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Staphyliniformia

Insecta - Coleoptera

Synonyms
Synonymy list
YearName and author
1900Staphyliniformia Lameere
1960Scarabaeiformia Crowson
1981Staphyliniformia Crowson p. 694
1981Scarabaeiformia Crowson p. 695
1995Scarabaeiformia Lawrence and Newton, Jr.
1995Staphyliniformia Lawrence and Newton, Jr.
2011Scarabaeiformia Bouchard et al.
2011Staphyliniformia Bouchard et al. p. 153
2022Staphyliniformia Cai et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
Dicondylia
Paranotalia
subclassPterygota()
RankNameAuthor
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
Holometabola
Aparaglossata
Neuropteriformia
Coleopterida
orderColeoptera
Mesocoleoptera
Metacoleoptera
suborderPolyphagaEmery 1886
infraorderStaphyliniformia
infraorderStaphyliniformia

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Infraor. Staphyliniformia Lameere 1900
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Superfm. Histeroidea Gyllenhal 1808
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Fm. †Cretohisteridae Zhou et al. 2018
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G. †Cretohister Zhou et al. 2018
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Cretohister sinensis Zhou et al. 2018
Fm. Histeridae Gyllenhal 1808 [clown beetle]
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Subfm. Abraeinae MacLeay 1819
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Tr. Acritini Wenzel 1944
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G. Acritus LeConte 1853
Tr. †Pantostictini Zhou et al. 2020
+
G. †Pantostictus Poinar and Brown 2009
G. †Amplectister Caterino and Maddison 2018
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Amplectister tenax Caterino and Maddison 2018
Amplectister terapoides Yamamoto and Caterino 2022
Subfm. †Antigracilinae Zhou et al. 2020
+
G. †Antigracilus Zhou et al. 2020
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Antigracilus costatus Zhou et al. 2020
G. Chaetabraeus Portevin 1929
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Chaetabraeus globulus Creutzer 1799
Subfm. Dendrophilinae Reitter 1909
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Tr. Anapleini Olexa 1982
+
G. Anapleus Horn 1873
G. †Cretanapleus Simon Pražák and Lackner 2023
Tr. Bacaniini Kryzhanovskij 1976
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G. Bacanius LeConte 1853
G. †Druantia Caterino 2021
+
Druantia aeterna Caterino 2021
G. †Olexum Simon Pražák and Lackner 2023
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Olexum complanatum Simon Pražák and Lackner 2023
Tr. Paromalini Reitter 1909
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G. Carcinops Marseul 1855
G. Xestipyge Marseul 1862
G. †Yethiha Caterino 2021
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Yethiha peregrina Caterino 2021
Yethiha pubescens Simon Pražák and Lackner 2023
Subfm. Haeteriinae Marseul 1857
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G. †Promyrmister Zhou et al. 2019
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Promyrmister kistneri Zhou et al. 2019
Subfm. Histerinae Gyllenhal 1808
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Tr. Histerini Gyllenhal 1808
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G. Hister Linnaeus 1758
G. †Platycretus Simon Pražák and Lackner 2023
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Platycretus muscularis Simon Pražák and Lackner 2023
Subfm. Onthophilinae MacLeay 1819
+
G. †Carinumerus Caterino 2021
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Carinumerus intricatus Caterino and Yamamoto 2023
Carinumerus maddisoni Caterino 2021
Carinumerus yingae Jiang et al. 2020
G. †Cretonthophilus Caterino et al. 2015
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Cretonthophilus tuberculatus Caterino et al. 2015
G. Onthophilus Leach 1817
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Onthophilus intermedius Handschin 1944
Onthophilus striatus Forster 1771
G. †Phasmister Caterino 2021
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Phasmister cristatus Caterino 2021
Phasmister hkamticus Caterino and Yamamoto 2023
Phasmister parallelus Caterino and Yamamoto 2023
G. Plegaderus Erichson 1834
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Plegaderus pitoni Théobald 1935
Subfm. Saprininae Blanchard 1845
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Tr. Myrmetini Portevin 1929
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G. Gnathoncus Jacquelin-Duval 1858
Subfm. Trypanaeinae Marseul 1857
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G. Trypanaeus Eschscholtz 1829
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Trypanaeus hispaniolus Chatzimanolis et al. 2006
Invalid names: Chlamydopsinae Bickhardt 1914 [empty], Hetaeriinae Marseul 1857 [empty], Niponiinae Fowler 1912 [empty], Tribalinae Bickhardt 1914 [empty], Trypeticinae Bickhardt 1914 [empty]
Fm. Synteliidae Lewis 1882
+
G. Syntelia Westwood 1864
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Syntelia sunwukong Jiang and Wang 2021
Invalid names: Sphaeritidae Shuckard 1839 [empty]
Superfm. Hydrophiloidea Latreille 1802 [water scavenger beetle]
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G. †Cretotaenia Ponomarenko 1977
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Cretotaenia pallida Ponomarenko 1977
Fm. Helophoridae Leach 1815
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G. Helophorus Fabricius 1775
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+
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Invalid names: Helophorus (Gephelophorus) lomnicki d'Orchymont 1927 [synonym], Helophorus (Trichelophorus) pleistocenicus Lomnicki 1894 [synonym]
Subg. Helophorus (Meghelophorus) Kuwert 1886
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Invalid names: Helophorus (Praehelephorus) dzieduszyckii Lomnicki 1894 [synonym]
Invalid names: Helophorus (Praehelephorus) Lomnicki 1894 [synonym]
Subg. †Helophorus (Mesohelophorus) Ponomarenko 1977
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Invalid names: Mesohelophorus sibiricus Ponomarenko 1977 [replaced]
Subg. †Helophorus (Mesosperchus) Ponomarenko 1977
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Helophorus (Mesosperchus) gracilis Prokin et al. 2010
Helophorus (Mesosperchus) inceptivus Fikáček et al. 2012
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Invalid names: Mesohelophorus elongatus Ponomarenko 1990 [synonym]
Helophorus (Mesosperchus) yixianus Fikáček et al. 2012
Subg. Helophorus (Orphelophorus) d'Orchymont 1927
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+
Invalid names: Helophorus (Orphelophorus) wandereri d'Orchymont 1927 [synonym]
Subg. Helophorus (Trichelophorus) Kuwert 1886
Helophorus auricollis Eschscholtz 1822
Helophorus browni McCorkle 1973
Helophorus eclectus d'Orchymont 1945
Helophorus lacustris Le Conte 1850
Helophorus nubilus Fabricius 1777
Helophorus oblongus Le Conte 1850
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Invalid names: Helophorus (Rhopalhelophorus) kuwerti Lomnicki 1894 [synonym]
Helophorus orientalis Motschulsky 1860
Helophorus praenanus Lomnicki 1894
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Invalid names: Helophorus polonicus Lomnicki 1894 [synonym]
Helophorus splendidus Sahlberg 1880
Helophorus tuberculatus Gyllenhal 1808
Fm. Hydrochidae Thomson 1859
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G. †Ponohydrochus Prokin and Strelnikova 2021
+
Ponohydrochus buryaticus Prokin and Strelnikova 2021
Fm. Hydrophilidae Latreille 1802 [water scavenger beetle]
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Subfm. Acidocerinae Zaitzev 1908
