PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Shamosaurinae (disused)

Reptilia - Ankylosauridae

Taxonomy

Shamosaurinae was named by Tumanova (1983). Its type is Shamosaurus.

It was assigned to Ankylosauridae by Tumanova (1983), Tumanova (1986), Tumanova (1987), Vickaryous and Ryan (1997), Ryan (1997), Carpenter and Kirkland (1998), Kirkland (1998), Ford (2000) and Arbour and Currie (2016).

Synonymy list

Year	Name and author
1983	Shamosaurinae Tumanova p. 110
1986	Shamosaurinae Tumanova p. 75
1987	Shamosaurinae Tumanova p. 23
1997	Shamosaurinae Ryan p. 442
1997	Shamosaurinae Vickaryous and Ryan p. 491
1998	Shamosaurinae Carpenter and Kirkland p. 268
1998	Shamosaurinae Kirkland p. 273 fig. 3
2000	Shamosaurinae Ford p. 176
2016	Shamosaurinae Arbour and Currie p. 402

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()

Rank	Name	Author
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Ornithischia	()
—	Genasauria
—	Thyreophora	()
—	Ankylosauria	()
family	Ankylosauridae
subfamily	Shamosaurinae
subfamily	Shamosaurinae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Shamosaurinae Tumanova 1983

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Diagnosis

Reference	Diagnosis
T. A. Tumanova 1983	Ankylosaurids with a tapered forward end of the snout; the anterior wall of the pterygoids slopes gently backward, the pterygoids fuse rigidly with the basisphenoid, the interpterygoidal cavity is small, the quadrate bones fuse with the lower edge of the paroccipital lobes (Fig. I b), the occipital condyle changes from a broad oval to a circular shape.
T. A. Tumanova 1987	Ankylosaurids with anterior part of snout narrow; angle of orbital plane with skull axis less than 25°; anterior wall of pterygoids inclined to the rear; occipital condyle wide oval to round; pterygoids fused with basisphenoid; interpterygoid fenestra very small; orbits at midpoint of skull length.

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

gregariouso

Environment:

terrestrialo

Locomotion:

actively mobileo

Life habit:

ground dwellingo

Diet:

herbivoreo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2004-12-14 12:50:52

Modified:

2004-12-14 14:50:52

Source: o = order

Reference: Marsh 1875

Age range: Early/Lower Cenomanian or 99.60000 to 93.50000 Ma

Collections: one only

Time interval	Ma	Country or state	Original ID and collection number
Early/Lower Cenomanian	99.6 - 93.5	Uzbekistan (Navoi)	Shamosaurinae indet. (58208)