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Hydrophilidae (water scavenger beetle)

Insecta - Coleoptera - Hydrophilidae

Synonyms
Synonymy list
YearName and author
1802Hydrophilidae Latreille p. 136
1831Sphaeridium melanarium Gistl p. 247
1847Escheria Heer p. 57
1847Escheria ovalis Heer p. 57 figs. Pl 7, fig 23
1862Escheria bella Heer p. 69 figs. Pl 5, fig 17
1864Escheria bella Heer p. 381 fig. 273
1880Escheriidae Kolbe p. 266
1885Escheria bella Scudder p. 802 fig. 1063
1916Hydrophilidae Tillyard
1918Hydrophilidae Tillyard
1923Hydrophilidae Tillyard
1981Hydrophilidae Crowson p. 694
1985Mesosperchus schultzi Ponomarenko p. 142 figs. 6; Pl 2 fig 3
1992Escheria Carpenter
1992Hydrophilidae Carpenter
1995Hydrophilidae Lawrence and Newton, Jr.
2010Hydrophilidae Fikáček et al.
2011Hydrophilidae Bouchard et al. p. 153
2016Hydrophilidae Webster
2017Hydrophilidae Fikáček et al. p. 50
2022Hydrophilidae Cai et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
RankNameAuthor
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderPolyphagaEmery 1886
infraorderStaphyliniformia
superfamilyHydrophiloidea
familyHydrophilidae
familyHydrophilidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Hydrophilidae Latreille 1802 [water scavenger beetle]
show all | hide all
Subfm. Acidocerinae Zaitzev 1908
+
G. Helochares Mulsant 1846
+
†Helochares fog Arriaga-Varela et al. 2021
†Helochares mydaleus Zhang 1989
G. †Hydroyixia Fikáček et al. 2014
+
†Hydroyixia elongata Fikáček et al. 2014
†Hydroyixia latissima Fikáček et al. 2014
Invalid names: Horelophopsinae Hansen 1997 [synonym]
G. †Alegorius Fikáček et al. 2014
+
†Alegorius yixianus Fikáček et al. 2014
G. †Aposphinctus Bode 1953
+
+
Invalid names: Spalacoides simplex Bode 1953 [synonym]
†Aposphinctus sibiricus Ponomarenko 1990
†Aposphinctus striatus Bode 1953
+
Invalid names: Hexameristus inflatus Bode 1953 [synonym]
Invalid names: Hexameristus Bode 1953 [synonym], Spalacoides Bode 1953 [synonym], Streblocardioides Bode 1953 [synonym]
Subfm. Chaetarthriinae Bedel 1881
+
Tr. Anacaenini Hansen 1991
+
G. Anacaena Thomson 1859
+
†Anacaena morla Arriaga-Varela et al. 2021
†Anacaena paleodominica Fikáček and Engel 2011
G. Crenitis Bedel 1881
+
†Crenitis profechuyi Arriaga-Varela et al. 2021
G. †Cretocrenis Fikáček et al. 2017
+
†Cretocrenis burmanicus Fikáček et al. 2017
Invalid names: Horelophinae Hansen 1991 [synonym]
Tr. Chaetarthriini Bedel 1881
+
G. Chaetarthria Stephens 1835
+
G. †Creniphilites Wickham 1913
+
†Creniphilites orpheus Wickham 1913
G. †Cretosperchus Ponomarenko 1997
+
†Cretosperchus medievalis Ponomarenko 1997
Subfm. Enochrinae Short and Fikáček 2013
+
G. Cymbiodyta Bedel 1881
+
†Cymbiodyta exstincta Scudder 1900
†Cymbiodyta samueli Fikáček et al. 2023
G. Enochrus Thomson 1859
+
†Enochrus innovatus Theobald 1937
†Enochrus lapsus Zhang 1989
†Enochrus pleistocenicus Lomnicki 1894
†Enochrus primaevus Scudder 1876
†Enochrus scudderi Wickham 1909
†Enochrus striatus Theobald 1937
Invalid names: Philydrus Solier 1834 [replaced]
