PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Elongatoolithus

Reptilia - Elongatoolithidae

Taxonomy

Elongatoolithus was named by Zhao (1975). It is considered to be a form taxon. It is the type genus of Elongatoolithidae.

It was assigned to Oviraptoridae by Zelenitsky and Currie (2004); and to Elongatoolithidae by Zhao (1975), Zeng and Zhang (1979), Zhao (1979), Mikhailov (1991), Zhao et al. (1991), Mikhailov (1994), Mikhailov et al. (1994), Zhao (1994), Mikhailov (2000), Zelenitsky et al. (2000), Amo Sanjuán et al. (2000), Cousin (2002), Fang et al. (2003), Watabe (2004), Mohabey (2005), Liang et al. (2009), Wang et al. (2010), Zhao et al. (2013), Wang et al. (2013), Huh et al. (2014), Zhang et al. (2017), Simon et al. (2019) and Jin et al. (2020).

Species

E. andrewsi, E. chichengshanensis, E. chimeiensis, E. elongatus, E. excellens, E. frustrabilis, E. jianchangensis, E. laijiaensis, E. magnus, E. sigillarius, E. subtitectorius, E. taipinghuensis, E. yangjiagouensis

Synonymy list

Year	Name and author
1975	Elongatoolithus Zhao p. 108
1979	Elongatoolithus Zeng and Zhang p. 133
1979	Elongatoolithus Zhao p. 336
1991	Elongatoolithus Mikhailov p. 221
1991	Elongatoolithus Zhao et al. p. 5
1994	Elongatoolithus Mikhailov p. 111
1994	Elongatoolithus Mikhailov et al. p. 103
1994	Elongatoolithus Zhao p. 187
2000	Elongatoolithus Amo Sanjuán et al. p. 9
2000	Elongatoolithus Mikhailov p. 569
2000	Elongatoolithus Zelenitsky et al. p. 130
2002	Elongatoolithus Cousin p. 43
2003	Elongatoolithus Fang et al. p. 517
2004	Elongatoolithus Watabe p. 17
2004	Elongatoolithus Zelenitsky and Currie p. 134A
2005	Elongatoolithus Mohabey p. 473
2009	Elongatoolithus Liang et al. p. 1590
2010	Elongatoolithus Wang et al. p. 114
2013	Elongatoolithus Wang et al. p. 199
2013	Elongatoolithus Zhao et al. p. 4665
2014	Elongatoolithus Huh et al. p. 220
2017	Elongatoolithus Zhang et al. p. 191
2019	Elongatoolithus Simon et al. p. 7
2020	Elongatoolithus Jin et al. p. 9

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida

Rank	Name	Author
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Theropoda	()
family	Elongatoolithidae
genus	Elongatoolithus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Elongatoolithus Zhao 1975

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†Elongatoolithus andrewsi Zhao 1975

†Elongatoolithus chichengshanensis Fang et al. 2003

†Elongatoolithus chimeiensis Fang 2007

†Elongatoolithus elongatus Young 1954

†Elongatoolithus excellens Mikhailov 1994

†Elongatoolithus frustrabilis Mikhailov 1994

†Elongatoolithus jianchangensis Fang 2007

†Elongatoolithus laijiaensis Fang et al. 2003

†Elongatoolithus magnus Zeng and Zhang 1979

†Elongatoolithus sigillarius Mikhailov 1994

†Elongatoolithus subtitectorius Mikhailov 1994

†Elongatoolithus taipinghuensis Yu 1998

†Elongatoolithus yangjiagouensis Fang 2007

Diagnosis

Reference	Diagnosis
K. E. Mikhailov 1994	Elongated-ellipsoidal eggs of medium size (up to 170 mm long) with "fine" ornamentation on surface (a = 8–11) and typically elongatoolithid microstructure of shell. Ratio of spongiose to mammilar layer ranging from 2:1 to 3:1.

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

solitaryo

Environment:

terrestrialo

Locomotion:

actively mobileo

Life habit:

ground dwellingo

Diet:

carnivoreo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2004-12-14 12:50:08

Modified:

2004-12-14 14:50:08

Source: o = order

Reference: Marsh 1875

Age range: base of the Early/Lower Barremian to the top of the Late/Upper Campanian or 130.00000 to 70.60000 Ma

Collections (38 total)

Time interval	Ma	Country or state	Original ID and collection number
Early/Lower Barremian	130.0 - 125.45	Spain (Aragón)	Elongatoolithidae indet. (61409)
Early/Lower Albian	112.03 - 109.0	Japan (Hyogo)	E. sp. (117980)
Cenomanian - Santonian	100.5 - 83.6	China (Guangdong)	E. sp. (188428)
Late/Upper Cretaceous	100.5 - 66.0	China (Hunan)	E. sp., E. magnus (71664)
Late/Upper Cretaceous	100.5 - 66.0	China (Xinjiang)	Oolithes elongatus (97613)
Late/Upper Cenomanian - Turonian	99.6 - 89.8	China (Zhejiang)	E. chichengshanensis (204721) E. laijiaensis (204718)
Campanian	83.6 - 72.1	China (Nei Mongol)	Elongatoolithidae indet. (46904) E. sp. (205652)
Campanian	83.6 - 72.1	China (Shandong)	Oolithes elongatus (80114 80115 80117) Oolithes elongatus, Reptilia indet. (24757)
Campanian	83.6 - 72.1	Mongolia (Omnogov)	E. frustrabilis (37211)
Campanian - Maastrichtian	83.6 - 66.0	China (Xinjiang)	Oolithes elongatus (159589) Oolithes sp. (24753)
Campanian - Maastrichtian	83.6 - 66.0	China (Hunan)	Oolithes elongatus (79741)
Late/Upper Campanian	83.5 - 70.6	Mongolia (Omnogov)	Elongatoolithidae indet. (53229 65344) E. sp. (118259 127668) Protoceratops andrewsi (46152)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Mongolia (Omnogov)	E. sp. (174156)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Mongolia (Dornogov)	Elongatoolithidae indet. (45331)
Maastrichtian	72.1 - 66.0	China (Guangdong)	Dinosauria indet. (71855) E. elongatus, E. andrewsi (24752 68149 68396 68397 68398 68399) Oolithes elongatus (71854 71859 71867)
Maastrichtian	72.1 - 66.0	Mongolia (Omnogov)	Elongatoolithidae indet. (55257 65335 165580)
Maastrichtian	72.1 - 66.0	China (Jiangxi)	E. elongatus (220036)