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Cupedidae (reticulated beetle)
Taxonomy
Cupedidae was named by Laporte (1836). It is extant. Its type is Cupes.
It was corrected as Cupidae by Lin (1982); it was misspelled as Cupesidae by Hong and Wang (1987), Carpenter (1992), Lawrence and Newton (1995), Kirejtshuk et al. (2010), Bouchard et al. (2011), Kirejtshuk (2020) and Cai et al. (2022).
It was assigned to Coleoptera by Lin (1976) and Lin (1982); to Cupesoidea by Hong and Wang (1987); to Archostemata by Carpenter (1992), Lawrence and Newton (1995), Kirejtshuk et al. (2010) and Bouchard et al. (2011); and to Cupedoidea by Crowson (1981), Kirejtshuk (2020) and Cai et al. (2022).
It was corrected as Cupidae by Lin (1982); it was misspelled as Cupesidae by Hong and Wang (1987), Carpenter (1992), Lawrence and Newton (1995), Kirejtshuk et al. (2010), Bouchard et al. (2011), Kirejtshuk (2020) and Cai et al. (2022).
It was assigned to Coleoptera by Lin (1976) and Lin (1982); to Cupesoidea by Hong and Wang (1987); to Archostemata by Carpenter (1992), Lawrence and Newton (1995), Kirejtshuk et al. (2010) and Bouchard et al. (2011); and to Cupedoidea by Crowson (1981), Kirejtshuk (2020) and Cai et al. (2022).
Subtaxa
Synonymy list
| Year | Name and author |
|---|---|
| 1836 | Cupedidae Laporte |
| 1976 | Cupidae Lin p. 112 |
| 1981 | Cupedidae Crowson p. 694 |
| 1982 | Cupidae Lin p. 331 |
| 1987 | Cupesidae Hong and Wang p. 259 |
| 1992 | Cupedidae Carpenter |
| 1995 | Cupedidae Lawrence and Newton, Jr. |
| 2010 | Cupedidae Kirejtshuk et al. |
| 2011 | Cupedidae Bouchard et al. p. 95 |
| 2020 | Cupedidae Kirejtshuk p. 21 |
| 2022 | Cupedidae Cai et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Cupedidae Laporte 1836 [reticulated beetle]
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Subfm. Cupedinae Lacordaire 1857
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G. †Anaglyphites Ponomarenko 1964
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†Anaglyphites admotus Ponomarenko 1964
†Anaglyphites capitatus Ponomarenko 1966
†Anaglyphites clavatus Ponomarenko 1964
†Anaglyphites crassipygus Ponomarenko 2014
†Anaglyphites laximandibule Lin 1980
†Anaglyphites minimus Ponomarenko 1997
†Anaglyphites minutissimus Ponomarenko 2014
†Anaglyphites mongolicus Ponomarenko 1997
G. †Apriacma Kirejtshuk et al. 2016
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†Apriacma clavata Tan et al. 2006
†Apriacma latidentata Tan et al. 2006
†Apriacma renaria Tan et al. 2006
†Apriacma tuberculosa Tan et al. 2006
G. †Cainomerga Kirejtshuk et al. 2016
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†Cainomerga brevicornis Kirejtshuk et al. 2016
†Cainomerga fraterna Kirejtshuk et al. 2016
†Cainomerga immaculata Piton 1940
†Cainomerga palaeocenica Kirejtshuk et al. 2016
†Cainomerga ponti Kirejtshuk et al. 2016
G. Cupes Fabricius 1801
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†Cupes balticus Yamamoto 2024
†Cupes distinctissimus Kirejtshuk et al. 2016
†Cupes eckfeldensis Tröster 1993
†Cupes golovatchi Kirejtshuk 2020
†Cupes groehni Kirejtshuk 2005
†Cupes hoffeinsorum Kirejtshuk 2005
†Cupes kerneggeri Kirejtshuk 2005
†Cupes komissari Kirejtshuk 2005
†Cupes legalovi Kirejtshuk 2020
†Cupes longus Hong and Wang 1987
†Cupes lutzi Kirejtshuk 2020
†Cupes manifestus Kirejtshuk et al. 2010
†Cupes messelensis Tröster 1993
†Cupes nabozhenkoi Kirejtshuk 2020
†Cupes ponomarenkoi Kirejtshuk et al. 2010
†Cupes praeglacialis Gersdorf 1976
†Cupes rohdendorfi Yablokov-Khnzorian 1960
†Cupes simillimus Kirejtshuk et al. 2016
†Cupes tesselatus Motschulsky 1856
†Cupes tessellatus Motschulsky 1856
†Cupes wedmannae Kirejtshuk 2020
†Cupes weitschati Kirejtshuk 2005
