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Staphyliniformia
Taxonomy
Staphyliniformia was named by Lameere (1900) [rank=series]. It is extant.
It was assigned to Polyphaga by Crowson (1981), Lawrence and Newton (1995), Bouchard et al. (2011) and Cai et al. (2022).
It was assigned to Polyphaga by Crowson (1981), Lawrence and Newton (1995), Bouchard et al. (2011) and Cai et al. (2022).
Subtaxa
Synonyms
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Synonymy list
Year | Name and author |
---|---|
1900 | Staphyliniformia Lameere |
1960 | Scarabaeiformia Crowson |
1981 | Staphyliniformia Crowson p. 694 |
1981 | Scarabaeiformia Crowson p. 695 |
1995 | Scarabaeiformia Lawrence and Newton, Jr. |
1995 | Staphyliniformia Lawrence and Newton, Jr. |
2011 | Scarabaeiformia Bouchard et al. |
2011 | Staphyliniformia Bouchard et al. p. 153 |
2022 | Staphyliniformia Cai et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Infraor. Staphyliniformia Lameere 1900
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Superfm. Histeroidea Gyllenhal 1808
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Fm. †Cretohisteridae Zhou et al. 2018
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Fm. Histeridae Gyllenhal 1808 [clown beetle]
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Subfm. Abraeinae MacLeay 1819
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G. †Amplectister Caterino and Maddison 2018
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†Amplectister tenax Caterino and Maddison 2018
†Amplectister terapoides Yamamoto and Caterino 2022
Subfm. †Antigracilinae Zhou et al. 2020
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Subfm. Dendrophilinae Reitter 1909
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G. †Yethiha Caterino 2021
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†Yethiha peregrina Caterino 2021
†Yethiha pubescens Simon Pražák and Lackner 2023
Subfm. Haeteriinae Marseul 1857
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Subfm. Histerinae Gyllenhal 1808
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G. †Platycretus Simon Pražák and Lackner 2023
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†Platycretus muscularis Simon Pražák and Lackner 2023
Subfm. Onthophilinae MacLeay 1819
+
G. †Carinumerus Caterino 2021
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†Carinumerus intricatus Caterino and Yamamoto 2023
†Carinumerus maddisoni Caterino 2021
†Carinumerus yingae Jiang et al. 2020
G. Onthophilus Leach 1817
+
†Onthophilus intermedius Handschin 1944
Onthophilus striatus Forster 1771
G. †Phasmister Caterino 2021
+
†Phasmister cristatus Caterino 2021
†Phasmister hkamticus Caterino and Yamamoto 2023
†Phasmister parallelus Caterino and Yamamoto 2023
Subfm. Trypanaeinae Marseul 1857
+
Invalid names: Chlamydopsinae Bickhardt 1914 [empty], Hetaeriinae Marseul 1857 [empty], Niponiinae Fowler 1912 [empty], Tribalinae Bickhardt 1914 [empty], Trypeticinae Bickhardt 1914 [empty]
Invalid names: Sphaeritidae Shuckard 1839 [empty]
Superfm. Hydrophiloidea Latreille 1802 [water scavenger beetle]
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Fm. Helophoridae Leach 1815
+
G. Helophorus Fabricius 1775
