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Ecdysozoa

Taxonomy
Ecdysozoa is an unranked clade.

It was assigned to Paracoelomata by Zrzavy et al. (1998); and to Protostomia by Peterson et al. (2005), Edgecombe et al. (2011) and Nielsen (2012).

Subtaxa
Cephalorhyncha, Cycloneuralia, Nematozoa, Panarthropoda, Scalidophora
Sister taxa
Cycloneuralia, Nematozoa, Panarthropoda, Scalidophora

Synonymy list
YearName and author
1998Ecdysozoa Zrzavy et al.
2005Ecdysozoa Peterson et al. p. 38
2011Ecdysozoa Edgecombe et al. p. 153 figs. Fig. 1
2012Ecdysozoa Nielsen p. 238 figs. Fig. 22.1

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
RankNameAuthor
—Protostomia
—Ecdysozoa
—Ecdysozoa

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. †Ecdysozoa
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Invalid names: Protocephalorhynca Adrianov and Malakhov 1995 [empty]
Unr. Cycloneuralia Ahlrichs 1995
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Ph. Nematoda Potts 1932 [roundworm]
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G. Anguillula Muller 1786
G. †Copalonema Poinar 2011
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†Copalonema colombiana Poinar 2011
G. Enchytraeus Henle 1837
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†Enchytraeus sepultus Menge 1866
Cl. Enoplea Inglis 1983
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Subcl. Dorylaimia Inglis 1983
Subcl. Enoplia Pearse 1942
G. Ingenia Gerlach 1957
G. †Vetus Taylor 1935
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†Vetus capillaceus Menge 1866
†Vetus duisburgi Taylor 1935
†Vetus elongatus Poinar 2011
†Vetus libani Poinar 2011
†Vetus maximus Poinar 2011
†Vetus microcosmus Poinar 2011
†Vetus pristinus Menge 1866
†Vetus robustus Poinar 1977
Invalid names: Nemata [synonym]
Ph. Nematomorpha Vejdovsky 1886
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Superfm. Gordioidea Rauther 1930
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Fm. Chordodidae May 1919
G. Gordius Linnaeus 1758
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†Gordius tenuifibrosus Voigt 1938
Invalid names: Gordiacea [synonym]
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Ph. Priapulida Théel 1906 [priapulid worm]
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G. †Acosmia Chen and Zhou 1997
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†Acosmia maotiania Chen and Zhou 1997
Unr. †Archaeopriapulida Conway Morris 1977
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Fm. †Fieldiidae Conway Morris 1977
G. †Lecythioscopa Conway Morris 1977
Fm. †Miskoidae Walcott 1911
Fm. †Ottoidae Walcott 1911
G. †Scolecofurca Conway Morris 1977
Fm. †Selkirkiidae Conway Morris 1977
G. †Singuuriqia Peel 2017
Invalid names: Ancalagonidae Conway Morris 1977 [empty]
G. †Archotuba Hou et al. 1999
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†Archotuba conoidalis Hou et al. 1999
G. †Corynetis Luo and Hu 1999
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†Corynetis brevis Luo and Hu 1999
†Corynetis fortis Hu et al. 2012
Invalid names: Anningvermis Huang et al. 2004 [synonym]
G. †Eximipriapulus Ma et al. 2014
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†Eximipriapulus globocaudatus Ma et al. 2014
G. †Omnidens Hou and Others 2006
Unr. †Palaeoscolecidomorpha Shi et al. 2022
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G. †Maotianshania Sun and Huo 1987
G. †Markuelia Valkov 1983
Cl. †Palaeoscolecida Conway-Morris and Robison 1986
G. †Paraselkirkia Luo and Hu 1999
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†Paraselkirkia jinningensis Hou et al. 1999
Fm. Priapulidae Gosse 1855
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G. Acanthopriapulus van der Land 1970
G. †Priapulites Schram 1973
G. Priapulopsis Koren and Danielssen 1875
G. Priapulus Lamarck 1816
G. †Xiaoheiqingella Hu 2002
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Invalid names: Yunnanpriapulus Huang et al. 2004 [synonym]
Or. †Priapulomorpha Slavini-Plawen 1974
