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Grallator tenuis

Reptilia - Grallatoridae

Taxonomy
Grallator tenuis was named by Hitchcock (1858). It is considered to be a form taxon. Its type specimen is AC 12/3-12/4, a footprint, and it is a trace fossil. Its type locality is below Turners Falls, tracksite, which is in a Hettangian/Sinemurian fluvial-lacustrine sandstone/shale in the Portland Formation of Massachusetts.

Sister species lacking formal opinion data
G. pisanus

Synonyms
  • Anchisauripus hitchcocki was named by Lull (1904). It is considered to be a form taxon. Its type specimen is AC 56/1, a footprint, and it is a trace fossil. Its type locality is Lily Pond Quarry, Gill, which is in a Hettangian fluvial-lacustrine siltstone/sandstone in the Turners Falls Formation of Massachusetts.

    It was recombined as Anchisauripus (Grallator) hitchcocki by Olsen (1981); it was recombined as Grallator (Anchisauripus) hitchcocki by Olsen and Galton (1984) and Lockley (1986); it was synonymized subjectively with Grallator tenuis by Weems (1992); it was considered a nomen dubium by Baird (1957).
  • Grallator formosus was named by Hitchcock (1858). It is considered to be a form taxon. Its type specimen is AC 3/1, 4/1, a footprint, and it is a trace fossil. Its type locality is Moody's Corner, which is in a Hettangian/Sinemurian fluvial-lacustrine mudstone/sandstone in the Portland Formation of Massachusetts.

    It was synonymized subjectively with Grallator tenuis by Weems (1992).
  • Grallator cuneatus was named by Hitchcock (1858). It is considered to be a form taxon. Its type specimen is AC 25/1, a footprint, and it is a trace fossil. Its type locality is Lily Pond Quarry, Gill, which is in a Hettangian fluvial-lacustrine siltstone/sandstone in the Turners Falls Formation of Massachusetts.

    It was synonymized subjectively with Grallator tenuis by Weems (1992).
  • Atreipus acadianus was named by Olsen and Baird (1986). It is considered to be a form taxon. Its type specimen is PU 21713B, a footprint, and it is a trace fossil. Its type locality is Paddy's Island tracksite, which is in a Carnian terrestrial sandstone/siltstone in the Wolfville Formation of Canada.