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G. Helochares Mulsant 1846
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Helochares fog Arriaga-Varela et al. 2021
Helochares mydaleus Zhang 1989
G. †Hydroyixia Fikáček et al. 2014
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Hydroyixia elongata Fikáček et al. 2014
Hydroyixia latissima Fikáček et al. 2014
Invalid names: Horelophopsinae Hansen 1997 [synonym]
G. †Alegorius Fikáček et al. 2014
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Alegorius yixianus Fikáček et al. 2014
G. †Aposphinctus Bode 1953
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+
Invalid names: Spalacoides simplex Bode 1953 [synonym]
Aposphinctus sibiricus Ponomarenko 1990
Aposphinctus striatus Bode 1953
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Invalid names: Hexameristus inflatus Bode 1953 [synonym]
Invalid names: Hexameristus Bode 1953 [synonym], Spalacoides Bode 1953 [synonym], Streblocardioides Bode 1953 [synonym]
Subfm. Chaetarthriinae Bedel 1881
+
Tr. Anacaenini Hansen 1991
+
G. Anacaena Thomson 1859
G. Crenitis Bedel 1881
G. †Cretocrenis Fikáček et al. 2017
Invalid names: Horelophinae Hansen 1991 [synonym]
Tr. Chaetarthriini Bedel 1881
+
G. Chaetarthria Stephens 1835
G. †Creniphilites Wickham 1913
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Creniphilites orpheus Wickham 1913
G. †Cretosperchus Ponomarenko 1997
+
Cretosperchus medievalis Ponomarenko 1997
Subfm. Enochrinae Short and Fikáček 2013
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G. Cymbiodyta Bedel 1881
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Cymbiodyta exstincta Scudder 1900
Cymbiodyta samueli Fikáček et al. 2023
G. Enochrus Thomson 1859
+
Enochrus innovatus Theobald 1937
Enochrus lapsus Zhang 1989
Enochrus pleistocenicus Lomnicki 1894
Enochrus primaevus Scudder 1876
Enochrus scudderi Wickham 1909
Enochrus striatus Theobald 1937
Invalid names: Philydrus Solier 1834 [replaced]
Subfm. Georissinae Laporte 1819
+
G. †Georissites Ponomarenko 2008
+
Georissites magnus Haupt 1956
Georissites negev Ponomarenko 2008
G. Georissus Latreille 1809
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G. Hydrocharis Berthold 1827
Subfm. Hydrochinae Thomson 1859
+
G. Hydrochus Germar 1824
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Hydrochus amictus Scudder 1890
Hydrochus elongatus Schaller 1783
Hydrochus relictus Scudder 1890
Subfm. Hydrophilinae Latreille 1802
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Tr. Berosini Mulsant 1844
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G. Berosus Leach 1817
Tr. Hydrobiusini Mulsant 1844
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G. †Baissalarva Fikáček et al. 2014
G. †Cretoxenus Fikáček et al. 2014
G. Hydrobius Linnaeus 1758
G. Limnoxenus Motshulsky 1853
Invalid names: Sperchopsini Hansen 1991 [synonym]
Tr. Hydrophilini Latreille 1802
+
Invalid names: Hydrophilina Latreille 1802 [invalid subgroup], Hydrophilopsis Heer 1862 [nomen dubium], Hydrophilopsis elongata Heer 1862 [nomen dubium], Hydrous ovalis Heer 1862 [nomen dubium]
Tr. Laccobiini Houlbert 1922
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G. Laccobius Erichson 1837
G. Paracymus Thomson 1867
Invalid names: Amphiopini Kuwert 1890 [empty], Cymbiodyta austerus Statz 1939 [nomen dubium]
G. †Orphnospercheus Lin 1992
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G. †Paraspercheus Ponomarenko 1977
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Paraspercheus asiaticus Ponomarenko 1977
Paraspercheus vitimensis Ponomarenko 1977
G. †Penitus Zhang et al. 1994
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Penitus gemellus Zhang et al. 1994
G. Philhydrus Solier 1834
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Philhydrus scudderi Wickham 1909
G. †Prospercheus Prokin 2009
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Prospercheus cristatus Prokin 2009
G. †Protochares Fikáček et al. 2014
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Protochares brevipalpis Fikáček et al. 2014
G. Regimbartia Zaitzev 1908
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Regimbartia attenuata Fabricius 1801
Subfm. Rygmodinae Orchymont 1916
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Invalid names: Andotypini Hansen 1991 [synonym], Borborophorini Hansen 1991 [synonym], Tormissini Hansen 1991 [synonym]
Subfm. Sphaeridiinae Latreille 1802
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Tr. Coelostomatini Heyden 1891
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G. Coelostoma Brullé 1835
Tr. Megasternini Mulsant 1844
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G. Cercyon Leach 1817
G. Cryptopleurum Mulsant 1844
G. Megasternum Mulsant 1844
Tr. Sphaeridiini Latreille 1802
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G. Sphaeridium Fabricius 1775
Invalid names: Omicrini Smetana 1975 [empty], Protosternini Hansen 1991 [empty]
G. Sternolophus Solier 1834
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Sternolophus rufipes Fabricius 1792
Invalid names: Escheria Heer 1847 [nomen dubium], Escheria bella Heer 1862 [nomen dubium], Escheria ovalis Heer 1847 [nomen dubium], Escheriidae Kolbe 1880 [synonym], Mesosperchus schultzi Ponomarenko 1985 [nomen dubium], Spercheinae Erichson 1837 [empty], Sphaeridium melanarium Gistl 1831 [nomen dubium]
G. †Hydrophilopsia Ponomarenko 1987
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Hydrophilopsia longitarsalis Ponomarenko 1987
G. †Laetopsia Fikáček et al. 2012
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Laetopsia baissensis Ponomarenko 1987
Laetopsia bontsaganica Prokin 2009
Laetopsia hydraenoides Prokin et al. 2010
Laetopsia mongolica Ponomarenko 1987
Laetopsia shatrovskiyi Prokin et al. 2010
G. †Sinosperchopsis Prokin et al. 2010
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Sinosperchopsis silinae Prokin et al. 2010
Fm. Spercheidae Erichson 1837
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G. Spercheus Kugelann 1798
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Spercheus punctatus Ponomarenko 2019
Spercheus wightensis Ponomarenko 2019
Invalid names: Epimetopidae Zaitzev 1908 [empty], Georissidae Laporte 1819 [empty]
Superfm. Scarabaeoidea Latreille 1802 [scarab beetle]
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G. †Aliscarabaeus Hong 1984
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Fm. †Alloioscarabaeidae Bai et al. 2012
+
G. †Alloioscarabaeus Bai et al. 2012
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Alloioscarabaeus cheni Bai et al. 2012
G. †Ateuchites Meunier 1898
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Ateuchites grandis Meunier 1898
Fm. Bolboceratidae Mulsant 1842 [dor beetle]
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Subfm. Athyreinae Lynch Arribálzaga 1878
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G. †Amberathyreus Bai and Zhang 2017
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Amberathyreus beuteli Bai and Zhang 2017
Subfm. Bolboceratinae Mulsant 1842
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Invalid names: Bolboceratini Mulsant 1842 [invalid subgroup]
G. †Cretobolbus Nikolajev 1996
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Cretobolbus rohdendorfi Nikolajev 1996
G. †Ceafornotensis Woolley 2016
+
G. †Cretaclopus Nikolajev 2004
+
Cretaclopus longipes Ponomarenko 1986
G. †Cretonitis Nikolajev 2007
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Cretonitis copripes Nikolajev 2007
Fm. Diphyllostomatidae Holloway 1972
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G. †Prodiphyllostoma Yamamoto 2024