+
Subg. Helophorus (Atracthelophorus) Kuwert 1890
+
Helophorus (Atracthelophorus) aspericollis Angus 1973
Helophorus (Atracthelophorus) glacialis Villa and Villa 1833
Subg. Helophorus (Gephelophorus) Sharp 1915
+
Helophorus (Gephelophorus) sibiricus Motschulsky 1860
+
Invalid names: Helophorus (Gephelophorus) lomnicki d'Orchymont 1927 [synonym], Helophorus (Trichelophorus) pleistocenicus Lomnicki 1894 [synonym]
Subg. Helophorus (Meghelophorus) Kuwert 1886
+
Helophorus (Meghelophorus) aquaticus Linnaeus 1758
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Invalid names: Helophorus (Praehelephorus) dzieduszyckii Lomnicki 1894 [synonym]
Helophorus (Meghelophorus) grandis Illiger 1798
Helophorus (Meghelophorus) khnzoriani Angus 1970
Invalid names: Helophorus (Praehelephorus) Lomnicki 1894 [synonym]
Subg. †Helophorus (Mesohelophorus) Ponomarenko 1977
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†Helophorus (Mesohelophorus) palaeosibiricus Fikáček et al. 2012
+
Invalid names: Mesohelophorus sibiricus Ponomarenko 1977 [replaced]
Subg. †Helophorus (Mesosperchus) Ponomarenko 1977
+
†Helophorus (Mesosperchus) costalis Ponomarenko 2014
†Helophorus (Mesosperchus) gracilis Prokin et al. 2010
†Helophorus (Mesosperchus) inceptivus Fikáček et al. 2012
†Helophorus (Mesosperchus) tarsalis Ponomarenko 1977
+
Invalid names: Mesohelophorus elongatus Ponomarenko 1990 [synonym]
†Helophorus (Mesosperchus) yixianus Fikáček et al. 2012
Subg. Helophorus (Orphelophorus) d'Orchymont 1927
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Helophorus (Orphelophorus) obscurellus Poppius 1907
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Invalid names: Helophorus (Orphelophorus) wandereri d'Orchymont 1927 [synonym]
Subg. Helophorus (Rhopalhelophorus) Kuwert 1886
Subg. Helophorus (Trichelophorus) Kuwert 1886
Helophorus auricollis Eschscholtz 1822
Helophorus brevipalpis Bedel 1881
Helophorus browni McCorkle 1973
Helophorus discrepans Rey 1885
Helophorus eclectus d'Orchymont 1945
Helophorus lacustris Le Conte 1850
Helophorus nubilus Fabricius 1777
Helophorus oblongus Le Conte 1850
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Invalid names: Helophorus (Rhopalhelophorus) kuwerti Lomnicki 1894 [synonym]
Helophorus orientalis Motschulsky 1860
Helophorus sempervarians Angus 1970
Helophorus splendidus Sahlberg 1880
Helophorus tuberculatus Gyllenhal 1808
Fm. Hydrochidae Thomson 1859
+
G. †Ponohydrochus Prokin and Strelnikova 2021
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†Ponohydrochus buryaticus Prokin and Strelnikova 2021
Fm. Hydrophilidae Latreille 1802 [water scavenger beetle]
+
Subfm. Acidocerinae Zaitzev 1908
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G. Helochares Mulsant 1846
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†Helochares fog Arriaga-Varela et al. 2021
†Helochares mydaleus Zhang 1989
†Helochares undecimstriatum Heer 1870
G. †Hydroyixia Fikáček et al. 2014
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†Hydroyixia elongata Fikáček et al. 2014
†Hydroyixia latissima Fikáček et al. 2014
Invalid names: Horelophopsinae Hansen 1997 [synonym]
G. †Aposphinctus Bode 1953
+
†Aposphinctus sibiricus Ponomarenko 1990