Subfm. Georissinae Laporte 1819
+
G. †Georissites Ponomarenko 2008
+
†Georissites magnus Haupt 1956
†Georissites negev Ponomarenko 2008
G. Georissus Latreille 1809
+
G. Hydrocharis Berthold 1827
Subfm. Hydrochinae Thomson 1859
+
G. Hydrochus Germar 1824
+
†Hydrochus amictus Scudder 1890
Hydrochus elongatus Schaller 1783
†Hydrochus relictus Scudder 1890
Subfm. Hydrophilinae Latreille 1802
+
Tr. Berosini Mulsant 1844
+
G. Berosus Leach 1817
+
Subg. Berosus (Berosus) Leach 1817
+
†Berosus (Berosus) morticinus von Heyden and von Heyden 1866
+
Invalid names: Berosus capitatus Statz 1939 [synonym]
†Berosus barclayi Ponomarenko and Soriano 2019
Berosus luridus Linnaeus 1761
†Berosus sexstriatus Scudder 1878
Berosus signaticollis Charpentier 1825
Berosus spinosus Steven 1808
†Berosus tenuis Scudder 1878
Tr. Hydrobiusini Mulsant 1844
+
G. †Baissalarva Fikáček et al. 2014
+
†Baissalarva hydrobioides Fikáček et al. 2014
G. †Cretoxenus Fikáček et al. 2014
+
†Cretoxenus australis Fikáček et al. 2014
G. Hydrobius Linnaeus 1758
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†Hydrobius confixus Scudder 1890
Hydrobius convexus Brullé 1835
†Hydrobius couloni Heer 1862
†Hydrobius decineratus Scudder 1878
Hydrobius fuscipes Linnaeus 1758
†Hydrobius longicollis Heer 1847
†Hydrobius maceratus Scudder 1900
†Hydrobius prisconatator Wickham 1911
†Hydrobius titan Wickham 1913
G. Limnoxenus Motshulsky 1853
+
†Limnoxenus olenus Fikáček et al. 2010
Invalid names: Sperchopsini Hansen 1991 [synonym]
Tr. Hydrophilini Latreille 1802
+
Invalid names: Hydrophilina Latreille 1802 [invalid subgroup], Hydrophilopsis Heer 1862 [nomen dubium], Hydrophilopsis elongata Heer 1862 [nomen dubium], Hydrous ovalis Heer 1862 [nomen dubium]
Tr. Laccobiini Houlbert 1922
+
G. Laccobius Erichson 1837
+
Laccobius ellipticus Le Conte 1855
†Laccobius elongatus Scudder 1878
†Laccobius flachii Lomnicki 1894
Laccobius minutus Linnaeus 1758
†Laccobius priscus Oustalet 1870
†Laccobius vetustus Oustalet 1874
G. Paracymus Thomson 1867
+
†Paracymus excitatus von Heyden and von Heyden 1866
Invalid names: Amphiopini Kuwert 1890 [empty], Cymbiodyta austerus Statz 1939 [nomen dubium]
G. †Orphnospercheus Lin 1992
+
G. †Paraspercheus Ponomarenko 1977
+
†Paraspercheus asiaticus Ponomarenko 1977
†Paraspercheus vitimensis Ponomarenko 1977
G. †Penitus Zhang et al. 1994
+
†Penitus gemellus Zhang et al. 1994
G. Philhydrus Solier 1834
+
†Philhydrus scudderi Wickham 1909
G. †Prospercheus Prokin 2009
+
†Prospercheus cristatus Prokin 2009
G. †Protochares Fikáček et al. 2014
+
†Protochares brevipalpis Fikáček et al. 2014
G. Regimbartia Zaitzev 1908
+
Regimbartia attenuata Fabricius 1801
Subfm. Rygmodinae Orchymont 1916
+
Invalid names: Andotypini Hansen 1991 [synonym], Borborophorini Hansen 1991 [synonym], Tormissini Hansen 1991 [synonym]
Subfm. Sphaeridiinae Latreille 1802
+
Tr. Coelostomatini Heyden 1891
+
G. Coelostoma Brullé 1835
+
Coelostoma orbiculare Fabricius 1775
Coelostoma stultum Walker 1858
Tr. Megasternini Mulsant 1844
+
G. Cercyon Leach 1817
+