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Fm. Geotrupidae Latreille 1802 [earth-boring dung beetle]
+
Subfm. Geotrupinae Latreille 1802
+
G. Ceratophyus Fischer von Waldheim 1823
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Ceratophyus yatagaii Aiba and Král 2023
Tr. †Cretogeotrupini Nikolajev 1996
+
G. †Cretogeotrupes Nikolajev 1992
Tr. Geotrupini Latreille 1802
+
G. Geotrupes Latreille 1796
+
Invalid names: Obitiscarabaeus Hong 1984 [synonym]
G. †Lithogeotrupes Nikolajev 2008
+
Lithogeotrupes confusus Nikolajev 2008
G. †Parageotrupes Nikolajev and Ren 2010
+
Parageotrupes incanus Nikolajev and Ren 2010
G. †Orrhodomala Zhang 1989
+
Orrhodomala protista Zhang 1989
G. Typhaeus Leach 1815
Invalid names: Lethrinae Mulsant and Rey 1871 [empty], Taurocerastinae [empty]
G. †Geotrupoides Handlirsch 1906
+
Geotrupoides lithographicus Deichmüller 1886
Geotrupoides saxosus Zhang 1997
Fm. Glaphyridae MacLeay 1819 [bumble bee scarab beetle]
+
Subfm. Amphicominae Blanchard 1845
+
G. Amphicoma Latreille 1807
Subfm. †Cretoglaphyrinae Nikolajev 2005
+
G. †Cretoglaphyrus Nikolajev 2005
+
Cretoglaphyrus calvescens Nikolajev 2005
Cretoglaphyrus leptopterus Nikolajev 2005
Cretoglaphyrus olenguicus Nikolajev 2005
Cretoglaphyrus rohdendorfi Nikolajev 2005
Cretoglaphyrus zherikhini Nikolajev 2005
G. †Cretohypna Yan et al. 2012
+
Cretohypna cristata Yan et al. 2012
Cretohypna puncta Zhao et al. 2016
Cretohypna robusta Zhao et al. 2016
Cretohypna yixianensis Nikolajev and Ren 2015
Subfm. Glaphyrinae MacLeay 1819
+
G. Glaphyrus Latreille 1802
+
Glaphyrus ancestralis Nikolajev and Ren 2011
Glaphyrus antiquus Heer 1862
G. Lichnanthe Burmeister 1844
+
Lichnanthe defuncta Wickham 1910
G. †Lithohypna Nikolajev et al. 2011
+
Lithohypna chifengensis Nikolajev et al. 2011
Lithohypna laoningensis Nikolajev 2014
Lithohypna lepticephala Nikolajev and Ren 2012
Lithohypna longula Nikolajev and Ren 2012
Lithohypna tuberculata Nikolajev and Ren 2012
Lithohypna yuxiana Nikolajev and Ren 2012
G. †Mesohypna Nikolajev and Ren 2013
+
Mesohypna lopatini Nikolajev and Ren 2013
Mesohypna probata Nikolajev 2014
G. †Paraglaphyrus Nikolajev 2015
+
Paraglaphyrus ovalis Nikolajev 2015
Paraglaphyrus robustus Nikolajev 2015
Paraglaphyrus subtilis Nikolajev 2015
Paraglaphyrus yixianensis Nikolajev 2015
Fm. Glaresidae Prudhomme de Borre 1886 [enigmatic scarab beetle]
+
G. †Cretoglaresis Nikolajev 2007
+
Cretoglaresis nana Nikolajev 2007
Cretoglaresis ovalis Nikolajev 2009
G. Glaresis Erichson 1848
+
Glaresis burmitica Cai and Huang 2018
Glaresis cretacea Nikolajev 2007
Glaresis orthochilus Bai et al. 2010
Glaresis tridentata Bai et al. 2014
G. †Lithoglaresis Nikolajev 2007
+
Lithoglaresis ponomarekoi Nikolajev 2007
Invalid names: Aphodiites Heer 1864 [nomen dubium]
G. †Holcorobeus Nikritin 1977
+
Holcorobeus atrosulcatus Nikolajev 1992
Holcorobeus evittatus Zhang 1992
Holcorobeus nigrovittatus Nikolajev 1992
Holcorobeus vittatus Nikritin 1977
Invalid names: Holcorobeus (Holcorobeus) Nikritin 1977 [invalid subgroup]
G. †Hongscarabaeus Krell 2000
+
Invalid names: Proscarabaeus Hong 1982 [replaced]
Fm. Hybosoridae Erichson 1847 [scavenger scarab beetle]
+
Subfm. Anaidinae Nikolajev 1996
+
G. Anaides Westwood 1842
+
Anaides howdeni Ocampo 2006
G. †Crassisorus Nikolajev et al. 2012
+
Crassisorus fractus Nikolajev et al. 2012
G. †Cretanaides Nikolajev 1996
+
Cretanaides trogopterus Nikolajev 1996
G. †Protanaides Nikolajev 2010
+
Protanaides sibiricus Nikolajev 2010
Subfm. Ceratocanthinae Martinez 1968
+
Tr. Ceratocanthini Martinez 1968
+
G. Ceratocanthus White 1842
G. Germarostes Paulian 1982
G. Nesopalla Paulian and Howden 1982
Invalid names: Acanthocerini [replaced]
G. †Palaeopycnus Lu et al. 2022
+
Palaeopycnus circus Lu et al. 2022
Palaeopycnus fushengii Lu et al. 2022
Invalid names: Ivieolini Howden and Gill 2000 [empty]
G. †Coprologus Heer 1847
+
Coprologus gracilis Heer 1847
Subfm. Hybosorinae Erichson 1847
+
G. †Cretohybosorus Nikolajev 1999
+
Cretohybosorus buryaticus Nikolajev 1999
Cretohybosorus striatulus Nikolajev 1999
G. †Fortishybosorus Yan et al. 2013
+
Fortishybosorus ericeusicus Yan et al. 2013
G. Hybosorus MacLeay 1819
+
Hybosorus lividus Heer 1862
Hybosorus ocampoi Bai and Zhang 2016
G. †Jurahybosorus Nikolajev 2005
+
Jurahybosorus kazakhstanicus Nikolajev 2008
Jurahybosorus mongolicus Nikolajev 2005
G. †Leptosorus Nikolajev 2006
+
Leptosorus fortus Ren et al. 1995
Leptosorus zherikhini Nikolajev 2006
G. Phaeochrous Laporte 1840
+
Phaeochrous tertiarium Deichmüller 1881
G. †Protohybosorus Nikolajev 2010
+
Protohybosorus grandissimus Nikolajev 2010
Protohybosorus karatavicus Nikolajev 2010
Protohybosorus mesoasiaticus Nikolajev 2010
Invalid names: Anaidini Nikolajev 1996 [empty], Hybosorini Erichson 1847 [empty]
Subfm. Liparochrinae Ocampo 2006
+
G. †Libanochrus Kirejtshuk et al. 2011
+
Libanochrus calvus Kirejtshuk et al. 2011
Subfm. †Mimaphodiinae Nikolajev 2007
+
G. †Mimaphodius Nikolajev 2007
+
Mimaphodius pusillus Nikolajev 2007
G. †Procoilodes Ocampo 2002
+
Procoilodes adrastus Ocampo 2002
G. †Pulcherhybosorus Yan et al. 2012
+
Pulcherhybosorus tridentatus Yan et al. 2012
G. †Sinochaetodus Lu et al. 2018
+
Sinochaetodus tridentatus Lu et al. 2018
G. †Sinohybosorus Nie et al. 2018
+
Sinohybosorus cheni Nie et al. 2018
G. †Tyrannasorus Ratcliffe and Ocampo 2001
+
Tyrannasorus rex Ratcliffe and Ocampo 2001
G. †Hybosorites Nikolajev 1996
+
Hybosorites fissuratus Nikolajev 1996
G. †Juraclopus Nikolajev 2004
+
Juraclopus rohdendorfi Nikolajev 2004
Fm. Lucanidae Latreille 1804 [stag beetle]
+
Subfm. Aesalinae MacLeay 1819
+
Tr. Aesalini MacLeay 1819
+
G. †Cretaesalus Nikolajev 1993
G. †Juraesalus Nikolajev et al. 2011
G. †Sinaesalus Nikolajev et al. 2011
Tr. Ceratognathini Sharp 1899
+
G. †Cretognathus Yamamoto 2023
G. †Oncelytris Li and Cai 2023
Tr. Nicagini LeConte 1861
+
G. †Protonicagus Cai et al. 2017
Subfm. †Ceruchitinae Nikolajev 2006
+
G. †Ceruchites Statz 1952
+
Ceruchites hahnei Statz 1952
G. †Cretolucanus Nikolajev 2007
+
Cretolucanus longus Nikolajev 2007
Cretolucanus ordinarius Nikolajev 2007
Cretolucanus sibericus Nikolajev 2007
G. †Dorcasoides Motschulsky 1856
+
Dorcasoides bilobus Motschulsky 1856
G. †Electraesalopsis Bai et al. 2017
+
Electraesalopsis beuteli Bai et al. 2017
Subfm. Lampriminae MacLeay 1819
+
G. †Protognathinus Chalumeau and Brochier 2001
+
Protognathinus spielbergi Chalumeau and Brochier 2001
Subfm. †Litholampriminae Nikolajev 2015
+
G. †Litholamprima Nikolajev and Ren 2015
+
Litholamprima longimana Nikolajev and Ren 2015
Litholamprima qizhihaoi Jiang et al. 2022
Subfm. Lucaninae Latreille 1804
+
G. †Anisoodontus Wu et al. 2022
+
Anisoodontus qizhihaoi Wu et al. 2022
Anisoodontus xiafangyuani Wu et al. 2022
Tr. Lucanini Latreille 1804
+
G. Dorcus MacLeay 1819
G. Lucanus Scopoli 1763
G. Penichrolucanus Deyrolle 1863
Invalid names: Figulidae Burmeister 1847 [synonym]
G. †Miocenidorcus Riou 1999
+
Tr. †Platycerini Mulsant 1842
+
G. Platycerus Geoffroy 1762
G. †Succiniplatycerus Nikolayev 1990
G. †Prolucanus Qi et al. 2022
+
Prolucanus beipiaoensis Qi et al. 2022
Invalid names: Chiasognathini Burmeister 1847 [empty], Lucanus fossilis Wickham 1913 [nomen dubium], Penichrolucaninae Arrow 1950 [synonym], Platycerus sepultus Germar 1837 [nomen dubium]
G. †Paleognathus Waga 1883
+
Paleognathus succini Waga 1883
Subfm. †Paralucaninae Nikolajev 2000
+
G. †Paralucanus Nikolajev 2000
+
Paralucanus mesozoicus Nikolajev 2000
G. †Prosinodendron Bai et al. 2012
+
Prosinodendron krelli Bai et al. 2012
Subfm. †Protolucaninae Nikolajev 2007
+
G. †Protolucanus Nikolajev 2007
+
Protolucanus jurassicus Nikolajev 2007
Subfm. Syndesinae MacLeay 1819
+
G. Ceruchus MacLeay 1819
+
Ceruchus fuchsii Wickham 1911