Invalid names: Hexameristus Bode 1953 [synonym], Spalacoides Bode 1953 [synonym], Streblocardioides Bode 1953 [synonym]
Subfm. Chaetarthriinae Bedel 1881
+
Tr. Anacaenini Hansen 1991
+
G. Anacaena Thomson 1859
G. Crenitis Bedel 1881
G. †Cretocrenis Fikáček et al. 2017
Invalid names: Horelophinae Hansen 1991 [synonym]
Subfm. Enochrinae Short and Fikáček 2013
+
G. Cymbiodyta Bedel 1881
+
†Cymbiodyta exstincta Scudder 1900
†Cymbiodyta samueli Fikáček et al. 2023
G. Enochrus Thomson 1859
+
Enochrus hamiltoni Horn 1890
†Enochrus innovatus Theobald 1937
†Enochrus lapsus Zhang 1989
†Enochrus pleistocenicus Lomnicki 1894
†Enochrus primaevus Scudder 1876
Enochrus quadripunctatus Herbst 1797
†Enochrus scudderi Wickham 1909
†Enochrus striatus Theobald 1937
Invalid names: Philydrus Solier 1834 [replaced]
Subfm. Georissinae Laporte 1819
+
G. †Georissites Ponomarenko 2008
+
†Georissites magnus Haupt 1956
†Georissites negev Ponomarenko 2008
G. Hydrocharis Berthold 1827
Subfm. Hydrochinae Thomson 1859
+
G. Hydrochus Germar 1824
+
†Hydrochus amictus Scudder 1890
Hydrochus elongatus Schaller 1783
†Hydrochus relictus Scudder 1890
Subfm. Hydrophilinae Latreille 1802
+
Tr. Hydrobiusini Mulsant 1844
+
G. †Baissalarva Fikáček et al. 2014
G. †Cretoxenus Fikáček et al. 2014
G. Hydrobius Linnaeus 1758
G. Limnoxenus Motshulsky 1853
Invalid names: Sperchopsini Hansen 1991 [synonym]
Tr. Hydrophilini Latreille 1802
+
Invalid names: Hydrophilina Latreille 1802 [invalid subgroup], Hydrophilopsis Heer 1862 [nomen dubium], Hydrophilopsis elongata Heer 1862 [nomen dubium], Hydrous ovalis Heer 1862 [nomen dubium]
Invalid names: Amphiopini Kuwert 1890 [empty], Cymbiodyta austerus Statz 1939 [nomen dubium]
G. †Paraspercheus Ponomarenko 1977
+
†Paraspercheus asiaticus Ponomarenko 1977
†Paraspercheus vitimensis Ponomarenko 1977
Subfm. Rygmodinae Orchymont 1916
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Invalid names: Andotypini Hansen 1991 [synonym], Borborophorini Hansen 1991 [synonym], Tormissini Hansen 1991 [synonym]
Subfm. Sphaeridiinae Latreille 1802
+
Tr. Megasternini Mulsant 1844
+
G. Cercyon Leach 1817
G. Cryptopleurum Mulsant 1844
G. Megasternum Mulsant 1844
Invalid names: Omicrini Smetana 1975 [empty], Protosternini Hansen 1991 [empty]
Invalid names: Escheria Heer 1847 [nomen dubium], Escheria bella Heer 1862 [nomen dubium], Escheria ovalis Heer 1847 [nomen dubium], Escheriidae Kolbe 1880 [synonym], Mesosperchus schultzi Ponomarenko 1985 [nomen dubium], Spercheinae Erichson 1837 [empty], Sphaeridium melanarium Gistl 1831 [nomen dubium]
G. †Laetopsia Fikáček et al. 2012
+
†Laetopsia baissensis Ponomarenko 1987
†Laetopsia bontsaganica Prokin 2009
†Laetopsia hydraenoides Prokin et al. 2010
†Laetopsia mongolica Ponomarenko 1987
†Laetopsia shatrovskiyi Prokin et al. 2010
Fm. Spercheidae Erichson 1837
+
G. Spercheus Kugelann 1798
+
†Spercheus punctatus Ponomarenko 2019
†Spercheus wightensis Ponomarenko 2019
Invalid names: Epimetopidae Zaitzev 1908 [empty], Georissidae Laporte 1819 [empty]
Superfm. Scarabaeoidea Latreille 1802 [scarab beetle]
+
Fm. †Alloioscarabaeidae Bai et al. 2012
+
Fm. Geotrupidae Latreille 1802 [earth-boring dung beetle]
+
G. Typhaeus Leach 1815