Cercyon analis Paykull 1798
Cercyon herceus Smetana 1978
Cercyon luniger Mannerheim 1853
Cercyon marinus Thomson 1853
Cercyon melanocephalus Linnaeus 1758
Cercyon pygmaeus Illiger 1801
Cercyon sternalis Sharp 1918
†Cercyon terrigena Scudder 1879
Cercyon tristis Illiger 1801
Cercyon ustulatus Preyssler 1790
G. Cryptopleurum Mulsant 1844
+
Cryptopleurum crenatum Kugelann 1794
Cryptopleurum minutum Fabricius 1775
G. Megasternum Mulsant 1844
+
Megasternum obscurum Marsham 1802
Tr. Sphaeridiini Latreille 1802
+
G. Sphaeridium Fabricius 1775
+
Invalid names: Omicrini Smetana 1975 [empty], Protosternini Hansen 1991 [empty]
G. Sternolophus Solier 1834
+
Sternolophus rufipes Fabricius 1792
Invalid names: Escheria Heer 1847 [nomen dubium], Escheria bella Heer 1862 [nomen dubium], Escheria ovalis Heer 1847 [nomen dubium], Escheriidae Kolbe 1880 [synonym], Mesosperchus schultzi Ponomarenko 1985 [nomen dubium], Spercheinae Erichson 1837 [empty], Sphaeridium melanarium Gistl 1831 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Toarcian to the top of the Holocene or 182.70000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 182.0 Ma

Collections (148 total)


Time interval Ma Country or state Original ID and collection number
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) Hexameristus inflatus (105002) Spalacoides simplex (107555) Streblocardioides striatus, Aposphinctus conservatus (107343)
Toarcian - Aalenian182.7 - 170.3Russian Federation (Kemerovo) Hydrophilidae indet. (117989)
Bathonian168.3 - 166.1Mongolia Hydrophilidae indet. (119621)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Paraspercheus asiaticus (105426 106101)
Oxfordian - Kimmeridgian163.5 - 152.1China (Yunnan) Hydrophilidae indet. (164859)
Tithonian152.1 - 145.0Australia (New South Wales) Protochares brevipalpis (128121)
Tithonian152.1 - 145.0Mongolia Prospercheus cristatus (121394 131803)
Tithonian152.1 - 145.0Russian Federation (Chita) Aposphinctus sibiricus (125407 152172)
Early/Lower Tithonian150.8 - 145.0Germany (Bayern) Mesosperchus schultzi (123144)
Berriasian145.0 - 139.8Israel Cretosperchus medievalis (84472)
Hauterivian132.9 - 129.4Russian Federation (Chita) Aposphinctus sibiricus (125409)
Late/Upper Barremian130.0 - 125.45China (Nei Mongol) Alegorius yixianus (125655)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Alegorius sp., Alegorius yixianus, Hydroyixia elongata, Hydroyixia latissima (111712) Alegorius yixianus, Hydroyixia elongata, Hydroyixia latissima (111716)
Aptian125.0 - 113.0China (Jilin) Orphnospercheus lonjingensis (172391)
Aptian125.0 - 113.0Russian Federation (Buryatia) Paraspercheus vitimensis, Baissalarva hydrobioides (105969)
Aptian125.0 - 113.0China (Nei Monggol) Hydrophilidae indet. (152245)
Late/Upper Aptian122.46 - 113.0Brazil (Ceará) Hydrophilidae indet. (122557)
Late/Upper Aptian122.46 - 113.0Australia (Victoria) Hydrophilidae indet., Hydrophilidae larva indet., Cretoxenus australis (77192)
Albian113.0 - 100.5Jordan Hydrophilidae indet. (124847)
Early/Lower Cenomanian99.6 - 93.5Myanmar Cretocrenis burmanicus (161726)