G. Syndesus Macleay 1819
+
Syndesus ambericus Woodruff 2009
G. †Mahengea Strümpher et al. 2023
+
Mahengea mckayi Strümpher et al. 2023
G. †Mesaclopus Nikolajev 1992
+
Mesaclopus mongolicus Nikolajev 1992
G. †Mesoscarabaeus Hong 1982
+
G. †Mongolrobeus Nikolajev 2004
+
Mongolrobeus zherikhini Nikolajev 2004
Fm. Ochodaeidae Streubel 1846 [sand-loving scarab beetle]
+
Subfm. Chaetocanthinae Scholtz 1988
+
Tr. Synochodaeini Scholtz 1988
+
G. †Yixianochodaeus Nikolajev 2015
Subfm. †Cretochodaeinae Nikolajev 1995
+
G. †Cretochodaeus Nikolajev 1995
+
Cretochodaeus mongolicus Nikolajev 1995
Cretochodaeus striatus Nikolajev 1995
G. †Mioochodaeus Nikolajev 1995
+
Mioochodaeus proaevus Germar 1849
G. †Mesochodaeus Nikolajev and Ren 2010
+
Mesochodaeus daohugouensis Nikolajev and Ren 2010
Subfm. Ochodaeinae Streubel 1846
+
G. †Lithochodaeus Nikolajev 2007
+
Lithochodaeus cretaceus Nikolajev 2007
G. †Opiselleipon Bode 1953
+
Opiselleipon gravis Bode 1953
Fm. Passalidae Leach 1815 [bess beetle]
+
Subfm. Aulacocyclinae Kaup 1868
+
Tr. Cylindrocaulini Zang 1905
+
Invalid names: Ceracyclini Boucher and Bai 2017 [synonym]
Subfm. Passalinae Leach 1815
+
Tr. Macrolinini Kaup 1871
+
G. Macrolinus Kaup 1868
G. †Serrulus Hong 1984
Tr. Passalini Leach 1815
+
G. Passalus Fabricius 1792
Tr. †Protopassalini Santos et al. 2020
+
G. †Protopassalus Santos et al. 2020
Fm. †Passalopalpidae Boucher and Bai 2016
+
G. †Mesoceratocanthus Nikolajev et al. 2010
+
Mesoceratocanthus tuberculifrons Nikolajev et al. 2010
G. †Passalopalpus Boucher and Bai 2016
+
Passalopalpus cheni Boucher et al. 2016
Fm. Pleocomidae Leconte 1861 [rain beetle]
+
Subfm. †Archescarabaeinae Nikolajev 2010
+
G. †Archescarabaeus Nikolajev 2010
+
Archescarabaeus mesozoicus Nikolajev 2010
Subfm. †Cretocominae Nikolajev 2002
+
G. †Cretocoma Nikolajev 2002
+
Cretocoma tologoica Nikolajev 2002
Subfm. Pleocominae Leconte 1861
+
G. Pleocoma Le Conte 1856
+
Pleocoma dolichophylla Nikolajev and Ren 2012
G. †Proteroscarabaeus Grabau 1923
+
Proteroscarabaeus magnus Nikolajev 1996
Proteroscarabaeus nikritini Nikolajev 1996
Proteroscarabaeus yeni Grabau 1923
G. †Prionocephale Lin 1980
+
G. †Prophaenognatha Bai et al. 2011
+
Prophaenognatha robusta Bai et al. 2011
Fm. Scarabaeidae Latreille 1802 [scarab beetle]
+
Subfm. Aclopinae Blanchard 1850
+
Invalid names: Aclopini Blanchard 1850 [empty]
Subfm. Aegialiinae Laporte 1840
+
G. Aegialia Latreille 1807
+
Subg. Aegialia (Psammoporus) Thomson 1859
Aegialia cylindrica Eschscholtz 1822
Aegialia lacustris Le Conte 1850
+
Invalid names: Loricera lutosa Scudder 1890 [synonym]
Aegialia rupta Scudder 1890
Aegialia sibirica Stebnicka 1977
G. †Archeopsammoporus Minkina 2020
+
G. †Cretaegialia Nikolayev 1993
+
Cretaegialia aphodiiformis Nikolayev 1993
Cretaegialia rhypariformis Nikolayev 1993
G. †Antiqusolidus Lu et al. 2023
+
Antiqusolidus maculatus Lu et al. 2023
Subfm. Aphodiinae Leach 1815
+
Tr. Aphodiini Leach 1815
+
G. Agoliinus Schmidt 1913
Subtr. Aphodiina Leach 1815
G. Caligodorus Gordon and Skelley 2007
G. Dialytes Harold 1869
G. Dialytodius Gordon and Skelley 2007
G. Pardalosus Gordon and Skelley 2007
G. Planolinoides Dellacasa and Dellacasa 2005
G. †Cretorabaeus Nikolajev 1995
+
Cretorabaeus elongatus Nikolajev 1995
Tr. Eupariini Schmidt 1910
+
G. Airapus Stebnicka and Howden 1996
G. Ataenius Harold 1867
G. Oxyomus Kirby and Spence 1828
G. Saprosites Redtenbacher 1858
Tr. †Psammaegialiini Nikolajev et al. 2014
+
G. †Psammaegialia Nikolajev et al. 2014
Tr. Psammodiini Mulsant 1842
+
Subtr. Psammodiina Mulsant 1842
+
Invalid names: Pleurophorina Mulsant 1842 [synonym]
Subtr. Rhyssemina Pittinno and Marianni 1986
Tr. Rhyparini Schmidt 1910
+
G. Leptorhyparus Howden 2003
G. †Priscyparus Skelley 2021
Subfm. Aulonocneminae Janssens 1946
+
G. †Protopalnia Nikolajev 2007
+
Protopalnia magadanica Nikolajev 2007
G. Caccobius Thompson 1859
+
Caccobius schreberi Linnaeus 1767
Subfm. Cetoniinae Leach 1815 [flower chafer]
+
Tr. Cetoniini Leach 1815
+
Subtr. Cetoniina Leach 1815
Tr. Schizorhinini Burmeister 1842
+
G. †Genonota Zhang 1989
G. Heterosoma Schaum 1845
Subtr. Lomapterina Burmeister 1842
G. †Paleotrichius Poinar 2011
Tr. Trichiini Fleming 1821
+
G. Gnorimus Lepeletier and Serville 1828
Subtr. Trichiina Fleming 1821
Tr. Valgini Mulsant 1842
+
Subtr. Valgina Mulsant 1842
Subfm. †Cretoscarabaeinae Nikolajev 1995
+
G. †Cretoscarabaeus Nikolajev 1995
+
Cretoscarabaeus gibbosus Nikolajev 1995
Cretoscarabaeus lentiginosus Nikolajev 1995
Subfm. Dynastinae MacLeay 1819 [rhinoceras beetle]
+
G. †Calcitoryctes Krell 2011
+
G. Cyclocephala Dejean 1821
+
G. Ligyrus Burmeister 1847
+
Ligyrus compositus Wickham 1911
Ligyrus effetus Wickham 1914
Tr. Oryctini Mulsant 1842
+
G. Oryctes Hellwig 1798
G. Strategus Kirby 1828
G. †Oryctoantiquus Ratcliffe and Smith 2005
+
Oryctoantiquus borealis Ratcliffe and Smith 2005
Tr. Pentodontini Mulsant 1842
+
G. Diloboderus Reiche 1859
Subtr. Pentodontina Mulsant 1842
Invalid names: Pentodon proserpinae Heer 1862 [nomen dubium]
Subfm. †Electrorubesopsinae Bai and Wang 2018
+
G. †Electrorubesopsis Bai and Wang 2018
+
Electrorubesopsis beuteli Bai and Wang 2018
Subfm. Eremazinae Stebnicka 1977
+
Subfm. †Yixianscarabaeinae Nikolajev 2015
+
G. †Yixianscarabaeus Nikolajev 2015
G. Liothorax Motschulsky 1859
+
Liothorax kraatzi Harold 1868
Subfm. †Lithoscarabaeinae Nikolajev 1992
+
G. †Baisarabaeus Nikolajev 2005
+
Baisarabaeus rugosus Nikolajev 2005
Baisarabaeus yixianensis Nikolajev et al. 2013
G. †Lithoscarabaeus Nikolajev 1992
+
Lithoscarabaeus baissensis Nikritin 1977
Subfm. Melolonthinae Leach 1819
+
Tr. †Cretomelolonthini Nikolajev 1998
+
G. †Cretomelolontha Nikolajev 1998
Tr. Diplotaxini Kirby 1837
+
G. Diplotaxis Kirby 1837
G. †Eophyllocerus Haupt 1950
+
Tr. Euchirini Hope 1840
+
G. Cheirotonus Hope 1840
Tr. Hopliini Latreille 1829
+
Subtr. Hopliina Latreille 1829
Subtr. Pachycnemina Laporte 1840
G. Listrochelus Blanchard 1851
+
Listrochelus puerilis Wickham 1914
Tr. Macrodactylini Kirby 1837
+
G. Macrodactylus Latreille 1827
Tr. Melolonthini Leach 1819
+
Subtr. Melolonthina Leach 1819
G. Phyllophaga Harris 1826
+
Invalid names: Lachnosterna Hope 1837 [synonym]
Subtr. Rhizotrogina Burmeister 1855
Invalid names: Schizonychina Burmeister 1855 [empty]
G. †Melolonthites Heer 1847
+
Melolonthites collinsi Wickham 1929
Melolonthites interemtus Cockerell 1926
Melolonthites kollari Heer 1847
Tr. Sericini Kirby 1837
+
G. †Cretoserica Nikolajev 1998
G. †Lithanomala Nikolajev 1992
G. Maladera Mulsant 1871
G. †Mioserica Zhang 1989
Subtr. Sericina Kirby 1837
G. †Mesydra Ponomarenko 1977
+
Mesydra elongata Ponomarenko 1977
Subfm. †Prototroginae Nikolajev 2000
+
G. †Prototrox Nikolajev 2000
+
Prototrox transbaikalicus Nikolajev 2000
Subfm. Rutelinae MacLeay 1819
+
Tr. Adoretini Burmeister 1844
+
Subtr. Adoretina Burmeister 1844
Tr. Anomalini Streubel 1839
+
Subtr. Anomalina Streubel 1839
G. †Anomalites Frič 1885
+
Anomalites fugitivus Frič 1885
Tr. Anoplognathini MacLeay 1819
+
Subtr. Anoplognathina MacLeay 1819
G. Holotrichia Hope 1837
+
Holotrichia cressona Zhang et al. 1994
Holotrichia spatha Zhang 1989
G. Mimela Kirby 1823
+
Mimela rhenana von Heyden 1862
Mimela testaceipes Motschulsky 1860
Invalid names: Rhombonyx Hope 1837 [synonym]
G. †Miolachnosterna Wickham 1914
+
G. †Pelidnotites Cockerell 1920
+
Pelidnotites atavus Cockerell 1920
G. †Petraeianus Zhang 1989
+
Petraeianus ruderatus Zhang 1989
G. †Petulantis Zhang et al. 1994
+
Petulantis yimengensis Zhang et al. 1994
G. †Zhangsunia Krell 2000
+
Zhangsunia extumida Zhang et al. 1994
Invalid names: Dolichopoda Zhang et al. 1994 [replaced]
Subfm. Scarabaeinae Latreille 1802
+
G. Anachalcos Hope 1837
+
Anachalcos mfwangani Paulian 1976