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G. †Ancalagon Conway Morris 1977
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†Sicyophorus rara Luo and Hu 1999
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Invalid names: Protopriapulites haikouensis Hou et al. 1999 [synonym]
Invalid names: Palaeopriapulites Hou et al. 1999 [synonym], Protopriapulites Hou et al. 1999 [synonym]
Fm. Tubiluchidae Van der Land 1970
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G. †Paratubiluchus Han et al. 2004
Invalid names: Halicryptidae Salvini-Plawen 1974 [empty], Maccabeidae Por and Bromley 1974 [empty], Xiaoheiqingidae Chen 2002 [empty]
Invalid names: Priapula [empty]
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Ph. Arthropoda Latreille 1829 [arthropod]
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G. †Actaeus
G. †Almenia Hou and Bergström 1997
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†Almenia spinosa Hou and Bergström 1997
Or. †Bradoriida Raymond 1935
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Fm. †Beyrichonidae Ulrich and Bassler 1931
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G. †Aluta Mathew 1896
G. †Beyrichona Mathew 1886
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Invalid names: Escasona Mathew 1902 [synonym]
G. †Eremos Moberg and Segerberg 1906
G. †Sellula Wiman 1903
Fm. †Bradoriidae Mathew 1902
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G. †Bradoria Mathew 1899
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Invalid names: Bradorona Mathew 1903 [synonym]
G. †Walcottella Ulrich and Bassler 1931
Fm. †Cambriidae Li 1975
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G. †Petrianna Siveter 1986
Fm. †Comptalutidae Opik 1968
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G. †Alutella
G. †Haoia Shu 1990
Fm. †Hipponicharionidae Sylvester-Bradley 1961
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G. †Albrunnicola Martinsson 1979
G. †Bicarinellata Stigall 2008
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Invalid names: Bicarinella Rode et al. 2003 [replaced]
G. †Cambria Netskaya and Ivanova 1956
G. †Hipponicharion Mathew 1886
G. †Polyphyma Groom 1902
G. †Pseudobeyrichona Cui 1990
G. †Quadricona Topper et al. 2011
G. †Voibokalina Khazanovich et al. 1984
G. †Sinskolutella Melnikova 1998
G. †Spinospitella Skovsted et al. 2006
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†Spinospitella coronata Skovsted et al. 2006
Fm. †Svealutidae Opik 1968
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G. †Liangshanella Huo 1956
G. †Yakutingella Melnikova 1998
Invalid names: Archaeocopida Sylvester-Bradley 1961 [synonym], Bradoriid Hu 2005 [synonym]
G. †Burgessia
G. †Clypecaris Hou 1997
G. †Cryptocaris Barrande 1872
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†Cryptocaris bohemica Barrande 1872
†Cryptocaris contracta Barrande 1872
†Cryptocaris obsoleta Barrande 1872
†Cryptocaris pulchra Barrande 1872
†Cryptocaris rhomboidea Barrande 1872
†Cryptocaris solida Barrande 1872
†Cryptocaris tardissima Barrande 1872
G. †Dongshanocaris Hou and Bergström 1998
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†Dongshanocaris foliiformis Hou and Bergström 1998
G. †Emeiella
G. †Ercaia Chen et al. 2001
G. †Esmeraldacaris Waggoner 2003
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†Esmeraldacaris richardsonae Waggoner 2003
Subcl. †Euthycarcinoidea Gall and Grauvogel 1964
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Or. †Euthycarcinida Gall and Grauvogel 1964
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Fm. †Euthycarcinidae Handlirsch 1914
Fm. †Kottixerxidae Starobogatov 1988
G. †Forfexicaris Hou 1999
G. †Goticaris
Fm. †Guangweicarididae Luo et al. 2007
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G. †Guangweicaris Luo et al. 2007
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†Guangweicaris spinatus Luo et al. 2007
G. †Habelia
G. †Hanburia Walcott 1916
G. †Kablikia
G. †Kleptothule Budd 1995
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G. †Liantuoia
G. †Longquania Luo et al. 2008