Invalid names: Lethrinae Mulsant and Rey 1871 [empty], Taurocerastinae [empty]
G. †Geotrupoides Handlirsch 1906
+
†Geotrupoides jiaoheense Hong 1992
†Geotrupoides kezuoensis Hong 1992
†Geotrupoides lithographicus Deichmüller 1886
†Geotrupoides saxosus Zhang 1997
†Geotrupoides songyingziensis Hong 1984
Fm. Glaphyridae MacLeay 1819 [bumble bee scarab beetle]
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Subfm. †Cretoglaphyrinae Nikolajev 2005
+
G. †Cretoglaphyrus Nikolajev 2005
+
†Cretoglaphyrus calvescens Nikolajev 2005
†Cretoglaphyrus leptopterus Nikolajev 2005
†Cretoglaphyrus olenguicus Nikolajev 2005
†Cretoglaphyrus rohdendorfi Nikolajev 2005
†Cretoglaphyrus transbaikalicus Nikolajev 2005
†Cretoglaphyrus zherikhini Nikolajev 2005
G. †Cretohypna Yan et al. 2012
+
†Cretohypna cristata Yan et al. 2012
†Cretohypna puncta Zhao et al. 2016
†Cretohypna robusta Zhao et al. 2016
†Cretohypna yixianensis Nikolajev and Ren 2015
Subfm. Glaphyrinae MacLeay 1819
+
G. Glaphyrus Latreille 1802
+
†Glaphyrus ancestralis Nikolajev and Ren 2011
†Glaphyrus antiquus Heer 1862
G. †Lithohypna Nikolajev et al. 2011
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†Lithohypna chifengensis Nikolajev et al. 2011
†Lithohypna laoningensis Nikolajev 2014
†Lithohypna lepticephala Nikolajev and Ren 2012
†Lithohypna longula Nikolajev and Ren 2012
†Lithohypna tuberculata Nikolajev and Ren 2012
†Lithohypna yuxiana Nikolajev and Ren 2012
G. †Mesohypna Nikolajev and Ren 2013
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†Mesohypna lopatini Nikolajev and Ren 2013
†Mesohypna probata Nikolajev 2014
G. †Paraglaphyrus Nikolajev 2015
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†Paraglaphyrus ovalis Nikolajev 2015
†Paraglaphyrus robustus Nikolajev 2015
†Paraglaphyrus subtilis Nikolajev 2015
†Paraglaphyrus yixianensis Nikolajev 2015
Fm. Glaresidae Prudhomme de Borre 1886 [enigmatic scarab beetle]
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G. †Cretoglaresis Nikolajev 2007
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†Cretoglaresis nana Nikolajev 2007
†Cretoglaresis ovalis Nikolajev 2009
G. Glaresis Erichson 1848
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†Glaresis burmitica Cai and Huang 2018
†Glaresis cretacea Nikolajev 2007
†Glaresis orthochilus Bai et al. 2010
†Glaresis tridentata Bai et al. 2014
Invalid names: Aphodiites Heer 1864 [nomen dubium]
G. †Holcorobeus Nikritin 1977
+
†Holcorobeus atrosulcatus Nikolajev 1992
†Holcorobeus evittatus Zhang 1992
†Holcorobeus nigrovittatus Nikolajev 1992
†Holcorobeus vittatus Nikritin 1977
Invalid names: Holcorobeus (Holcorobeus) Nikritin 1977 [invalid subgroup]
G. †Hongscarabaeus Krell 2000
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†Hongscarabaeus brunneus Hong 1982
Invalid names: Proscarabaeus Hong 1982 [replaced]
Fm. Hybosoridae Erichson 1847 [scavenger scarab beetle]
+
Subfm. Anaidinae Nikolajev 1996
+
Subfm. Ceratocanthinae Martinez 1968
+
Tr. Ceratocanthini Martinez 1968
+
G. Ceratocanthus White 1842
G. Germarostes Paulian 1982
G. Nesopalla Paulian and Howden 1982
Invalid names: Acanthocerini [replaced]
G. †Palaeopycnus Lu et al. 2022
+
†Palaeopycnus circus Lu et al. 2022
†Palaeopycnus fushengii Lu et al. 2022
