PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Dortoka

Reptilia - Testudines

Taxonomy

Dortoka was named by Lapparent de Broin and Murelaga (1996).

It was assigned to Dortokidae by Lapparent de Broin and Murelaga (1996), de Lapparent de Broin and Murelaga (1999), Lapparent de Broin et al. (2004), Gaffney et al. (2006) and Pérez-García et al. (2012); and to Dortokidae by Cadena and Joyce (2015) and Augustin et al. (2021).

Species

D. botanica, D. vasconica, D. vremiri

Synonyms

Ronella was named by Gheerbrant et al. (1999).

It was synonymized subjectively with Dortoka by Cadena and Joyce (2015).

It was assigned to Dortokidae by Gheerbrant et al. (1999), Lapparent de Broin et al. (2004), Gaffney et al. (2006), Pérez-García (2012) and Vremir (2013).

Synonymy list

Year	Name and author
1996	Dortoka Lapparent de Broin and Murelaga
1999	Ronella Gheerbrant et al.
1999	Dortoka de Lapparent de Broin and Murelaga
2004	Dortoka Lapparent de Broin et al.
2004	Ronella Lapparent de Broin et al.
2006	Dortoka Gaffney et al. p. 29
2006	Ronella Gaffney et al. p. 29
2012	Ronella Pérez-García
2012	Dortoka Pérez-García et al. p. 151
2013	Ronella Vremir
2015	Dortoka Cadena and Joyce p. 15
2021	Dortoka Augustin et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda

Rank	Name	Author
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
—	Testudinata	(Oppel 1811)
order	Testudines	Batsch 1788
—	Panpleurodira
—	Dortokidae	()
genus	Dortoka

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Dortoka Lapparent de Broin and Murelaga 1996

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†Dortoka botanica Gheerbrant et al. 1999

†Dortoka vasconica Lapparent de Broin and Murelaga 1996

†Dortoka vremiri Augustin et al. 2021

Invalid names: Ronella Gheerbrant et al. 1999 [synonym]

Diagnosis

Reference	Diagnosis
E. Cadena and W. G. Joyce 2015	Dortoka can be diagnosed as a pan-pleurodire by the sutural articulation of the pelvis with the shell, presence of a well-developed anal notch, a single gular scute, central articulation in the cervical column and the loss of inframarginals. Dortoka is currently differentiated from other pan-pleurodires by the distinct microsculpturing of the shell consisting of a microreticulate pattern, and the distinct macrosculpturing of the carapace, particularly of the neurals, consisting of anteroposteriorly elongate pits and ridges, absence of contact between peripheral I and costal I, irregularly shaped neurals, a lapping of pleural II onto costal I and the absence of mesoplastra in some taxa.
F. J. Augustin et al. 2021	Members of the genus are characterized by the possession of the following synapomorphies: (1) distinct microsculpturing of the shell, consisting of a micro-reticulate pattern; (2) distinct macrosculpturing of the neurals, consisting of anteroposteriorly elongate pits and ridges; (3) absence of contact between the first peripheral and the first costal; (4) irregularly shaped neurals; (5) the second pleural overlaps the first costal; and (6) absence of mesoplastra

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

coastal, freshwaterg

Locomotion:

actively mobilec

Comments:

From Cadena and Joyce (2015): All known dortokids originate from littoral or riverine sediments and are found associated with continental faunas (Lapparent de Broin and Murelaga 1996, 1999; Lapparent de Broin et al. 2004; Pérez-García et al. 2014). The shell histology and bone density of D. vasconica is comparable with that of freshwater forms (Pérez-García et al. 2012).g

Created:

2017-04-29 07:44:12

Modified:

2017-04-29 07:44:12

Source: g = genus, c = class, subp = subphylum

References: Cadena and Joyce 2015, Carroll 1988, Hendy et al. 2009

Age range: base of the Late/Upper Campanian to the top of the Lutetian or 83.50000 to 41.30000 Ma

Collections (18 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Campanian	83.5 - 70.6	Spain (Castilla y Leon)	Chelonia indet. (48861) D. vasconica (28406 28765)
Middle Campanian	83.5 - 70.6	France (Provence-Alpes-Côte d'Azur)	D. sp. (48849)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	France (Languedoc-Roussillon)	D. sp. (74660)
Late/Upper Campanian - Early/Lower Maastrichtian	83.5 - 66.0	Spain (Valencia)	D. sp. (53918)
Late/Upper Campanian - Early/Lower Maastrichtian	83.5 - 66.0	France (Languedoc-Roussillon)	D. sp. (59687) D. vasconica (48870)
Late/Upper Campanian - Late/Upper Maastrichtian	83.5 - 66.0	France (Provence-Alpes-Cote d'Azur)	D. sp. (59778)
Early/Lower Maastrichtian	70.6 - 66.0	Romania (Hunedoara)	D. vremiri (221768)
Thanetian	59.2 - 56.0	Romania	Ronella botanica (83307 138239 138240 138242 138243 138244 138245)
Ypresian - Lutetian	56.0 - 41.3	Romania	Ronella sp. (178481)