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†Longquania bispinosa Luo et al. 2008
Cl. †Marrelomorpha Beurlen 1930
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Or. †Acercostraca Lehmann 1955
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Fm. †Vachonisiidae Tasch 1969
Invalid names: Mimetasterida Beurlen 1934 [empty]
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Invalid names: Emeraldellida Størmer 1944 [empty], Limulavida Walcott 1912 [empty]
G. †Molaria
G. †Motina
Cl. Myriapoda Latreille 1802
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G. †Anthracerpes Meek and Worthen 1865
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†Anthracerpes typus Meek and Worthen 1865
G. †Archicarabides Fritsch 1901
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†Archicarabides pater Fritsch 1901
Cl. Chilopoda Latreille 1817 [centipede]
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G. †Nisaukankoil Malkani 2019
G. †Phailawaghkankoil Malkani 2019
G. †Rhyniognatha Tillyard 1928
Invalid names: Anamorpha Haase 1880 [empty]
Unr. Dignatha Tiegs 1947
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Subcl. Diplopoda Gervais 1844 [millipede]
G. †Eileticus Scudder 1882
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†Eileticus aequalis Scudder 1890
†Eileticus anthracinus Scudder 1882
G. †Eojulus Kušta 1886
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†Eojulus fragilis Kušta 1886
G. †Hemiphoberia Fritsch 1899
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†Hemiphoberia alternans Fritsch 1899
G. †Heterovorhoeffia Fritsch 1899
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†Heterovorhoeffia crassa Fritsch 1899
G. †Juloites Haupt 1956
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G. †Kralupia Fritsch 1907
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†Kralupia carbonaria Fritsch 1907
G. †Palenarthrus Scudder 1890
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†Palenarthrus impressus Scudder 1890
G. †Sandtneria Fritsch 1899
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†Sandtneria gemmata Fritsch 1899
Cl. Symphyla Ryder 1880
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Or. Scolopendrellida Hoffman 1969
Fm. Scutigerellidae Bagnall 1913
Invalid names: Eoscolopendridae Scudder 1890 [invalid subgroup], Gerascutigeridae Scudder 1890 [invalid subgroup], Tiphoscorpionoidea Kjellesvig-Waering 1986 [invalid subgroup]
G. †Naptile
G. †Occacaris Hou 1999
Or. †Odaraiida Simonetta and Delle Cave 1975
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Fm. †Pectocariidae Hou 1999
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G. †Pectocaris Hou 1999
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Invalid names: Odaraia [synonym]
G. †Panlongia Liu et al. 2006
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†Panlongia spinosa Liu et al. 2006
†Panlongia tetranodusa Liu et al. 2006
G. †Parioscorpio Wendruff et al. 2020
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†Parioscorpio venator Wendruff et al. 2020
Unr. †Parvancorinomorpha Lin et al. 2006
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Fm. †Parvancorinidae Glaessner 1979
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G. †Primicaris Zhang et al. 2003
G. †Skania Walcott 1931
G. †Pauloterminus Taylor 2002
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G. †Pisinnocaris Hou and Bergström 1998
G. †Platicytes Garvey and Hasiotis 2008
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†Platicytes lioparadus Garvey and Hasiotis 2008
G. †Priscansermarinus Collins and Rudkin 1981
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†Priscansermarinus barnetti Collins and Rudkin 1981
G. †Proricaris Baily 1870
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†Proricaris machenrici Baily 1870
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†Protocaris marshi Walcott 1884
G. †Pseudoiulia Hou and Bergström 1998
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†Pseudoiulia cambriensis Hou and Bergström 1998
Cl. Pycnogonida Latreille 1810 [sea spider]