Tr. Ateuchini Perty 1830
+
Subtr. Ateuchina Perty 1830
G. †Lobateuchus Montreuil et al. 2010
G. Canthochilum Chapin 1934
+
Canthochilum alleni Tarasov and Vaz-de-Mello 2016
Canthochilum philipsivieorum Tarasov and Vaz-de-Mello 2016
Tr. Coprini Leach 1815
+
G. Copris Geoffroy 1762
Tr. Deltochilini Lacordaire 1856
+
G. Canthon Hoffmannsegg 1817
G. Deltochilum Eschscholtz 1822
G. Scybalophagus Martínez 1953
Tr. Gymnopleurini Lacordaire 1856
+
G. Gymnopleurus Illiger 1803
G. Heliocopris Hope 1837
+
Heliocopris antiquus Fujiyama 1968
G. Metacatharsius Paulian 1939
+
Metacatharsius rusingae Paulian 1976
Tr. Oniticellini Kolbe 1905
+
Subtr. Drepanocerina van Lansberge 1875
Subtr. Oniticellina Kolbe 1905
Tr. Onitini Laporte 1840
+
G. Onitis Fabricius 1798
Tr. Onthophagini Burmeister 1846
+
G. Onthophagus Latreille 1802
G. Phalops Erichson 1848
Tr. Phanaeini Hope 1838
+
G. Phanaeus Macleay 1819
+
Invalid names: Palaeocopris Pierce 1946 [synonym]
Tr. Scarabaeini Latreille 1802
+
G. Scarabaeus Linnaeus 1758
Invalid names: Onthophagus spitsbergeniensis Krell 2010 [nomen dubium]
G. Triodonta Mulsant 1842
Invalid names: Allidiostomatinae Arrow 1940 [empty], Coprophaginae [empty], Dynamopodinae Arrow 1911 [empty], Oniticellus amplicollis Heer 1862 [nomen dubium], Orphninae Erichson 1847 [empty], Pachypodinae Erichson 1840 [empty], Phaenomeridinae Erichson 1847 [empty]
G. †Scelocopris Zhang 1989
+
Scelocopris enertheus Zhang 1989
Fm. †Septiventeridae Bai et al. 2013
+
G. †Septiventer Bai et al. 2013
+
Septiventer quadridentatus Bai et al. 2013
Fm. Trogidae MacLeay 1819 [hide beetle]
+
Subfm. †Kresnikinae Tihelka et al. 2019
+
G. †Kresnikus Tihelka et al. 2019
+
Kresnikus beynoni Tihelka et al. 2019
Subfm. Omorginae Nikolajev 2005
+
G. †Cretomorgus Nikolajev 2007
+
Cretomorgus ikhbogdensis Nikolajev 2007
Subfm. Troginae MacLeay 1819
+
Tr. †Avitortorini Nikolajev 2007
+
G. †Avitortor Ponomarenko 1977
G. †Paratrox Nikolajev 2009
G. Trox Fabricius 1775
+
Trox antiquus Wickham 1909
Trox cretaceus Nikolajev 2007
Trox minutus Nikolajev 2008
Trox sibericus Nikolajev 2007
Invalid names: Trox oustaleti Scudder 1879 [nomen dubium]
Invalid names: Belohinidae Paulian 1959 [empty], Copris subterraneus Heer 1862 [nomen dubium], Gymnopleurus deperditus Heer 1862 [nomen dubium], Holcorobeini Nikolajev 1992 [invalid subgroup], Onthophagus urusheeri Krell 2000 [nomen dubium]
Superfm. Staphylinoidea Latreille 1802
+
Fm. Agyrtidae Thomson 1859 [primitive carrion beetle]
+
Subfm. Agyrtinae Thomson 1859
+
G. Agyrtes Frölich 1799
G. Ipelates Reitter 1884
+
Ipelates kerneggeri Perkovsky 2005
Ipelates weitshati Perkovsky 2007
G. †Eosilphites Haupt 1950
+
Eosilphites decoratus Haupt 1950
G. †Mesecanus Newton 1982
+
Mesecanus communis Ponomarenko 1977
Invalid names: Mesagyrtes Ponomarenko 1977 [replaced]
Subfm. Necrophilinae Newton 1997
+
G. †Palaeonecrophilus Strelnikova et al. 2020
+
Palaeonecrophilus buryaticus Strelnikova et al. 2020
G. †Ponomarenkia Perkovsky 2001
+
Ponomarenkia parva Ponomarenko 1990
G. †Prosilpha Hong 1982
+
Prosilpha nigrita Hong 1982
Subfm. Pterolomatinae Thomson 1862
+
G. Pteroloma Gyllenhal 1827
+
Pteroloma forsstromii Gyllenhal 1810
G. †Sinosilphia Hong and Wang 1990
+
Sinosilphia punctata Hong and Wang 1990
Fm. Colonidae Horn 1880
+
G. Colon Herbst 1797
+
Colon burmiticum Yamamoto 2018
G. Colonellus Szymczakowski 1964
+
Colonellus burmiticus Cai and Huang 2017
Fm. Hydraenidae Mulsant 1844 [minute moss beetle]
+
Subfm. Hydraeninae Mulsant 1844
+
G. †Archaeodraena Jäch and Yamamoto 2017
+
Archaeodraena cretacea Jäch and Yamamoto 2017
G. †Davidraenites Kirejtshuk 2019
+
Davidraenites gratshevi Kirejtshuk 2019
Davidraenites interruptus Kirejtshuk 2019
Davidraenites spurcus Kirejtshuk 2019
G. †Eolimnebius Kirejtshuk 2019
+
Eolimnebius fossilis Kirejtshuk 2019
Tr. Hydraenini Mulsant 1844
+
G. Hydraena Kugelann 1794
G. †Hydraenites Kirejtshuk 2019
+
Hydraenites gracilimmus Kirejtshuk 2019
Tr. Limnebiini [minute moss beetle]
+
G. Limnebius Leach 1815
G. †Metacoxites Kirejtshuk 2019
+
Metacoxites ventritalis Kirejtshuk 2019
G. †Ochtebiites Ponomarenko 1977
+
Ochtebiites altus Ponomarenko 1977
Ochtebiites incertus Ponomarenko 1977
Ochtebiites manlaicus Ponomarenko 1980
Ochtebiites minor Ponomarenko 1985
Subfm. Ochthebiinae Thomson 1859
+
Tr. Ochthebiini Thomson 1859
+
Subtr. Ochthebiina Thomson 1859
Invalid names: Prosthetopinae Perkins 1994 [empty]
Fm. Jacobsoniidae Heller 1926
+
Subfm. Derolathrinae Sen Gupta 1979
+
G. Derolathrus Sharp 1908
+
Derolathrus abyssus Yamamoto and Parker 2017
Derolathrus capdoliensis Tihelka et al. 2022
Derolathrus groehni Cai et al. 2016
G. Sarothrias Grouvelle 1918
+
Sarothrias cretaceus Cai et al. 2017
Invalid names: Sarothriidae [replaced]
Fm. Leiodidae Fleming 1821 [round fungus beetle]
+
G. Agaricophagus Schmidt 1841
+
Subfm. Camiarinae Jeannel 1911
+
Tr. Agyrtodini Jeannel 1936
+
G. †Cretagyrtodes Cai and Huang 2017
Subfm. Catopocerinae Hatch 1927
+
G. †Archaeocerus Perreau 2019
+
Archaeocerus uenoi Perreau 2019
Subfm. Cholevinae Kirby 1837
+
Tr. Anemadini Hatch 1927
+
G. Nemadus Thomson 1867
Tr. Cholevini Kirby 1837
+
G. Catops Paykull 1798
Subtr. Cholevina Kirby 1837
G. Prionochaeta Horn 1880
Invalid names: Catopina Jeannel 1922 [empty]
Tr. Oritocatopini Jeannel 1936
+
Invalid names: Cretaciella Perreau 2019 [nomen nudum]
Tr. Ptomaphagini Jeannel 1911
+
G. †Cretoptomaphagus Bao and Antunes-Carvalho 2020
Subtr. Ptomaphagina Jeannel 1911
Subtr. Ptomaphaginina Szymczakowski 1964
G. Cholevinus Reitter 1901
+
Cholevinus sibiricus Jeannel 1923
Subfm. Leiodinae Fleming 1821
+
Tr. Anisotomini Horaninow 1834
+
Invalid names: Agathidiini Westwood 1838 [synonym]
Tr. Leiodini Fleming 1821
+
G. Leiodes Latreille 1796
Tr. Pseudoliodini Portevin 1926
+
G. †Mesagyrtoides Perkovsky 1999
G. †Tafforeus Perreau 2012
+
G. Aglyptinus Cockerell 1906
Tr. Sogdini Lopatin 1961
+
G. Hydnobius Schmidt 1841
Invalid names: Leptininae LeConte 1866 [empty], Platypsyllinae Ritsema 1869 [empty]
Fm. Ptiliidae Erichson 1845 [feather-winged beetle]
+
G. Microptilium Matthews 1872
+
Subfm. Nossidiinae Sörensson and Delgado 2019
+
G. †Crenossidium Li et al. 2022
+
Crenossidium slipinskii Li et al. 2022
Subfm. Ptiliinae Erichson 1845
+
Tr. Acrotrichini Reitter 1909
+
G. Acrotrichis Moltschulsky 1848
Tr. Cephaloplectini Sharp 1883
+
Invalid names: Limulodinae Ganglbauer 1899 [synonym]
Tr. Discheramocephalini Grebennikov 2009
+
G. †Kekveus Yamamoto et al. 2018
Tr. Ptenidiini Flach 1889
+
G. Ptenidium Erichson 1845
Tr. Ptiliini Erichson 1845
+
G. Micridium Motschulsky 1868
G. Ptilium Gyllenhal 1827
Tr. Ptinellini Reitter 1906
+
G. Ptinella Motschulsky 1844
Invalid names: Nanosellini Barber 1924 [empty]
Fm. Staphylinidae Latreille 1802 [rove beetle]
+
Subfm. Aleocharinae Fleming 1821
+
Tr. Aleocharini Fleming 1821
+
G. Aleochara Gravenhorst 1802
Tr. Athetini Casey 1910
+
Subtr. Athetina Casey 1910
Tr. Corotocini Fenyes 1918
+
Subtr. Corotocina Fenyes 1918
Tr. Gymnusini Heer 1839
+
G. Adinopsis Cameron 1919
G. †Cretodeinopsis Cai and Huang 2015
G. †Electrogymnusa Wolf-Schwenninger 2004
G. Gymnusa Gravenhorst 1806
Invalid names: Deinopsini Sharp 1883 [synonym]
Tr. Homalotini Heer 1839
+
Subtr. Bolitocharina Thomson 1859
Subtr. Gyrophaenina Kraatz 1856
Subtr. Homalotina Heer 1839
G. Leptusa Kraatz 1856
Tr. Hygronomini Thomson 1859
+
Subtr. Hygronomina Thomson 1859
Tr. Hypocyphtini Laporte 1835
+
G. †Baltioligota Pasnik 2005
Tr. Lomechusini Fleming 1821
+
Subtr. Lomechusina Fleming 1821
Tr. Mesoporini Cameron 1959
+
G. †Ambracyptus Lundgren 1984
+
Invalid names: Paracyptus Seevers 1971 [replaced]
G. Dictyon Fauvel 1900
G. †Mesosymbion Yamamoto et al. 2016