Invalid names: Ivieolini Howden and Gill 2000 [empty]
Subfm. Hybosorinae Erichson 1847
+
G. †Cretohybosorus Nikolajev 1999
+
†Cretohybosorus buryaticus Nikolajev 1999
†Cretohybosorus striatulus Nikolajev 1999
G. †Jurahybosorus Nikolajev 2005
+
†Jurahybosorus kazakhstanicus Nikolajev 2008
†Jurahybosorus mongolicus Nikolajev 2005
G. †Leptosorus Nikolajev 2006
+
†Leptosorus fortus Ren et al. 1995
†Leptosorus zherikhini Nikolajev 2006
G. †Protohybosorus Nikolajev 2010
+
†Protohybosorus grandissimus Nikolajev 2010
†Protohybosorus karatavicus Nikolajev 2010
†Protohybosorus mesoasiaticus Nikolajev 2010
Invalid names: Anaidini Nikolajev 1996 [empty], Hybosorini Erichson 1847 [empty]
Subfm. Liparochrinae Ocampo 2006
+
Subfm. †Mimaphodiinae Nikolajev 2007
+
Fm. Lucanidae Latreille 1804 [stag beetle]
+
Subfm. Aesalinae MacLeay 1819
+
Tr. Aesalini MacLeay 1819
+
G. †Cretaesalus Nikolajev 1993
G. †Juraesalus Nikolajev et al. 2011
G. †Sinaesalus Nikolajev et al. 2011
G. †Cretolucanus Nikolajev 2007
+
†Cretolucanus longus Nikolajev 2007
†Cretolucanus ordinarius Nikolajev 2007
†Cretolucanus sibericus Nikolajev 2007
Subfm. Lampriminae MacLeay 1819
+
G. †Protognathinus Chalumeau and Brochier 2001
+
†Protognathinus spielbergi Chalumeau and Brochier 2001
Subfm. †Litholampriminae Nikolajev 2015
+
G. †Litholamprima Nikolajev and Ren 2015
+
†Litholamprima longimana Nikolajev and Ren 2015
†Litholamprima qizhihaoi Jiang et al. 2022
Subfm. Lucaninae Latreille 1804
+
G. †Anisoodontus Wu et al. 2022
+
†Anisoodontus qizhihaoi Wu et al. 2022
†Anisoodontus xiafangyuani Wu et al. 2022
Tr. Lucanini Latreille 1804
+
G. Dorcus MacLeay 1819
G. Lucanus Scopoli 1763
G. Penichrolucanus Deyrolle 1863
Invalid names: Figulidae Burmeister 1847 [synonym]
Invalid names: Chiasognathini Burmeister 1847 [empty], Lucanus fossilis Wickham 1913 [nomen dubium], Penichrolucaninae Arrow 1950 [synonym], Platycerus sepultus Germar 1837 [nomen dubium]
Subfm. †Paralucaninae Nikolajev 2000
+
Subfm. †Protolucaninae Nikolajev 2007
+
G. †Mesoscarabaeus Hong 1982
+
†Mesoscarabaeus corneus Hong 1982
†Mesoscarabaeus morulosus Hong 1982
Fm. Ochodaeidae Streubel 1846 [sand-loving scarab beetle]
+
Subfm. Chaetocanthinae Scholtz 1988
+
Subfm. †Cretochodaeinae Nikolajev 1995
+
G. †Cretochodaeus Nikolajev 1995
+
†Cretochodaeus mongolicus Nikolajev 1995
†Cretochodaeus striatus Nikolajev 1995
Subfm. Ochodaeinae Streubel 1846
+
Fm. Passalidae Leach 1815 [bess beetle]
+
Subfm. Aulacocyclinae Kaup 1868
+
Subfm. Passalinae Leach 1815
+
Fm. †Passalopalpidae Boucher and Bai 2016
+
G. †Mesoceratocanthus Nikolajev et al. 2010
+
†Mesoceratocanthus tuberculifrons Nikolajev et al. 2010
Fm. Pleocomidae Leconte 1861 [rain beetle]
+
Subfm. †Archescarabaeinae Nikolajev 2010
+
Subfm. †Cretocominae Nikolajev 2002
+
Subfm. Pleocominae Leconte 1861
+
G. †Proteroscarabaeus Grabau 1923
+
†Proteroscarabaeus magnus Nikolajev 1996
†Proteroscarabaeus nikritini Nikolajev 1996
†Proteroscarabaeus robustus Zhang 1997
†Proteroscarabaeus yeni Grabau 1923
Fm. Scarabaeidae Latreille 1802 [scarab beetle]
+
Subfm. Aegialiinae Laporte 1840
+
G. Aegialia Latreille 1807
+
Subg. Aegialia (Psammoporus) Thomson 1859
Aegialia cylindrica Eschscholtz 1822