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G. †Cambropycnogon Waloszek and Dunlop 2002
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†Cambropycnogon klausmuelleri Waloszek and Dunlop 2002
G. †Flagellopantopus Poschmann and Dunlop 2006
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†Flagellopantopus blocki Poschmann and Dunlop 2006
G. †Haliestes Siveter et al. 2004
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†Haliestes dasos Siveter et al. 2004
Fm. †Palaeoisopodidae Dubinin 1957
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G. †Palaeomarachne Rudkin et al. 2013
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†Palaeomarachne granulata Rudkin et al. 2013
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Fm. †Palaeopantopodidae Hedgpeth 1955
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†Palaeopantopus maucheri Broili 1928
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†Palaeothea devonica Bergstroem et al. 1980
Or. Pantopoda Gerstaecker 1863
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Fm. Ammotheidae Dohrn 1881
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Invalid names: Ammothellidae Fry 1978 [synonym], Ephyrogymnidae Fry 1978 [synonym], Eurycididae Sars 1891 [synonym], Oorhynchidae Schimkewitsch 1913 [synonym], Paranymphonidae Fry 1978 [synonym], Sericosuridae Fry 1978 [synonym], Tanystylidae Shimkevich 1913 [synonym], Trygaeidae Fry 1978 [synonym]
Fm. Callipallenidae Hilton 1942
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Invalid names: Cheilapallenidae Fry 1978 [synonym], Clavigeropallenidae Fry 1978 [synonym], Hannonidae Fry 1978 [synonym], Metapallenidae Fry 1978 [synonym], Pallenidae Wilson 1878 [synonym], Queubidae Fry 1978 [synonym], Stylopallenidae Fry 1978 [synonym]
Fm. Colossendeidae Hoek 1881
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Invalid names: Pasithoidae Sars 1891 [synonym], Rhopalorhynchidae Fry 1978 [synonym]
Fm. Endeidae Norman 1904
Fm. Phoxichilidiidae Sars 1891
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Invalid names: Anoplodactylidae Fry 1978 [synonym], Phoxiphilyridae Fry 1978 [synonym]
Invalid names: Austrodecidae Stock 1954 [empty], Nymphonidae Wilson 1878 [empty], Pallenopsidae Fry 1978 [empty], Pycnogonidae Wilson 1878 [empty], Rhynchothoracidae Thompson 1909 [empty]
Invalid names: Arachnopoda Dana 1853 [synonym], Palaeoisopoda [empty]
G. †Pygmaclypeatus Zhang et al. 2000
G. †Rhombicalvaria Hou 1987
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G. †Rhynimonstrum Anderson and Trewin 2003
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†Rhynimonstrum dunlopi Anderson and Trewin 2003
G. †Schizodiscus Clarke 1888
Cl. Scorpionida Latreille 1817 [scorpion]
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G. †Waeringoscorpio Størmer 1970
Invalid names: Euscorpionina Petrunkevitch 1949 [empty], Neoscorpionina [empty], Palaeoscorpiones Stockwell 1989 [empty], Proscorpionidae Scudder 1885 [empty]
G. †Shankouia Waloszek et al. 2005
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†Shankouia zhenghei Chen et al. 2004
G. †Tanglangia Lou and Hu 1999
Or. †Tuzoida Simonetta and Delle Cave 1975
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Fm. †Tuzoiidae Raymond 1935
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G. †Tuzoia Walcott 1912
G. †Utahcaris Conway Morris and Robison 1988
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†Utahcaris orion Conway Morris and Robison 1988