Invalid names: Cupoides Motschulsky 1856 [synonym], Distocupes Neboiss 1984 [synonym], Tenomerga Neboiss 1984 [synonym]
G. †Gracilicupes Tan et al. 2006
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†Gracilicupes crassicruralis Tan et al. 2006
†Gracilicupes minimus Strelnikova 2019
†Gracilicupes tenuicruralis Tan et al. 2006
G. †Kirghizocupes Ponomarenko 1966
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†Kirghizocupes cellulosus Ponomarenko 1966
†Kirghizocupes indistinctus Ponomarenko 1969
†Kirghizocupes proporeius Ponomarenko 1969
Invalid names: Mesocupoides Ponomarenko 1969 [synonym]
G. †Latocupes Ren and Tan 2006
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†Latocupes angustilabialis Tan et al. 2007
†Latocupes bellus Ren and Tan 2006
†Latocupes collaris Tan et al. 2007
†Latocupes fortis Ren and Tan 2006
†Latocupes jiensis Ren 1995
†Latocupes latilabialis Tan et al. 2007
Invalid names: Pulchicupes Ren 1995 [synonym]
G. †Mallecupes Jarzembowski et al. 2017
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Subg. †Mallecupes (Mallecupes) Jarzembowski et al. 2017
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†Mallecupes (Mallecupes) qingqingae Jarzembowski et al. 2017
Subg. †Mallecupes (Mallecupoides) Jarzembowski et al. 2017
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†Mallecupes (Mallecupoides) cleevelyi Jarzembowski et al. 2017
†Mallecupes cheni Wang et al. 2023
†Mallecupes prokini Kirejtshuk 2020
†Mallecupes zhangi Song et al. 2022
G. †Menatops Kirejtshuk et al. 2016
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†Menatops bartenevi Kirejtshuk et al. 2016
†Menatops orbiculatus Kirejtshuk et al. 2010
G. †Mesocupes Martynov 1926
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†Mesocupes bidens Ponomarenko 1964
†Mesocupes minor Ponomarenko 1968
†Mesocupes paulus Ponomarenko 1964
†Mesocupes primitivus Martynov 1926
†Mesocupes spinosus Ponomarenko 1964
†Mesocupes zherikhini Soriano and Delclos 2006
Invalid names: Mesocupes (Mesocupes) Martynov 1926 [invalid subgroup]
G. †Pterocupes Ponomarenko 1966
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†Pterocupes antennatus Ponomarenko 1966
†Pterocupes leptocerus Ponomarenko 1966
Invalid names: Anaglyphites pluricavus Soriano and Delclos 2006 [nomen dubium], Cupedini Crowson 1962 [empty], Platycupes sogdianus Ponomarenko 1966 [nomen dubium], Priacma sanzii Soriano and Delclos 2006 [nomen dubium]
Invalid names: Mesocupedinae Ponomarenko 1969 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Kiessling 2004, Bush and Bambach 2015 | |||||
Age range
Maximum range based only on fossils: base of the Ladinian to the top of the Piacenzian or 242.00000 to 2.58800 Ma
Minimum age of oldest fossil (stem group age): 237.0 Ma
Minimum age of oldest fossil (stem group age): 237.0 Ma
Collections (73 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Ladinian | Kyrgyzstan (Osh) | Mesocupoides proporeius, Mesocupoides indistinctus (110763) Mesocupoides proporeius, Pterocupes antennatus, Pterocupes leptocerus, Kirghizocupes cellulosus (106090) Platycupes sogdianus, Asimma rara (113468) | |
| Hettangian | USA (New Jersey) | Coleoptera indet. (153635) | |
| Bajocian - Bathonian | China (Anhui) | Cupedidae indet. (106695) | |
| Callovian - Oxfordian | China (Nei Mongol) | Gracilicupes crassicruralis, Gracilicupes tenuicruralis (113376) Mesocupes angustilabialis, Mesocupes latilabialis, Mesocupes collaris (113371) Mesocupes collaris (113360) | |
| Callovian - Oxfordian | Kazakhstan (Chimkent) | Anaglyphites admotus, Mesocupes bidens (114238) Anaglyphites clavatus, Anaglyphites admotus, Mesocupes spinosus, Mesocupes primitivus (105971) Anaglyphites paulus (31311) Cupidium abavum (106102) Mesocupes minor (106101) Mesocupes primitivus (113402 115792) | |