†Aegialia rupta Scudder 1890
Aegialia sibirica Stebnicka 1977
G. †Cretaegialia Nikolayev 1993
+
†Cretaegialia aphodiiformis Nikolayev 1993
†Cretaegialia rhypariformis Nikolayev 1993
Subfm. Aphodiinae Leach 1815
+
Tr. Aphodiini Leach 1815
+
G. Agoliinus Schmidt 1913
Subtr. Aphodiina Leach 1815
G. Caligodorus Gordon and Skelley 2007
G. Dialytes Harold 1869
G. Dialytodius Gordon and Skelley 2007
G. Pardalosus Gordon and Skelley 2007
G. Planolinoides Dellacasa and Dellacasa 2005
Tr. Eupariini Schmidt 1910
+
G. Airapus Stebnicka and Howden 1996
G. Ataenius Harold 1867
G. Oxyomus Kirby and Spence 1828
G. Saprosites Redtenbacher 1858
Subfm. Aulonocneminae Janssens 1946
+
Subfm. Cetoniinae Leach 1815 [flower chafer]
+
Tr. Schizorhinini Burmeister 1842
+
G. †Genonota Zhang 1989
G. Heterosoma Schaum 1845
Subtr. Lomapterina Burmeister 1842
G. †Paleotrichius Poinar 2011
Subfm. †Cretoscarabaeinae Nikolajev 1995
+
G. †Cretoscarabaeus Nikolajev 1995
+
†Cretoscarabaeus gibbosus Nikolajev 1995
†Cretoscarabaeus lentiginosus Nikolajev 1995
Subfm. Dynastinae MacLeay 1819 [rhinoceras beetle]
+
Invalid names: Pentodon proserpinae Heer 1862 [nomen dubium]
Subfm. †Electrorubesopsinae Bai and Wang 2018
+
Subfm. Eremazinae Stebnicka 1977
+
Subfm. †Lithoscarabaeinae Nikolajev 1992
+
G. †Baisarabaeus Nikolajev 2005
+
†Baisarabaeus rugosus Nikolajev 2005
†Baisarabaeus yixianensis Nikolajev et al. 2013
Subfm. Melolonthinae Leach 1819
+
G. †Eophyllocerus Haupt 1950
+
†Eophyllocerus glaucinus Haupt 1950
†Eophyllocerus scrobiculatus Haupt 1950
Tr. Melolonthini Leach 1819
+
Subtr. Melolonthina Leach 1819
Subtr. Rhizotrogina Burmeister 1855
Invalid names: Schizonychina Burmeister 1855 [empty]
G. †Melolonthites Heer 1847
+
†Melolonthites aciculata Heer 1847
†Melolonthites collinsi Wickham 1929
†Melolonthites deperdita Heer 1847
†Melolonthites interemtus Cockerell 1926
†Melolonthites kollari Heer 1847
†Melolonthites laterosinuatus Piton 1935
†Melolonthites lavateri Heer 1847
†Melolonthites obsoleta Heer 1847
†Melolonthites parschlugiana Heer 1847
Tr. Sericini Kirby 1837
+
G. †Cretoserica Nikolajev 1998
G. †Lithanomala Nikolajev 1992
G. Maladera Mulsant 1871
G. †Mioserica Zhang 1989
Subtr. Sericina Kirby 1837
Subfm. †Prototroginae Nikolajev 2000
+
Subfm. Rutelinae MacLeay 1819
+
G. Mimela Kirby 1823
+
†Mimela rhenana von Heyden 1862
Mimela testaceipes Motschulsky 1860
Invalid names: Rhombonyx Hope 1837 [synonym]
G. †Zhangsunia Krell 2000
+
†Zhangsunia extumida Zhang et al. 1994
Invalid names: Dolichopoda Zhang et al. 1994 [replaced]
Subfm. Scarabaeinae Latreille 1802
+
G. Canthochilum Chapin 1934
+
†Canthochilum alleni Tarasov and Vaz-de-Mello 2016
†Canthochilum philipsivieorum Tarasov and Vaz-de-Mello 2016
Tr. Deltochilini Lacordaire 1856
+
G. Canthon Hoffmannsegg 1817
G. Deltochilum Eschscholtz 1822
G. Scybalophagus Martínez 1953
Tr. Oniticellini Kolbe 1905
+
Subtr. Drepanocerina van Lansberge 1875
Subtr. Oniticellina Kolbe 1905
Tr. Phanaeini Hope 1838
+
Invalid names: Onthophagus spitsbergeniensis Krell 2010 [nomen dubium]
G. Triodonta Mulsant 1842
Invalid names: Allidiostomatinae Arrow 1940 [empty], Coprophaginae [empty], Dynamopodinae Arrow 1911 [empty], Oniticellus amplicollis Heer 1862 [nomen dubium], Orphninae Erichson 1847 [empty], Pachypodinae Erichson 1840 [empty], Phaenomeridinae Erichson 1847 [empty]
