PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Macroolithus

Reptilia - Elongatoolithidae

Taxonomy

Macroolithus was named by Zhao (1975). It is considered to be a form taxon.

It was assigned to Elongatoolithidae by Zhao (1975), Zhao (1979), Mikhailov (1991), Zhao et al. (1991), Mikhailov (1994), Mikhailov et al. (1994), Zhao (1994), Mikhailov (2000), Zelenitsky et al. (2000), Amo Sanjuán et al. (2000), Cousin (2002), Watabe (2004), Cheng et al. (2008), Liang et al. (2009), Wang et al. (2010), Wang et al. (2013), Huh et al. (2014), He et al. (2019), Simon et al. (2019) and Jin et al. (2020).

Species

M. mutabilis, M. rugustus, M. yaotunensis

Synonymy list

Year	Name and author
1975	Macroolithus Zhao p. 106
1979	Macroolithus Zhao pp. 336-337
1991	Macroolithus Mikhailov p. 221
1991	Macroolithus Zhao et al. p. 5
1994	Macroolithus Mikhailov p. 117
1994	Macroolithus Mikhailov et al. p. 103
1994	Macroolithus Zhao p. 187
2000	Macroolithus Amo Sanjuán et al. p. 8
2000	Macroolithus Mikhailov p. 569
2000	Macroolithus Zelenitsky et al. p. 130
2002	Macroolithus Cousin p. 44
2004	Macroolithus Watabe p. 17
2008	Macroolithus Cheng et al. p. 1090
2009	Macroolithus Liang et al. p. 1590
2010	Macroolithus Wang et al. p. 114
2013	Macroolithus Wang et al. p. 199
2014	Macroolithus Huh et al. p. 220
2019	Macroolithus He et al. p. 353
2019	Macroolithus Simon et al. p. 7
2020	Macroolithus Jin et al. p. 9

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida

Rank	Name	Author
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Theropoda	()
family	Elongatoolithidae
genus	Macroolithus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Macroolithus Zhao 1975

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†Macroolithus mutabilis Mikhailov 1994

†Macroolithus rugustus Young 1965

†Macroolithus yaotunensis Zhao 1975

Diagnosis

Reference	Diagnosis
K. E. Mikhailov 1994	Large elongate-ellipsoidaieggs (more than 170 mm long) with "rough" ornament on surface (a = 6-8) and typically elongatolithidmicrostructure of shell. Ratio of spongiose to mammillar layers ranging from 2:1 to 3:1.

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

solitaryo

Environment:

terrestrialo

Locomotion:

actively mobileo

Life habit:

ground dwellingo

Diet:

carnivoreo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2004-12-14 12:50:08

Modified:

2004-12-14 14:50:08

Source: o = order

Reference: Marsh 1875

Age range: base of the Late/Upper Hauterivian to the top of the Late/Upper Campanian or 136.40000 to 70.60000 Ma

Collections (24 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Hauterivian - Early/Lower Barremian	136.4 - 125.45	Spain (Aragón)	M. sp. (84212)
Late/Upper Cretaceous	100.5 - 66.0	China (Jiangxi)	Oolithes rugustus (71868)
Santonian - Campanian	86.3 - 72.1	China (Henan)	M. yaotunensis (232203)
Late/Upper Campanian	83.5 - 70.6	Mongolia (Omnogov)	M. yaotunensis (24754)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Mongolia (Dornogov)	M. mutabilis (65327)
Maastrichtian	72.1 - 66.0	China (Guangdong)	Dinosauria indet. (71852) M. yaotunensis, M. rugustus (24752 68149 68396 68397 68398 68399) Oolithes rugustus (71853 71856 71862)
Maastrichtian	72.1 - 66.0	Kazakhstan (Aqmola)	Dinosauria indet. (13349)
Maastrichtian	72.1 - 66.0	Mongolia (Omnogov)	M. rugustus (65335 165580) M. yaotunensis (168417)
Maastrichtian	72.1 - 66.0	Mongolia (Ovorhangai)	Hadrosauridae indet. (13347)
Maastrichtian	72.1 - 66.0	China (Jiangxi)	M. rugustus (224481) M. sp. (147070 177457) M. yaotunensis (49393)