| Tithonian | Mongolia | Anaglyphites crassipygus (121393) Anaglyphites minutissimus (104748 121394) | |
| Late/Upper Hauterivian | United Kingdom (England) | Cupedidae indet. (119697) | |
| Hauterivian | Russian Federation (Chita) | Cupedidae indet. (125409) | |
| Early/Lower Barremian | Spain | Anaglyphites pluricavus (123437) | |
| Late/Upper Barremian | Spain (Castilla-La Mancha) | Priacma sanzii, Anaglyphites zherikhini (28006) | |
| Late/Upper Barremian | China (Liaoning) | Latocupes fortis, Latocupes bellus, Priacma latidentata, Priacma tuberculosa (111717) Priacma clavata, Priacma renaria (111710) Priacmopsis subtilis, Furcicupes raucus (119055) | |
| Aptian | China (Jilin) | Ensicupes obtusus (151876) | |
| Aptian | China (Nei Mongol) | Ensicupes guyangensis (156005) | |
| Aptian | Russian Federation (Buryatia) | Cupedidae indet., Anaglyphites capitatus, Priacmopsis adumbrata (105536) Gracilicupes minimus (113350) | |
| Aptian | China (Zhejiang) | Anaglyphites laximandibule (155596) | |
| Aptian | Mongolia | Anaglyphites mongolicus (113359 124342) Anaglyphites mongolicus, Priacmopsis minimus (113355) Priacmopsis minimus (113356) | |
| Late/Upper Aptian | Brazil (Ceará) | Cupedidae indet. (122557) | |
| Late/Upper Aptian | China (Beijing Shi) | Pulchicupes jiensis (106965) | |
| Cenomanian | Canada (Newfoundland and Labrador) | Cupedidae indet. (119438) | |
| Early/Lower Cenomanian | Myanmar | Mallecupes prokini (172900) Mallecupes qingqingae (167760) Mallecupes qingqingae, Barbaticupes combertiae, Mallecupes cleevelyi, Priacma megapuncta (161726) Mallecupes zhangi, Mallecupes cheni (224419) | |
| Early/Lower Cenomanian | France | Cupedidae indet. (123019) | |
| Turonian | USA (New Jersey) | Paracupes svitkoi (124293) | |
| Late/Upper Coniacian | Russian Federation (Khabarovsk) | Cupedinae indet. (123528 138054) | |
| Selandian | France (Auvergne) | Cupedinae indet. 1, Mesocupes brevicornis (123436) Cupedinae indet. 2, Cupedinae indet. 3, Cupes manifestus, Mesocupes sp. 1, Mesocupes spp., Mesocupes fraternus, Menatops bartenevi, Cupes distinctissimus, Cupes simillimus (188751) Cupedinae indet. 4, Cupes manifestus, Mesocupes spp., Mesocupes sp. 2, Mesocupes sp. 3, Mesocupes palaeocenicus, Mesocupes ponti (120579) Cupes manifestus, Cupes orbiculatus (123463) Zonabris immaculatus (113685) | |
| Ypresian | France (Oise) | Cupes ponomarenkoi (123850) | |
| MP 11 | Germany (Hessen) | Cupes nabozhenkoi, Cupes wedmannae (207885) Tenomerga messelense (157394) | |
| MP 13 | Germany (Eifel) | Cupedidae indet. 1, Cupedidae indet. 2, Tenomerga eckfeldense (138912) Cupedidae indet. 2 (135644) Cupes golovatchi, Cupes legalovi, Cupes lutzi (24201) Tenomerga eckfeldense (135643) | |
| Priabonian | United Kingdom (England) | Cupedinae indet. (113829) | |
| Priabonian | Poland | Cupoides tessellatus (120811) | |
| Priabonian | Ukraine | Cupes groehni (153587) | |
| Priabonian | Lithuania | Cupes rohdendorfi (113470) | |
| Priabonian | Russian Federation | Cupes balticus (234391) Cupes groehni, Cupes hoffeinsorum, Cupes rohdendorfi, Cupes tesselatus (123419) | |
| Priabonian | Russian Federation (Kaliningrad) | Cupes groehni, Cupes rohdendorfi (123770) Cupes groehni, Cupes rohdendorfi, Cupes tesselatus, Cupes weitschati (110735) Cupes kerneggeri, Cupes rohdendorfi (124201) Cupes motschulskyi (120610) Cupes motschulskyi, Cupes tesselatus, Cupes komissari (123420) Priacma tessellata (152640) | |
| Chattian | France (Bouches-Du-Rhone) | Cainocups aixensis (127415) | |
| Burdigalian | China (Shandong) | Cupes longus (128550) | |
| MN 4 | Czech Republic | Miocupes rihai (105501) | |
| Piacenzian | Germany (Niedersachsen) | Cupes praeglacialis (109196) |