Fm. †Septiventeridae Bai et al. 2013
+
Fm. Trogidae MacLeay 1819 [hide beetle]
+
Subfm. †Kresnikinae Tihelka et al. 2019
+
Subfm. Omorginae Nikolajev 2005
+
Subfm. Troginae MacLeay 1819
+
G. Trox Fabricius 1775
+
†Trox antiquus Wickham 1909
†Trox cretaceus Nikolajev 2007
†Trox minutus Nikolajev 2008
†Trox sibericus Nikolajev 2007
Invalid names: Trox oustaleti Scudder 1879 [nomen dubium]
Invalid names: Belohinidae Paulian 1959 [empty], Copris subterraneus Heer 1862 [nomen dubium], Diphyllostomatidae Holloway 1972 [empty], Gymnopleurus deperditus Heer 1862 [nomen dubium], Holcorobeini Nikolajev 1992 [invalid subgroup], Onthophagus urusheeri Krell 2000 [nomen dubium]
Superfm. Staphylinoidea Latreille 1802
+
Fm. Agyrtidae Thomson 1859 [primitive carrion beetle]
+
G. †Mesecanus Newton 1982
+
†Mesecanus communis Ponomarenko 1977
†Mesecanus lintouensis Lin 1985
Invalid names: Mesagyrtes Ponomarenko 1977 [replaced]
Subfm. Necrophilinae Newton 1997
+
G. †Palaeonecrophilus Strelnikova et al. 2020
+
†Palaeonecrophilus buryaticus Strelnikova et al. 2020
Subfm. Pterolomatinae Thomson 1862
+
Fm. Colonidae Horn 1880
+
Fm. Hydraenidae Mulsant 1844 [minute moss beetle]
+
Subfm. Hydraeninae Mulsant 1844
+
G. †Davidraenites Kirejtshuk 2019
+
†Davidraenites gratshevi Kirejtshuk 2019
†Davidraenites interruptus Kirejtshuk 2019
†Davidraenites spurcus Kirejtshuk 2019
G. †Ochtebiites Ponomarenko 1977
+
†Ochtebiites altus Ponomarenko 1977
†Ochtebiites incertus Ponomarenko 1977
†Ochtebiites manlaicus Ponomarenko 1980
†Ochtebiites minor Ponomarenko 1985
Invalid names: Prosthetopinae Perkins 1994 [empty]
Fm. Jacobsoniidae Heller 1926
+
Subfm. Derolathrinae Sen Gupta 1979
+
G. Derolathrus Sharp 1908
+
†Derolathrus abyssus Yamamoto and Parker 2017
†Derolathrus capdoliensis Tihelka et al. 2022
Derolathrus cavernicolus Peck 2010
†Derolathrus groehni Cai et al. 2016
Invalid names: Sarothriidae [replaced]
Fm. Leiodidae Fleming 1821 [round fungus beetle]
+
Subfm. Cholevinae Kirby 1837
+
Tr. Cholevini Kirby 1837
+
G. Catops Paykull 1798
Subtr. Cholevina Kirby 1837
G. Prionochaeta Horn 1880
Invalid names: Catopina Jeannel 1922 [empty]
Tr. Ptomaphagini Jeannel 1911
+
G. †Cretoptomaphagus Bao and Antunes-Carvalho 2020
Subtr. Ptomaphagina Jeannel 1911
Subtr. Ptomaphaginina Szymczakowski 1964
Subfm. Leiodinae Fleming 1821
+
Invalid names: Leptininae LeConte 1866 [empty], Platypsyllinae Ritsema 1869 [empty]
Fm. Ptiliidae Erichson 1845 [feather-winged beetle]
+
Subfm. Nossidiinae Sörensson and Delgado 2019
+
Fm. Staphylinidae Latreille 1802 [rove beetle]
+
Subfm. Aleocharinae Fleming 1821
+
Tr. Gymnusini Heer 1839
+
G. Adinopsis Cameron 1919
G. †Cretodeinopsis Cai and Huang 2015
G. †Electrogymnusa Wolf-Schwenninger 2004
G. Gymnusa Gravenhorst 1806
Invalid names: Deinopsini Sharp 1883 [synonym]
Tr. Homalotini Heer 1839
+
Subtr. Bolitocharina Thomson 1859
Subtr. Gyrophaenina Kraatz 1856
Subtr. Homalotina Heer 1839
G. Leptusa Kraatz 1856
Tr. Mesoporini Cameron 1959
+
G. Dictyon Fauvel 1900
G. †Mesosymbion Yamamoto et al. 2016
G. †Palaeomesoporus Yamamoto and Maruyama 2017