Early/Lower Cenomanian99.6 - 93.5Myanmar (Kachin) Cretocrenis burmanicus (221886)
Middle Turonian93.5 - 89.3Israel Georissites negev (156985)
Danian - Selandian66.0 - 59.2Svalbard and Jan Mayen Elytridium undecimstriatum (122011)
Ypresian56.0 - 47.8Canada (British Columbia) Cercyon terrigena (122190)
Bridgerian50.3 - 46.2USA (Wyoming) Hydrochus relictus, Hydrobius confixus (122090) Laccobius elongatus, Philhydrus primaevus (121306) Tropisternus saxialis, Tropisternus sculptilis, Berosus tenuis, Berosus sexstriatus, Philhydrus sp., Hydrobius decineratus (121322)
MP 1148.6 - 40.4Germany (Hessen) Hydrobiomorpha eopalpalis (165522) Hydrophilus sp., Hydrobiomorpha eopalpalis (120036) Hydrophilus sp., Hydrobiomorpha sp. 1, Hydrochara sp., Hydrophilina indet., Hydrobiomorpha eopalpalis (120798)
Lutetian47.8 - 41.3Germany (Sachsen-Anhalt) Georyssus magnus (114060)
Priabonian38.0 - 33.9Poland Crenitis profechuyi, Helochares fog (110737)
Priabonian38.0 - 33.9United Kingdom (Isle of Wight) Berosus barclayi (156628)
Priabonian38.0 - 33.9France Enochrus striatus (113709)
Priabonian38.0 - 33.9Russian Federation Anacaena morla (123419)
Priabonian38.0 - 33.9Lithuania Cymbiodyta samueli (230326)
Priabonian38.0 - 33.9United Kingdom (England) Berosus sp., Hydrochara woodwardi, Berosus barclayi (123959) Hydrochara woodwardi, Berosus barclayi (21976 123960)
Chadronian37.2 - 33.9USA (Colorado) Hydrobius prisconatator (106381 123119) Hydrobius titan, Creniphilites orpheus (113749) Philydrus scudderi (124002) Tropisternus limitatus, Tropisternus vanus, Philhydrus scudderi, Hydrobius titan, Hydrocharis extricatus, Hydrobius maceratus (106384)
MP 2333.9 - 28.4France Hydrophilidae indet., Philydrus innovatus, Laccobius vetustus (122535)
Rupelian33.9 - 28.1Germany Hydrobius convexus (122407)
Rupelian33.9 - 28.1France (Auvergne) Laccobius priscus (127630)
Late/Upper Oligocene28.4 - 23.03France (Puy-de-Dome) Hydrophilus magnificus (155204)
MP 3028.4 - 23.03Germany (Nordrhein-Westfalen) Hydrophilus fraternus, Hydrous miserandus (127628)
MP 3028.4 - 23.03Germany Coccinella protogaeae (115802) Philydrus morticinus, Laccobius excitatus, Hydrous neptunus (123945)
Chattian28.1 - 23.03Brazil Hydrophilidae indet. (153054)
Chattian28.1 - 23.03France (Bouches-Du-Rhone) Laccobius vetustus, Hydrophilus antiquus (123944) Limnoxenus olenus (127415)
Chattian28.1 - 23.03Germany (North Rhine-Westphalia) Berosus capitatus, Paracymus excitatus, Hydrophilus fraternus, Hydrophilus neptunus, Hydrous rottensis, Cymbiodyta austerus, Berosus morticinus (122010)
Chattian28.1 - 23.03France (Provence-Alpes-Côte d'Azur) Hydrophilopsis incerta (122487)
Chattian - Aquitanian28.1 - 20.44Germany (Sachsen) Hydrobiomorpha enspelense (26765)
Aquitanian23.03 - 20.44Switzerland Hydrophilus gaudini, Hydrophilus ruminianus (154053)
Aquitanian23.03 - 20.44Germany Hydrous sp., Hydrophilus sp. (123872)
Aquitanian23.03 - 20.44New Zealand Hydrophilidae indet. (50035)
Aquitanian - Burdigalian23.03 - 15.97Russian Federation Hydrophilidae indet. (117611)