G. †Palaeomesoporus Yamamoto and Maruyama 2017
Tr. Oxypodini Thomson 1859
+
G. Oxypoda Mannerheim 1831
Tr. Trichopseniini Le Conte and Horn 1883
+
G. †Cretotrichopsenius Cai et al. 2017
G. Prorhinopsenius Pasteels and Kistner 1971
+
Invalid names: Palaeopsenius Seevers 1971 [synonym]
G. †Aleocharopsis Wickham 1913
+
Aleocharopsis caseyi Wickham 1913
Aleocharopsis secunda Wickham 1913
Subfm. Apateticinae Fauvel 1895
+
G. †Miosilpha Wickham 1912
+
Miosilpha necrophiloides Wickham 1912
G. †Apticax Schomann and Solodovnikov 2012
+
Apticax solidus Schomann and Solodovnikov 2012
Apticax volans Schomann and Solodovnikov 2012
G. †Bembicidiodes Schaufuss 1889
+
Bembicidiodes inaequicollis Schaufuss 1889
G. Boletobius Leach 1819
G. †Cardiolita Strand 1933
G. †Caririderma Martins-Neto 1990
+
Caririderma pilosa Martins-Neto 1990
Subfm. Coomaniinae Żyła and Solodovnikov 2020
+
G. Coomania Cameron 1939
+
Coomania enkarsios Jenkins Shaw et al. 2022
Coomania megistos Jenkins Shaw et al. 2022
Coomania yini Jenkins Shaw et al. 2022
Subfm. Dasycerinae Reitter 1887
+
G. †Cedasyrus Yin and Cai 2020
+
Cedasyrus dimorphus Yin and Cai 2020
G. †Protodasycerus Yamamoto 2016
+
Protodasycerus aenigmaticus Yamamoto 2016
Protodasycerus corpulentus Yin and Cai 2020
Protodasycerus gigas Yin and Cai 2020
Protodasycerus tuberculatus Yamamoto et al. 2020
G. †Vetudasycerus Cai et al. 2017
+
Vetudasycerus burmiticus Cai et al. 2017
G. Ellipsotomus Motschulsky 1845
Subfm. Euaesthetinae Thomson 1859
+
Tr. Euaesthetini Thomson 1859
+
G. Euaesthetus Gravenhorst 1806
G. Octavius Fauvel 1873
Tr. Nordenskioldiini Bernhauer and Schubert 1911
+
G. Nordenskioldia Sahlberg 1880
+
Invalid names: Libanoeuaesthetus Lefebvre et al. 2005 [synonym]
Subfm. Glypholomatinae Jeannel 1962
+
G. †Juroglypholoma Cai et al. 2012
+
Juroglypholoma antiquum Cai et al. 2012
Juroglypholoma talbragarense Cai et al. 2013
G. Heterosoma Bernhauer 1903
G. †Laostaphylinus Zhang 1988
+
Laostaphylinus fuscus Zhang 1988
Subfm. Megalopsidiinae Leng 1920
+
G. Megalopinus Eichelbaum 1915
+
Megalopinus extinctus Yamamoto and Solodovnikov 2016
G. †Mesapatetica Cai et al. 2014
+
Mesapatetica aenigmatica Cai et al. 2014
Subfm. Micropeplinae Leach 1815
+
G. Arrhenopeplus Koch 1937
+
G. Kalissus Leconte 1874
+
Kalissus nitidus LeConte 1874
G. Micropeplus Latreille 1809
+
Micropeplus caelatus Erichson 1839
Micropeplus cribratus LeConte 1863
Micropeplus fulvus Erichson 1840
Micropeplus hoogendorni Matthews 1970
Micropeplus hopkinsi Matthews 1970
Micropeplus macrofulvus Gersdorf 1976
Micropeplus nelsoni Campbell 1989
Micropeplus pengweii Jiang et al. 2020
Micropeplus porcatus Paykull 1789
Micropeplus punctatus LeConte 1863
G. †Protopeplus Cai and Huang 2014
+
Protopeplus cretaceus Cai and Huang 2014
Subfm. Mycetoporinae Thomson 1859
+
G. Bolitobius Leach 1819
G. †Cuneocharis Ryvkin 1990
+
Cuneocharis elongatus Ryvkin 1990
G. †Glabrimycetoporus Yue et al. 2009
+
Glabrimycetoporus amoenus Yue et al. 2009
G. Ischnosoma Stephens 1829
+
Ischnosoma flavicolle LeConte 1863
G. Lordithon Thomson 1859
+
Lordithon durabilis Scudder 1900
Lordithon funditus Scudder 1900
Lordithon longiceps LeConte 1863
Lordithon lyelli Scudder 1900
Lordithon stygis Scudder 1900
G. Mycetoporus Mannerheim 1830
+
Mycetoporus demersus Scudder 1900
G. Parabolitobius Li et al. 2000
+
Parabolitobius antiquus Yamamoto 2023
Parabolitobius groehni Schülke 2000
G. †Ryvkinius Herman 2001
+
Ryvkinius gracilis Ryvkin 1990
Invalid names: Mesoporus Ryvkin 1990 [replaced]
G. †Undiatina Ryvkin 1990
+
Undiatina pilosa Ryvkin 1990
Invalid names: Bolitobiini Horn 1877 [synonym]
Subfm. Olisthaerinae Thomson 1858
+
G. †Anicula Ryvkin 1985
+
Anicula inferna Ryvkin 1985
G. †Protolisthaerus Cai et al. 2015
+
Protolisthaerus jurassicus Cai et al. 2015
Subfm. Omaliinae MacLeay 1825
+
Tr. Anthophagini Thomson 1859
+
G. Acidota Stephens 1829
G. Anthobium Leach 1819
+
Invalid names: Lathrimaeum Erichson 1839 [synonym]
G. Anthophagus Gravenhorst 1802
G. Arpedium Erichson 1839
G. Artochia Casey 1893
G. Eucnecosum Reitter 1909
G. Geodromicus Redtenbacher 1857
G. Lesteva Latreille 1796
G. Microedus Leconte 1874
G. Olophrum Erichson 1839
G. Orobanus LeConte 1878
G. Pelecomalium Casey 1886
G. Phlaeopterus Motschulsky 1853
G. †Sinanthobium Cai and Huang 2013
G. Unamis Casey 1893
Invalid names: Lestevini Mulsant and Rey 1880 [synonym]
G. †Archodromus Tikhomirova 1968
+
Archodromus brachypterus Tikhomirova 1968
Archodromus comptus Tikhomirova 1968
Tr. Coryphiini Jakobson 1908
+
Subtr. Boreaphilina Zerche 1990
Subtr. Coryphiina Jakobson 1908
G. †Daiodromus Ryvkin 1990
+
Daiodromus umbrosus Ryvkin 1990
G. †Eophyllodrepa Ryvkin 1985
+
Eophyllodrepa propinqua Ryvkin 1985
Tr. Eusphalerini Hatch 1957
+
G. Eusphalerum Kraatz 1857
G. †Globoides Tikhomirova 1968
+
Globoides microps Tikhomirova 1968
Globoides oculatus Tikhomirova 1968
G. †Mesodeliphrum Ryvkin 1990
+
Mesodeliphrum rufulum Ryvkin 1990
G. †Morsum Ryvkin 1985
+
Morsum abdominale Ryvkin 1985
Tr. Omaliini MacLeay 1825
+
G. Acrulia Thomson 1858
G. †Duocalcar Peris and Thayer 2014
G. Hapalaraea Thomson 1858
G. Micralymma Westwood 1838
G. Omalium Gravenhorst 1802
G. Paraphloeostiba Steel 1960
G. Phyllodrepa Thomson 1859
G. Pycnoglypta Thomson 1858 [winter rove beetle]
G. †Porrhodromus Tikhomirova 1968
+
Porrhodromus communis Tikhomirova 1968
G. †Prodaia Ryvkin 1990
+
Prodaia inflata Ryvkin 1990
G. †Protostaphylinus Lin 1976
+
G. †Pseudolesteua Schaufuss 1890
+
Pseudolesteua insinuans Schaufuss 1890
Subfm. Osoriinae Erichson 1839
+
Tr. Eleusinini Sharp 1887
+
G. Eleusis Laporte 1835
Tr. Leptochirini Sharp 1887
+
G. †Cretochirus Yamamoto 2019
G. Priochirus Sharp 1887
Tr. Osoriini Erichson 1839
+
G. Neosorius Fagel 1959
G. Nototorchus McColl 1985
G. Osoriellus Fagel 1959
G. Osorius Guerin-Meneville 1829
G. †Sinolispinodes Zhang 1989
+
Sinolispinodes torosus Zhang 1989
Tr. Thoracophorini Reitter 1909
+
Subtr. Clavilispinina Newton and Thayer 1992
Subtr. Lispinina Bernhauer and Schubert 1910
G. †Lispinomimus Irmler 2003
G. †Paleosorius Ortega-Blanco et al. 2013
Subtr. Thoracophorina Reitter 1909
Invalid names: Lispinini Bernhauer and Schubert 1910 [synonym]
Subfm. Oxyporinae Fleming 1821
+
G. †Cretoxyporus Cai and Huang 2014
+
Cretoxyporus extraneus Cai and Huang 2014
G. Oxyporus Fabricius 1775
+
Oxyporus blumenbachii Gravenhorst 1806
Oxyporus cretaceous Yamamoto 2017
Oxyporus impressus Piton 1940
Oxyporus seuberti Heer 1862
Oxyporus stiriacus Scudder 1890
Oxyporus vulcanus von Heyden and von Heyden 1866
Oxyporus yixianus Solodovnikov and Yue 2011
G. †Protoxyporus Cai and Huang 2014
+
Protoxyporus grandis Cai and Huang 2014
Subfm. Oxytelinae Fleming 1821
+
Tr. Blediini Ádám 2001
+
G. Bledius Leach 1819
G. †Kupakara Chen et al. 2023
Tr. Coprophilini Heer 1839
+
G. †Gollandia Makranczy et al. 2018
G. †Mesocoprophilus Cai and Huang 2013
G. †Sinoxytelus Yue et al. 2010
Tr. Deleasterini Reitter 1909
+
G. Deleaster Erichson 1839
G. †Protodeleaster Cai et al. 2013
G. Syntomium Curtis 1828
G. †Megalymma Tikhomirova 1980
+
Megalymma gigantea Tikhomirova 1980
Megalymma rohdendorfi Tikhomirova 1980
G. †Mesoxytelus Tikhomirova 1968
+
Mesoxytelus mandibularis Tikhomirova 1968
Mesoxytelus parvus Tikhomirova 1968
G. †Morda Ryvkin 1990
+
Morda mora Ryvkin 1990
Tr. Oxytelini Fleming 1821
+
G. Anotylus Thomson 1859
G. Carpelimus Leach 1819
+
Invalid names: Trogophloeus Mannerheim 1830 [synonym]
G. †Dolichoxenus Engel and Chatzimanolis 2009
G. Oxytelus Gravenhorst 1802
G. Platystethus Mannerheim 1830
Tr. Syntomiini Böving and Craighead 1931
+
G. †Pseudanotylus Cai and Huang 2013
G. †Vidya Chen et al. 2023
Tr. Thinobiini Sahlberg 1876
+
G. Aploderus Stephens 1833
G. †Prajna Lu et al. 2017