Invalid names: Agonata Fabricius 1775 [invalid subgroup], Aptera Linnaeus 1758 [invalid subgroup], Arachnoidea Guthörl 1934 [empty], Blattina splendens Göppert 1865 [nomen dubium], Branchiata Fitzinger 1843 [empty], Devonopilio Tihelka et al. 2020 [nomen dubium], Devonopilio hutchinsoni Tihelka et al. 2020 [nomen dubium], Dinopilio parvus Petrunkevitch 1953 [nomen dubium], Euarthropoda [synonym], Geracus Matthew 1897 [nomen dubium], Geracus tubifer Matthew 1897 [nomen dubium], Gigantopleurida Guthörl 1934 [empty], Graeophonus Scudder 1890 [nomen dubium], Kampecaris obanensis Peach 1899 [nomen dubium], Kounicia Frič 1901 [nomen dubium], Libellula carbonaria Scudder 1876 [nomen dubium], Monoculus Linnaeus 1758 [nomen oblitum], Orstenocarida Müller and Walossek 1988 [empty], Paratrilobita [empty], Pedestria Scopoli 1763 [invalid subgroup], Pleurolycosa Fritsch 1904 [nomen vanum], Pleurolycosa prolifera Fritsch 1899 [nomen vanum], Poliochera pustulatus Laurentiaux-Vieira and Laurentiaux 1963 [nomen dubium], Schizoramia Bergström 1976 [synonym], Skaracarida Müller and Walossek 1985 [empty], Synistata Fabricius 1775 [invalid subgroup], Unogata Fabricius 1775 [invalid subgroup]
Or. †Luolishaniida Hou and Bergström 1995
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G. †Facivermis Hou and Chen 1989
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†Facivermis yunnanicus Hou and Chen 1989
Invalid names: Xishania Hu 2002 [replaced]
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†Luolishania longicruris Hou and Chen 1989
Invalid names: Miraluolishania Liu and Shu 2004 [synonym]
G. †Ovatiovermis Carib and Aria 2017
Fm. †Teratopodidae Caron and Aria 2020
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G. †Acinocricus Conway-Morris and Robinson 1988
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†Acinocricus stichus Conway-Morris and Robison 1988
G. †Collinsium Yang et al. 2015
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†Collinsium ciliosum Yang et al. 2015
G. †Collinsovermis Caron and Aria 2020 [Collins monster]
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†Collinsovermis monstruosus Caron and Aria 2020 [Collins' monster]
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†Onychodictyon ferox Hou et al. 1991
Ph. Onychophora Grube 1853 [velvet worm]
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G. †Antennacanthopodia Ou et al. 2011
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†Cardiodictyon catenulum Hou et al. 1991
G. †Diania Liu et al. 2011
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†Diania cactiformis Liu et al. 2011
Or. Euonychophora Hutchinson 1930
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Fm. Peripatidae Evans 1901
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G. †Cretoperipatus Engel and Grimaldi 2002
Invalid names: Ontonychophora Poinar 2000 [synonym]
Unr. †Hallucishaniids Smith and Caron 2015
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G. †Carbotubulus Haug et al. 2012
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†Carbotubulus waloszeki Haug et al. 2012
G. †Hallucigenia Conway Morris 1977
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†Hallucigenia fortis Hou and Bergström 1995
†Hallucigenia sparsa Conway Morris 1977
G. †Orstenotubulus Maas et al. 2007
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†Orstenotubulus evamuellerae Maas et al. 2007
G. †Rushtonites Hinz 1987
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†Rushtonites spinosus Hinz 1987
Invalid names: Hallucigeniidae Conway Morris 1977 [empty], Luolishaniidae Hou and Bergstrom 1995 [empty]
G. †Helenodora Thompson and Jones 1980
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†Helenodora inopinata Thompson and Jones 1980
G. †Microdictyon Bengston et al. 1981
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†Microdictyon cuneum Wotte and Sundberg 2017
†Microdictyon depressum Bengtson et al. 1990
†Microdictyon effusum Bengtson et al. 1981
†Microdictyon montezumaensis Wotte and Sundberg 2017
†Microdictyon rhomboidale Bengtson et al. 1986
†Microdictyon robisoni Bengtson et al. 1986
†Microdictyon sinicum Chen et al. 1989
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†Paucipodia haikouensis Luo and Hu 1999
G. †Xenusion Pompeckj 1927
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†Xenusion auerswaldae Pompeckj 1927
Invalid names: Udeonychophora Poinar 2000 [empty]
Fm. †Opabiniidae Walcott 1912
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G. †Opabinia Walcott 1912
G. †Utaurora Pates et al. 2022
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†Utaurora comosa Pates et al. 2022