    It was synonymized subjectively with Grallator tenuis by Weems (1992); it was synonymized subjectively with Atreipus milfordensis by Hunt and Lucas (2007).
Synonymy list
YearName and author
1858Grallator tenuis Hitchcock pp. 73, Pls. 13, 53
1858Grallator cuneatus Hitchcock p. 74 figs. Pls. 13, 41
1858Grallator formosus Hitchcock p. 75
1861Grallator tenuis Deane p. 36
1861Grallator cuneatus Deane p. 38
1861Grallator formosus Deane p. 39
1871Grallator cuneatus Hitchcock p. xxi
1871Grallator formosus Hitchcock p. xxi
1871Grallator tenuis Hitchcock p. xxi
1889Grallator cuneatus Eyerman p. 32
1889Grallator tenuis Eyerman p. 32
1889Grallator cuneatus Hitchcock p. 118
1889Grallator formosus Hitchcock p. 118
1889Grallator tenuis Hitchcock p. 118
1894Grallator cuneatus Lyman p. 205
1894Grallator tenuis Lyman p. 205
1894Grallator formosus Lyman p. 206
1898Grallator cuneatus Hitchcock p. 401
1898Grallator formosus Hitchcock p. 401
1898Grallator tenuis Hitchcock p. 401
1902Grallator cuneatus Hay p. 545
1902Grallator formosus Hay p. 545
1902Grallator tenuis Hay p. 545
1904Anchisauripus hitchcocki Lull p. 488 fig. 5
1904Grallator cuneatus Lull pp. 495-496
1904Grallator tenuis Lull p. 495 fig. 14
1904Grallator formosus Lull p. 496
1915Anchisauripus hitchcocki Lull p. 182 fig. 39
1915Grallator tenuis Lull p. 201 fig. 54
1915Grallator cuneatus Lull p. 203 fig. 56
1915Grallator formosus Lull p. 205 fig. 57
1917Anchisauripus hitchcocki Lull p. 114
1917Grallator cuneatus Lull p. 116
1917Grallator tenuis Lull p. 116
1917Grallator formosus Lull p. 117
1927Anchisauripus hitchcocki Hitchcock and Grier p. 186
1927Grallator cuneatus Hitchcock and Grier p. 186
1929Grallator formosus Thorpe p. 298
1930Anchisauripus hitchcocki Hay p. 3
1930Grallator cuneatus Hay p. 13
1930Grallator formosus Hay p. 13
1930Grallator tenuis Hay p. 13
1937Grallator tenuis Anonymous p. 12
1952Grallator cuneatus Bock p. 402
1952Grallator tenuis Bock p. 402
1953Grallator tenuis Lull pp. 154-155
1953Grallator cuneatus Lull p. 157
1953Grallator formosus Lull p. 157
1953Anchisauripus hitchcocki Lull pp. 168-169
1957Grallator tenuis Baird p. 470
1958Grallator tenuis Kuhn p. 22
1963Grallator cuneatus Kuhn p. 78
1963Grallator formosus Kuhn p. 78
1963Grallator tenuis Kuhn p. 78
1963Anchisauripus hitchcocki Kuhn p. 81
1965Grallator formosus Nelson p. 7
1969Grallator cuneatus Haubold p. 100 fig. 8
1970Grallator tenuis Demathieu p. 184
1971Anchisauripus hitchcocki Haubold p. 65
1971Grallator cuneatus Haubold p. 67
1971Grallator formosus Haubold p. 67
1971Grallator tenuis Haubold p. 67
1971Grallator formosus Welles p. 29
1974Anchisauripus hitchcocki Haubold p. 128
1979Grallator tenuis Eriksen p. 6
1981Anchisauripus (Grallator) hitchcocki Olsen p. 558
1984Grallator (Anchisauripus) hitchcocki Olsen and Galton p. 98 fig. 4
1984Grallator formosus Thulborn and Wade p. 449
1986Grallator tenuis Lockley p. 22
1986Grallator (Anchisauripus) hitchcocki Lockley p. 36
1986Atreipus acadianus Olsen and Baird p. 698 fig. 6.3C
1986Grallator cuneatus Pannell p. 22
1986Grallator formosus Pannell p. 22
1987Grallator cuneatus Weems p. 18
1989Atreipus acadianus Olsen p. 141
1990Grallator cuneatus Thulborn p. 156
1992Grallator tenuis Weems p. 124
1993Atreipus acadianus Thulborn p. 462
1999Grallator tenuis Gierlinski and Pienkowski p. 336
2000Atreipus acadianus Gand et al. p. 615
2000Anchisauripus hitchcocki Gand et al. p. 617
2002Atreipus acadianus Gierlinski and Niedzwiedzki p. 471
2003Atreipus acadianus Szajna and Hartline p. 266
2006Grallator tenuis Lockley and Eisenberg p. 267
2006Atreipus acadianus Thulborn p. 291 fig. 8
2006Atreipus acadianus Weems p. 372
2006Grallator tenuis Weishampel p. 132
2006Atreipus acadianus Weishampel p. 134
2015Grallator tenuis Pienkowski et al. p. 22
2016Grallator cuneatus Wagensommer et al. p. 171
2016Grallator tenuis Wagensommer et al. p. 171
2021Grallator tenuis Khosla and Bajpai p. 199
2021Atreipus acadianus Klein and Lucas p. 6

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
RankNameAuthor
classReptilia
subclassEureptilia()
—Romeriida
—Diapsida()
—Archosauromorpha(Huene 1946)
—Crocopoda
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
informalAvemetatarsalia
—Ornithodira
—Dinosauromorpha
—Dinosauriformes
—Dinosauria()
—Theropoda()
familyGrallatoridae
genusGrallator
speciestenuisHitchcock 1858

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Grallator tenuis Hitchcock 1858
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Invalid names: Anchisauripus hitchcocki Lull 1904 [synonym], Atreipus acadianus Olsen and Baird 1986 [synonym], Grallator cuneatus Hitchcock 1858 [synonym], Grallator formosus Hitchcock 1858 [synonym]
Diagnosis
ReferenceDiagnosis
R. S. Lull 1904 (Anchisauripus hitchcocki)Divarication of digits I and II, 138°; of II and III, 19°; of III and IV, 14°; of II and IV, 33°. Length of digit II, .060 m.; of III, .088 m.; of IV, .075 m.; of the foot, exclusive of hallux, .119 m.; including the hallux, .140 m. Distance between lateral tips, .067 m.
R. S. Lull 1915 (Anchisauripus hitchcocki)Pes: Length of digit II, 60 mm.; III, 88 mm.; IV, 75 mm.; foot exclusive of hallux, 119 mm., including hallux, 140 mm. Distance between lateral tips, 67 mm. Divarication of digits I and II, 138°; II and III, 19°; II and IV, 33°.
Length of step unknown.