G. †Turgaphloeus Ryvkin 1990
+
Turgaphloeus pubescens Ryvkin 1990
Invalid names: Euphaniini Reitter 1909 [empty], Piestini Erichson 1839 [empty]
Subfm. Paederinae Fleming 1821
+
Tr. Lathrobiini Laporte 1835
+
Subtr. Lathrobiina Laporte 1835
Subtr. Medonina Casey 1905
G. Micrillus Erichson 1839
Subtr. Scopaeina Mulsant and Rey 1878
G. Scymbalium Erichson 1839
G. †Lithoplanes Scudder 1886
+
Lithoplanes deleta Oustalet 1874
Lithoplanes elongata Oustalet 1874
Invalid names: Erinnys Oustalet 1874 [replaced]
G. †Mesostaphylinus Zhang 1988
+
Mesostaphylinus antiquus Solodovnikov and Yue 2013
Mesostaphylinus elongatus Solodovnikov and Yue 2013
Mesostaphylinus yixianus Solodovnikov and Yue 2013
Tr. Paederini Fleming 1821
+
Subtr. Cryptobiina Casey 1905
Subtr. Paederina Fleming 1821
Subtr. Stilicina Casey 1905
Invalid names: Lithocharina Casey 1905 [empty], Stilicopsina Casey 1905 [empty]
Tr. Pinophilini Nordmann 1837
+
Subtr. Procirrina Bernhauer and Schubert 1912
Invalid names: Pinophilina Nordmann 1837 [empty]
G. †Paroxyporus Cai 2021
+
Subfm. Phloeocharinae Erichson 1839
+
G. †Angucharcotes Li et al. 2022
+
Angucharcotes thayerae Li et al. 2022
G. Charhyphus Sharp 1887
+
Charhyphus balticus Shavrin 2020
Charhyphus serratus Yamamoto and Shavrin 2021
Invalid names: Triga Fauvel 1878 [replaced], Trigites Handlirsch 1907 [synonym]
G. Phloeocharis Mannerheim 1830
+
Phloeocharis agerata Chatzimanolis et al. 2013
Phloeocharis burmana Yamamoto and Newton 2023
Subfm. Piestinae Erichson 1839
+
G. †Eopiestus Cai and Lu 2016
+
Eopiestus groehni Cai and Lu 2016
G. †Paleosiagonium Yue et al. 2016
+
Paleosiagonium adaequatum Yue et al. 2016
Paleosiagonium brevelytratum Yue et al. 2016
G. †Propiestus Yamamoto et al. 2018
+
Propiestus archaicus Yamamoto et al. 2018
G. Siagonium Kirby and Spence 1815
+
Siagonium crassum Giebel 1856
Invalid names: Prognathus Blondel 1827 [synonym]
Subfm. †Protactinae Heer 1847
+
G. †Protactus Heer 1847
+
Protactus erichsonii Heer 1847
Protactus minor Heer 1862
Subfm. Proteininae Erichson 1839
+
Tr. Proteinini Erichson 1839
+
G. Megarthrus Curtis 1829
G. Proteinus Latreille 1796
Tr. †Vetuproteinini Cai et al. 2016
+
G. †Vetuproteinus Cai et al. 2016
Subfm. Protopselaphinae Newton and Thayer 1995
+
G. Protopselaphus Newton and Thayer 1995
+
Protopselaphus newtoni Liu et al. 2020
Protopselaphus thayerae Liu et al. 2020
Subfm. Pselaphinae Latreille 1802
+
Unr. Batrisitae Reitter 1882
+
Tr. Batrisini Reitter 1882
Unr. Clavigeritae Leach 1815
+
Tr. Clavigerini Leach 1815
Tr. †Protoclavigerini Parker and Grimaldi 2014
Invalid names: Tiracerini Besuchet 1986 [empty]
G. †Duterotyrus Schaufuss 1890
Unr. Euplectitae Streubel 1839
+
Tr. Bythinoplectini Schaufuss 1890
Tr. Euplectini Streubel 1839
Tr. Metopiasini Raffray 1904
Tr. Trichonychini Reitter 1882
Tr. Trogastrini Brendel and Wickham 1890
Unr. Faronitae Reitter 1882
+
G. †Cretasonoma Peris et al. 2014
G. Faronus Aubé 1844
G. Sagola Sharp 1874
Invalid names: Faronini Reitter 1882 [invalid subgroup]
Unr. Goniaceritae Reitter 1882
+
Tr. Brachyglutini Raffray 1904
Tr. Bythinini Raffray 1890
Tr. Goniacerini Reitter 1882
+
Invalid names: Goniastini Schaufuss 1872 [replaced]
Tr. Tychini Raffray 1904
G. †Greys Schaufuss 1890
+
Greys conciliator Schaufuss 1890
G. †Penarhytus Peris et al. 2014
+
Penarhytus tenebris Peris et al. 2014
G. †Protrichonyx Parker 2016
+
Protrichonyx rafifrons Parker 2016
G. Pselaphaulax Reitter 1909
+
Unr. Pselaphitae Latreille 1802
+
Tr. Ctenistini Blanchard 1845
Tr. Hybocephalini Raffray 1890
Tr. Pselaphini Latreille 1802
Tr. Tmesiphorini Jeannel 1949
Tr. Tyrini Reitter 1882
Invalid names: Arhytodini Raffray 1890 [empty]
Subfm. Pseudopsinae Ganglbauer 1895
+
G. †Cretaceonanobius Liu et al. 2022
+
Cretaceonanobius fossilis Liu et al. 2022
G. †Cretopseudopsis Liu et al. 2020
+
Cretopseudopsis maweii Liu et al. 2020
G. Pseudopsis Newman 1834
+
Pseudopsis sulcata Newman 1834
Subfm. Scaphidiinae Latreille 1807
+
Tr. Scaphidiini Latreille 1806
+
G. Scaphidium Olivier 1790
G. †Scaphidiopsis Handlirsch 1906
+
Scaphidiopsis aequivoca Weyenbergh 1869
Scaphidiopsis hageni Weyenbergh 1869
Tr. Scaphisomatini Casey 1893
+
G. Scaphisoma Leach 1815
G. †Scaphylinus Kaddumi 2005
+
Scaphylinus ammanicus Kaddumi 2005
Subfm. Scydmaeninae Leach 1815 [antlike stone beetle]
+
Unr. Cephenniitae Reitter 1882
+
Tr. Cephenniini Reitter 1882
Tr. Eutheiini Casey 1897
G. †Cryptodiodon Schaufuss 1890
+
Cryptodiodon corticaroides Schaufuss 1890
G. †Electroscydmaenus Schaufuss 1890
+
Unr. †Hapsomelitae Poinar and Brown 2004
+
G. †Ektatotricha Chatzimanolis et al. 2010
G. †Electroatopos Chatzimanolis et al. 2010
G. †Hapsomela Poinar and Brown 2004
G. †Hetereuthia Schaufuss 1890
+
Hetereuthia elegans Schaufuss 1890
G. †Heuretus Schaufuss 1890
+
Heuretus coriaceus Schaufuss 1890
Unr. Mastigitae Fleming 1821
+
G. †Aenictosoma Schaufuss 1891
Tr. †Baltostigini Jałoszyński and Brunke 2018
Tr. Clidicini Casey 1897
Tr. Leptochromini Jałoszyński and Brunke 2018
Tr. Leptomastacini Casey 1897
Tr. Mastigini Fleming 1821
G. †Palaeomastigus Schaufuss 1890
Invalid names: Clidicus balticus Schaufuss 1896 [nomen dubium], Clidicus cretaceus Kirejtshuk et al. 2015 [nomen dubium], Papusini Jałoszyński and Brunke 2018 [empty]
G. †Microlynx Kohring and Schlüter 1989
+
Microlynx franzbesucheti Kohring and Schlüter 1989
G. †Palaeothia Schaufuss 1890
+
Palaeothia tenuitarsus Schaufuss 1890
G. †Pinitoides Motschulsky 1856
+
Pinitoides scydmaeniformis Motschulsky 1856
Unr. Scydmaenitae Leach 1815
+
Tr. Chevrolatiini Reitter 1882
G. †Kachinus Chatzimanolis et al. 2010
Tr. Scydmaenini Leach 1815
+
Invalid names: Eumicrini Reitter 1882 [synonym]
Tr. Stenichnini Fauvel 1885
+
Invalid names: Cyrtoscydmini Schaufuss 1889 [synonym], Glandulariini Schaufuss 1889 [synonym]
G. †Scydmaenoides Motschulsky 1856
+
Scydmaenoides nigrescens Motschulsky 1856
G. †Semnodioceras Schaufuss 1890
+
Semnodioceras halticaeforme Schaufuss 1890
Invalid names: Mastiginae Fleming 1821 [empty]
G. †Seniaulus von Heyden and von Heyden 1866
+
Seniaulus scaphioides von Heyden and von Heyden 1866
Subfm. Silphinae Latreille 1807
+
G. Aclypea Reitter 1884
+
Aclypea altaica Gebler 1830
+
Invalid names: Blitophaga jacituca Rjabukhin 1990 [synonym], Blitophaga jacutica Rjabukhin 1990 [synonym]
Aclypea bituberosa Le Conte 1859
Aclypea calva Reitter 1885
Aclypea opaca Linnaeus 1758
Aclypea sericea Zubkov 1833
Invalid names: Blitophaga Reitter 1885 [synonym]
G. Dendroxena Motschulsky 1858
+
Dendroxena sexcarinata Motschulsky 1861
Invalid names: Xylodrepa Thomson 1859 [synonym]
Tr. Necrodini Portevin 1926
+
G. Necrodes Leach 1815
G. Oiceoptoma Leach 1815
G. Phosphuga Leach 1817
+
Phosphuga atrata Linnaeus 1758
G. †Pliosilpha Gersdorf 1969
+
Pliosilpha strausi Gersdorf 1969
Tr. Silphini Latreille 1807
+
G. Necrophila Kirby and Spence 1828
G. Silpha Linnaeus 1758
G. Thanatophilus Leach 1815
G. †Sinostaphylina Hong and Wang 1990
+
Sinostaphylina nanligezhuangensis Hong and Wang 1990
G. †Sinostaphylius Hong 1992
+
Subfm. Solieriinae Newton and Thayer 1992
+
G. †Prosolierius Thayer et al. 2012
+
Prosolierius crassicornis Thayer et al. 2012
Prosolierius mixticornis Thayer et al. 2012
Prosolierius parvus Peris et al. 2014
Prosolierius tenuicornis Thayer et al. 2012
Prosolierius thayerae Yamamoto 2023
Subfm. Staphylininae Latreille 1802
+
Tr. Arrowinini Solodovnikov and Newton 2005
+
G. †Paleowinus Solodovnikov and Yue 2013
G. †Cretoprosopus Solodovnikov and Yue 2013
+
Cretoprosopus problematicus Solodovnikov and Yue 2013
Tr. Diochini Casey 1906
+
G. Diochus Erichson 1839
G. †Megolisthaerus Solodovnikov and Yue 2010
+
Megolisthaerus chinensis Solodovnikov and Yue 2010
Megolisthaerus minor Cai and Huang 2013