Ph. Tardigrada Doyère 1840 [water bear]
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Cl. Eutardigrada Richters 1926
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Or. Apochela Schuster et al. 1980
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Fm. Milnesiidae Ramazzotti 1962
Or. Parachela Schuster et al. 1980
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Fm. †Beornidae Cooper 1964
Superfm. Isohypsibioidea Marley et al. 2011
Invalid names: Heterotardigrada Marcus 1927 [empty]
Cl. †Xenusia Dzik and Krumbiegel 1989
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Or. †Scleronychophora Huo and Bergström 1995
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Invalid names: Eoconchariidae Hou and Shu 1987 [empty]
†Xenusia auerswaldae Pompeckj 1927
Or. †Xenusiida Dzik and Krumbiegel 1989
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Invalid names: Xenusiidae Dzik and Krumbiegel 1989 [empty]
Invalid names: Lobopodia [invalid subgroup]
Unr. †Scalidophora
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Ph. Chaetognatha Leuckart 1854 [arrow worm]
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Fm. †Amiskwidae Walcott 1911
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G. †Amiskwia Walcott 1911
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†Amiskwia sagittiformis Walcott 1911
G. †Eognathacantha Chen and Huang 2002
G. †Paucijaculum Schram 1973
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†Paucijaculum samamithion Schram 1973
Cl. †Protoconodonta Landing 1995
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G. †Mongolodus Missarzhevsky 1977
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†Mongolodus maximi Gravestock et al. 2001
†Mongolodus sulcus Parkhaev and Demidenko 2010
Invalid names: Maldeotaia Singh and Shukla 1981 [synonym]
G. †Protohertzina Missarzhevsky 1973
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†Protohertzina anabarica Missarzhevsky 1973
†Protohertzina cultrata Missarzhevsky 1977
†Protohertzina dabashanensis Yang and He 1984
†Protohertzina robusta Qian 1977
†Protohertzina siciformis Missarzhevsky 1973
†Protohertzina unguliformis Missarzhevsky 1973
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Invalid names: Emeidus primitivus Chen 1982 [synonym], Hastina bialata Yang and He 1984 [synonym]
Invalid names: Emeidus Chen 1982 [synonym], Hastina Yang and He 1984 [synonym]
Or. †Protoconodontida Landing 1995
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†Ganloudina longispina Yang and He 1984
†Ganloudina platybasala Yang and He 1984
G. †Glauderia Poulsen 1967
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†Glauderia multifidus Missarzhevsky and Mambetov 1981
G. †Kijacus Missarzhevsky and Mambetov 1981
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†Kijacus kijanicus Missarzhevsky 1966
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Invalid names: Leguminella He 1984 [synonym]
G. †Titerina
G. †Eokinorhynchus Zhang et al. 2015
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†Eokinorhynchus rarus Zhang et al. 2015
G. †Eolympia Han et al. 2010
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†Eolympia pediculata Han et al. 2010
G. †Eopriapulites Liu et al. 2014
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†Eopriapulites sphinx Liu et al. 2014
Unr. †Loricifera
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G. †Eolorica Harvey and Butterfield 2017
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†Eolorica deadwoodensis Harvey and Butterfield 2017
Invalid names: Kinorhyncha [empty]
Invalid names: Nematozoa Zrzavy et al. 1998 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
No ecological data are available

Age range: base of the Pridoli to the top of the Kungurian or 423.00000 to 272.30000 Ma

Collections (2 total)


Time interval Ma Country or state Original ID and collection number
Pridoli423.0 - 419.2China (Xinjiang) Sciadophyton pristinum (10931)
Sakmarian - Kungurian295.5 - 272.3USA (Kansas) Eukaryota indet. (32659)