Early/Lower Miocene23.03 - 15.97Mexico Anacaena sp. (21977)
Burdigalian20.44 - 15.97China (Shandong) Enochrus lapsus, Helochares mydaleus, Hydrous pergrandis (128552) Penitus gemellus (138945)
Burdigalian20.44 - 15.97Greece Hydrophilus vexatorius (144526)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Anacaena paleodominica (122982)
MN 320.0 - 16.9Czech Republic Hydrophilus pistaceus (105508)
MN 515.97 - 13.65Germany (Baden-Württemberg) Hydrophilidae indet. (109058)
Badenian13.65 - 12.7Austria (Steiermark) Hydrophilus carbonarius (115306)
Sarmatian12.7 - 11.608Croatia Hydrobius longicollis (117246)
Sarmatian12.7 - 11.608Germany (Wurttemberg) Hydrophilus kuntzeni, Hydrophilus caraboides (113752)
Sarmatian12.7 - 11.608Germany (Baden-Württemberg) Hydrobiomorpha heeri (144245) Hydrophilus rehmanni (117338) Hydrophilus spectabilis (172253) Hydrophilus spectabilis, Hydrous escheri, Hydrobiomorpha heeri, Hydrophilus giganteus, Hydrophilopsis elongata, Hydrous ovalis, Escheria bella, Hydrous rehmanni, Hydrous braunii, Hydrobius couloni (117252) Hydrophilus vexatorius, Hydrophilus knorrii, Hydrophilus braunii (115305) Hydrophilus vexatorius, Hydrophilus spectabilis, Hydrophilus noachicus, Escheria ovalis (115304)
Late/Upper Miocene11.608 - 5.333Russian Federation Hydrophilidae indet., Coelostoma orbiculare (113713)
Messinian7.246 - 5.333USA (Alaska) Georyssus sp. (123352)
Zanclean5.333 - 3.6Japan Coelostoma sp., Regimbartia attenuata (165737)
Pliocene5.333 - 2.588Japan Coelostoma sp., Hydrochara libera, Sternolophus rufipes (165744) Coelostoma sp., Regimbartia attenuata (165743)
Piacenzian3.6 - 2.588USA (Alaska) Georyssus sp. (162796)
Piacenzian3.6 - 2.588Japan Coelostoma sp. (165742) Hydrophilidae indet., Hydrophilus sp., Hydrochara sp., Coelostoma sp., Coelostoma stultum, Regimbartia attenuata, Sternolophus rufipes (165738)
Gelasian2.588 - 1.806Greenland Laccobius sp. (160109) Megasternum obscurum (160166)
Gelasian2.588 - 1.806France Georyssus nigrobrunneus (113996)
Early/Lower Pleistocene2.588 - 0.781Japan Coelostoma sp. (153617)
Early/Lower Pleistocene2.588 - 0.781Russian Federation Hydrophilidae indet. (163089) Laccobius sp., Cercyon sp. (163069)
Calabrian1.8 - 0.774Japan Hydrochara sp., Coelostoma orbiculare (160539)
Chibanian0.774 - 0.129Japan (Kumamoto) Coelostoma sp. (164901)
Chibanian0.774 - 0.129Japan Hydrophilidae indet. (165705) Hydrophilidae indet., Coelostoma sp. (165704) Hydrophilidae indet., Hydrophilus sp., Hydrochara sp., Coelostoma sp., Hydrochara libera (165700)
Middle Pleistocene0.774 - 0.129United Kingdom Coelostoma orbiculare, Sphaeridium scarabaeoides, Hydrophilus caraboides, Cercyon melanocephalus, Georissus crenulatus, Megasternum boletophagum (168267) Enochrus sp., Cercyon sp., Cercyon sternalis (168272) Enochrus sp., Coelostoma orbiculare, Hydrobius fuscipes, Sphaeridium scarabaeoides, Cercyon melanocephalus, Megasternum boletophagum, Chaetarthria seminulum (168266) Enochrus sp., Georissus crenulatus, Hydrochus elongatus, Cercyon