Tr. Othiini Thomson 1859
+
G. Atrecus Jacquelin du Val 1856
G. †Vetatrecus Kypke et al. 2018
G. †Paleothius Solodovnikov and Yue 2013
+
Paleothius gracilis Solodovnikov and Yue 2013
Tr. Staphylinini Latreille 1802
+
Subtr. Acylophorina Outerelo and Gamarra 1985
Subtr. Algonina Schillhammer and Brunke 2018
Subtr. Amblyopinina Seevers 1944
Subtr. Anisolinina Hayashi 1993
Subtr. †Baltognathina Brunke et al. 2019
G. †Cretoquedius Ryvkin 1988
Subtr. Cyrtoquediina Brunke and Solodovnikov 2016
G. †Durothorax Solodovnikov and Yue 2013
Subtr. Hyptiomina Casey 1906
Subtr. Philonthina Kirby 1837
Subtr. Quediina Kraatz 1857
Subtr. Staphylinina Latreille 1802
Invalid names: Quediini Horn 1878 [synonym], Xanthopygina Sharp 1884 [empty]
Tr. †Thayeralinini Solodovnikov and Yue 2013
+
G. †Thayeralinus Solodovnikov and Yue 2013
Tr. Xantholinini Erichson 1839
+
G. Gyrohypnus Leach 1819
G. Leptacinus Erichson 1839
G. Neohypnus Coiffait and Saiz 1964
G. Neoxantholinus Cameron 1945
G. Xantholinus Dejean 1821
Invalid names: Platyprosopinae Lynch Arribálzaga 1884 [synonym], Platyprosopini Lynch Arribálzaga 1884 [empty], Xantholininae Erichson 1839 [synonym]
G. †Staphylinites Scudder 1876
+
Staphylinites obsoletum Scudder 1876
Subfm. Steninae Macleay 1825
+
G. †Afristenus Mnguni et al. 2022
+
Afristenus orapensis Mnguni et al. 2022
G. Dianous Leach 1819
+
Dianous nitidulus Le Conte 1849
G. †Eocenostenus Cai et al. 2014
+
Eocenostenus fossilis Cai et al. 2014
G. †Festenus Żyła et al. 2017
+
Festenus gracilis Żyła et al. 2017
Festenus robustus Żyła et al. 2017
G. Stenus Latreille 1796
+
Subg. Stenus (Hemistenus) Motschulsky 1860
+
Invalid names: Stenus (Parastenus) Heyden 1905 [synonym]
Subg. Stenus (Nestus) Rey 1883
Subg. Stenus (Tesnus) Rey 1884
Stenus abraham Puthz 2010
Stenus archetypus Puthz 2010
Stenus atavus Puthz 2010
Stenus avus Puthz 2010
Stenus biguttatus Linnaeus 1758
Stenus brunnipes Stephens 1833
Stenus comma LeConte 1863
Stenus groehni Puthz 2010
Stenus gypsi Oustalet 1874
Stenus juno Paykull 1789
Stenus ketura Puthz 2010
Stenus methusalem Puthz 2010
Stenus morsei Scudder 1900
Stenus noach Puthz 2010
Stenus ornatus Förster 1891
Stenus pinguis Casey 1884
Stenus prodromus Heer 1856
Stenus scribai von Heyden and von Heyden 1866
Stenus scrupeus Casey 1884
Stenus sordidus Puthz 1988
Invalid names: Areus Casey 1884 [synonym]
G. †Sulcelytrinus Tikhomirova 1968
+
Sulcelytrinus antiquus Tikhomirova 1968
Subfm. Tachyporinae MacLeay 1825
+
G. †Abscondus Tikhomirova 1968
+
Abscondus regularis Tikhomirova 1968
Abscondus similis Tikhomirova 1968
Tr. Tachinusini Fleming 1821
+
G. †Hesterniasca Zhang et al. 1992
G. †Mesotachinus Tikhomirova 1968
G. Nitidotachinus Campbell 1993
G. †Protachinus Cai et al. 2013
G. Tachinus Gravenhorst 1802
+
Invalid names: Elliptoma Motschulsky 1845 [synonym]
Tr. Tachyporini MacLeay 1825
+
Subtr. Euconosomatina Cameron 1918
Subtr. Tachyporina MacLeay 1825
G. †Tachyporoides Tikhomirova 1968
+
Tachyporoides villosus Tikhomirova 1968
Tr. Vatesini Seevers 1958
+
G. Coproporus Kraatz 1857
G. †Procileoporus Yamamoto 2016
Invalid names: Acylophorus immotus Scudder 1900 [nomen dubium]
Subfm. Trichophyinae Thomson 1858
+
G. Trichophya Mannerheim 1830
+
Trichophya minor Yamamoto and Newton 2021
Subfm. Trigonurinae Reiche 1821
+
G. †Abolescus Tikhomirova 1968
+
Abolescus glabratus Tikhomirova 1968
G. †Kovalevia Ryvkin 1990
+
Kovalevia onokhoica Ryvkin 1990
G. †Trigunda Ryvkin 1990
+
Trigunda lata Ryvkin 1990
G. †Tunicopterus Tikhomirova 1968
+
Tunicopterus sigara Tikhomirova 1968
Invalid names: Empelinae Newton and Thayer 1992 [empty], Habrocerinae Mulsant and Rey 1877 [empty], Leptotyphlinae Fauvel 1874 [empty], Microsilphinae Crowson 1950 [empty], Neophoninae Fauvel 1905 [empty], Silphidae Latreille 1807 [synonym], Triga coeni Scudder 1900 [nomen dubium]
Invalid names: Dasyceridae Reitter 1887 [empty], Empelidae [empty]
Invalid names: Scarabaeiformia Crowson 1960 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Angulata to the top of the Holocene or 201.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 196.5 Ma

Collections (615 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Angulata201.3 - 196.5Switzerland Aphodiites protogaeus (131766)
Pliensbachian190.8 - 182.7Russian Federation (Kemerovo) Ochtebiites minor (125341)
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) Hexameristus inflatus, Opiselleipon gravis (105002) Spalacoides simplex (107555) Streblocardioides striatus, Aposphinctus conservatus (107343)
Toarcian - Aalenian182.7 - 170.3Russian Federation (Kemerovo) Hydrophilidae indet. (117989)
Bajocian - Bathonian170.3 - 166.1China (Anhui) Mesagyrtes lintouensis (106695)
Bajocian - Bathonian170.3 - 166.1Mongolia (Uver-Khangay Aymag) Jurahybosorus mongolicus (118011)
Bajocian - Bathonian170.3 - 166.1Russian Federation (Krasnoyarsk) Morsum abdominale (118010)
Bathonian168.3 - 166.1Mongolia Hydrophilidae indet. (119621)
Callovian - Oxfordian166.1 - 157.3China (Liaoning) Protostaphylinus mirus (151526)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Abolescus glabratus, Mesotachinus major, Globoides microps, Archodromus brachypterus, Archodromus comptus, Mesoxytelus mandibularis, Tunicopterus sigara (105970) Jurahybosorus kazakhstanicus, Abolescus glabratus, Mesotachinus major, Mesotachinus minor, Abscondus similis, Porrhodromus communis, Archodromus brachypterus, Mesoxytelus parvus (105971) Paraspercheus asiaticus, Abolescus glabratus, Mesotachinus major, Mesotachinus medius, Abscondus regularis, Tachyporoides villosus, Globoides oculatus, Archodromus brachypterus, Sulcelytrinus antiquus (106101) Paraspercheus asiaticus, Ochtebiites incertus (105426) Protohybosorus grandissimus (112846) Protohybosorus karatavicus (113439) Protohybosorus mesoasiaticus, Abolescus glabratus, Mesotachinus major, Archodromus comptus, Mesoxytelus mandibularis (106102)
Callovian - Oxfordian166.1 - 157.3China (Hebei) Prostaphylinus mirus (154824)
Callovian - Oxfordian166.1 - 157.3China (Nei Mongol) Alloioscarabaeus cheni (113366) Juraesalus atavus, Juroglypholoma antiquum, Sinanthobium daohugouense, Mesapatetica aenigmatica, Protolisthaerus jurassicus, Silphidae indet. (113371) Mesochodaeus daohugouensis (113365)
Oxfordian163.5 - 157.3Russian Federation (Buryat) Ochtebiites altus, Mesagyrtes communis, Eophyllodrepa propinqua, Anicula inferna (105017)
Oxfordian163.5 - 157.3Russian Federation Mesagyrtes communis (123219)
Oxfordian - Kimmeridgian163.5 - 152.1China (Yunnan) Hydrophilidae indet. (164859)
Tithonian152.1 - 145.0Australia (New South Wales) Protochares brevipalpis, Juroglypholoma talbragarense, Protachinus minor, Olisthaerinae indet. (128121)
Tithonian152.1 - 145.0Russian Federation (Chita) Aposphinctus sibiricus (152172) Mesohelophorus elongatus, Aposphinctus sibiricus (125407) Mesosperchus tarsalis (119424 152174) Mesosperchus tarsalis, Mesohelophorus elongatus, Hydrophilopsia longitarsalis (114307) Mesosperchus tarsalis, Trigunda lata, Undiatina pilosa (123197) Staphylinidae indet., Prodaia inflata, Daiodromus umbrosus, Morda mora, Cuneocharis elongatus, Mesoporus gracilis (119397)
Tithonian152.1 - 145.0Mongolia Helophorus inceptivus, Prospercheus cristatus (131803) Helophorus spp., Helophorus costalis, Paralucanus mesozoicus, Hydraenidae indet. (104748) Helophorus spp., Helophorus inceptivus, Helophorus costalis, Prospercheus cristatus, Hydraenidae indet. (121394) Mesagyrtoides fulvus (152133)
Tithonian152.1 - 145.0Russian Federation Mesosperchus tarsalis (125408 138150)
Early/Lower Tithonian150.8 - 145.0Germany (Bayern) Geotrupes lithographicus (133578) Mesosperchus schultzi (123144) Scaphidium hageni, Cassida aequivoca (123906)