upper Aptian | Japan (Iwate) | H. explanata, H. tetraseptata, Miyakosmilia densa (91282) H. hideshimaensis (91327) H. hideshimaensis, H. mojiensis (91324) H. sp., H. hideshimaensis (91328) Miyakosmilia ishidai (91325) | |
Late/Upper Aptian | Spain | Hexasmilia elmari (67606) Hexasmilia sp., Hexasmilia elmari (67607) | |
Early/Lower Aptian | Greece | H. sp. (34719 60375) | |
Early/Lower Aptian | Germany | H. sp., H. minima (221477) | |
Early/Lower Aptian | Switzerland | H. minima (221503) | |
Early/Lower Aptian | Romania | H. sp., H. minima (69201) | |
Albian | Greece | H. sp., H. minuta (74811) | |
Early/Lower Albian | Mexico | H. exigua (21254) | |
Early/Lower Albian | USA (Texas) | H. hilli (74765) | |
Late/Upper Albian | USA (Texas) | H. washitaensis (123322) | |
Cenomanian | France | Pachycoenia sp. (74200) | |
Early/Lower Cenomanian | Spain | H. provincialis, H. sp. (23537) | |
Early/Lower Cenomanian | Greece | H. distans (203930) | |
Late/Upper Cenomanian | Germany | Pachycoenia clemens (74193) | |
Turonian - Campanian | Georgia | H. fuchsi (91629) | |
Late/Upper Turonian - Coniacian | Austria | H. erecta (27296) | |
Early/Lower Coniacian | Austria | H. provincalis (24265) | |
Santonian | Spain | H. reussi (67610) | |
Santonian | Armenia | H. exigua (86675) | |
Santonian | Austria | H. grandis, H. exigua, H. verrucosa (74895) | |
Late/Upper Santonian | France (Provence-Alpes-Cote d'Azur) | Hexasmilia ferryi (84709) | |
Campanian | Serbia and Montenegro | H. exigua (79226) | |
Campanian | Spain | H. dendroides, H. exigua, H. verrucosa (67084) | |
Late/Upper Campanian | Bulgaria | Pachycoenia corbariensis (67576) | |
Maastrichtian | Spain (Cataluna) | H. garumnica (88323) | |
Late/Upper Maastrichtian | Netherlands (Limburg) | H. bacillaris (85271 85272 85273) | |
Thanetian | Slovakia | Scleractinia indet. (88184) | |
Ypresian | Mexico (Baja California Sur) | H. sp. (5495) | |
Lutetian | Senegal (Thies) | Bacillastraea senegalensis (76723) |