pygmaeus, Cercyon sternalis (168270) Enochrus sp., Laccobius sp., Coelostoma orbiculare, Cercyon sp., Hydrobius fuscipes, Cercyon ustulatus, Cercyon sternalis, Megasternum boletophagum, Chaetarthria seminulum (168264) Enochrus spp., Laccobius sp., Coelostoma orbiculare, Cercyon sp., Sphaeridium scarabaeoides, Georissus crenulatus, Megasternum boletophagum, Chaetarthria seminulum (168265) Hydrochus elongatus, Cercyon sternalis, Megasternum boletophagum (168271) Laccobius sp., Coelostoma orbiculare, Cryptopleurum minutum, Cercyon analis, Megasternum boletophagum (168268) Paracymus sp., Coelostoma orbiculare, Hydrobius fuscipes, Cercyon analis (168269)
Middle Pleistocene0.774 - 0.129United Kingdom (Warwickshire) Cercyon sp. (169389) Cercyon sp., Cryptopleurum crenatum (169391)
Middle Pleistocene0.774 - 0.129Japan (Kumamoto) Coelostoma sp. (165716)
Late/Upper Pleistocene0.129 - 0.0117USA (Colorado) Enochrus hamiltoni (169353) Enochrus sp. (169350) Hydrobius fuscipes (169333 169336)
Late/Upper Pleistocene0.129 - 0.0117Ukraine Hydrobius fuscipes, Philhydrus pleistocenicus, Laccobius flachii, Sphaeridium scarabaeoides (153476)
Late/Upper Pleistocene0.129 - 0.0117Canada (Ontario) Cymbiodyta exstincta (152577)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation (Astrakhan) Hydrophilidae indet., Enochrus sp., Hydrochara sp., Anacaena sp., Cercyon marinus, Cercyon tristis, Hydrochara caraboides, Hydrochara flavipes, Berosus luridus, Berosus spinosus, Enochrus quadripunctatus, Berosus signaticollis (125794)
Late/Upper Pleistocene0.129 - 0.0117USA (Pennsylvania) Hydrophilidae indet. (137207)
Late/Upper Pleistocene0.129 - 0.0117USA (California) Hydrophilus sp. (44954)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (England) Cercyon tristis (164904 164906 164908 164921) Cercyon tristis, Hydrobius fascipes (164905 164920)
Late/Upper Pleistocene0.129 - 0.0117Japan Coelostoma sp., Coelostoma orbiculare, Hydrochara libera (166082) Hydrophilidae indet. (164902)
Late/Upper Pleistocene0.129 - 0.0117USA (Wisconsin) Cercyon sp., Cercyon herceus (169406)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation Hydrobius fuscipes (153435 163102 173216)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Worcestershire) Cercyon spp., Hydrobius fuscipes, Sphaeridium sp., Cercyon tristis, Cercyon melanocephalus, Cercyon ustulatus, Cryptopleurum minutum, Laccobius minutus, Georissus crenulatus (163070) Hydrobius fuscipes, Enochrus quadripunctatus (163075)
Late/Upper Pleistocene0.129 - 0.0117USA (Ohio) Hydrochus amictus (152574)
Late/Upper Pleistocene0.129 - 0.0117USA (Washington) Hydrobius fuscipes, Laccobius ellipticus, Cercyon luniger (167662) Hydrophilidae indet. (169375 169378 169379) Laccobius sp., Cymbiodyta sp., Cercyon sp., Georissus sp., Hydrobius fuscipes (157092)
Late/Upper Pleistocene0.129 - 0.0117Peru Hydrophilus sp., Gyrinidae indet. (71346)
Tarantian0.129 - 0.0117USA (Pennsylvania) Hydrophilidae indet., Enochrus sp. (137204 137205 137206)
Holocene0.0117 - 0.0Brazil Sphaeridium melanarium (155717)