G. †Aleocharopsis Wickham 1913
+
†Aleocharopsis caseyi Wickham 1913
†Aleocharopsis secunda Wickham 1913
Subfm. Apateticinae Fauvel 1895
+
G. †Apticax Schomann and Solodovnikov 2012
+
†Apticax solidus Schomann and Solodovnikov 2012
†Apticax volans Schomann and Solodovnikov 2012
G. Boletobius Leach 1819
G. †Cardiolita Strand 1933
Subfm. Coomaniinae Żyła and Solodovnikov 2020
+
G. Coomania Cameron 1939
+
†Coomania enkarsios Jenkins Shaw et al. 2022
†Coomania megistos Jenkins Shaw et al. 2022
†Coomania yini Jenkins Shaw et al. 2022
Subfm. Dasycerinae Reitter 1887
+
G. †Protodasycerus Yamamoto 2016
+
†Protodasycerus aenigmaticus Yamamoto 2016
†Protodasycerus corpulentus Yin and Cai 2020
†Protodasycerus gigas Yin and Cai 2020
†Protodasycerus tuberculatus Yamamoto et al. 2020
G. Ellipsotomus Motschulsky 1845
Subfm. Glypholomatinae Jeannel 1962
+
G. †Juroglypholoma Cai et al. 2012
+
†Juroglypholoma antiquum Cai et al. 2012
†Juroglypholoma talbragarense Cai et al. 2013
G. Heterosoma Bernhauer 1903
G. †Laostaphylinus Zhang 1988
+
†Laostaphylinus fuscus Zhang 1988
†Laostaphylinus nigritellus Zhang 1988
Subfm. Megalopsidiinae Leng 1920
+
Subfm. Micropeplinae Leach 1815
+
G. Micropeplus Latreille 1809
+
Micropeplus caelatus Erichson 1839
Micropeplus cribratus LeConte 1863
Micropeplus fulvus Erichson 1840
†Micropeplus hoogendorni Matthews 1970
†Micropeplus hopkinsi Matthews 1970
Micropeplus laticollis Maklin 1853
†Micropeplus macrofulvus Gersdorf 1976
Micropeplus nelsoni Campbell 1989
†Micropeplus pengweii Jiang et al. 2020
Micropeplus porcatus Paykull 1789
Micropeplus punctatus LeConte 1863
Micropeplus staphylinoides Marsham 1802
Subfm. Mycetoporinae Thomson 1859
+
G. Bolitobius Leach 1819
G. Lordithon Thomson 1859
+
†Lordithon durabilis Scudder 1900
†Lordithon funditus Scudder 1900
Lordithon longiceps LeConte 1863
†Lordithon lyelli Scudder 1900
†Lordithon stygis Scudder 1900
G. Mycetoporus Mannerheim 1830
+
†Mycetoporus demersus Scudder 1900
Mycetoporus punctatissimus Hatch 1957
G. Parabolitobius Li et al. 2000
+
†Parabolitobius antiquus Yamamoto 2023
†Parabolitobius groehni Schülke 2000
G. †Ryvkinius Herman 2001
+
†Ryvkinius gracilis Ryvkin 1990
Invalid names: Mesoporus Ryvkin 1990 [replaced]
Invalid names: Bolitobiini Horn 1877 [synonym]
Subfm. Olisthaerinae Thomson 1858
+
Subfm. Omaliinae MacLeay 1825
+
Tr. Anthophagini Thomson 1859
+
G. Acidota Stephens 1829
G. Anthophagus Gravenhorst 1802
G. Arpedium Erichson 1839
G. Artochia Casey 1893
G. Eucnecosum Reitter 1909
G. Geodromicus Redtenbacher 1857
G. Lesteva Latreille 1796
G. Microedus Leconte 1874
G. Olophrum Erichson 1839
G. Orobanus LeConte 1878
G. Pelecomalium Casey 1886
G. Phlaeopterus Motschulsky 1853
G. †Sinanthobium Cai and Huang 2013
G. Unamis Casey 1893
Invalid names: Lestevini Mulsant and Rey 1880 [synonym]
G. †Archodromus Tikhomirova 1968
+
†Archodromus brachypterus Tikhomirova 1968
†Archodromus comptus Tikhomirova 1968
G. †Globoides Tikhomirova 1968
+
†Globoides microps Tikhomirova 1968
†Globoides oculatus Tikhomirova 1968
Tr. Omaliini MacLeay 1825
+
G. Acrulia Thomson 1858
G. †Duocalcar Peris and Thayer 2014
G. Hapalaraea Thomson 1858
G. Micralymma Westwood 1838
G. Omalium Gravenhorst 1802
G. Paraphloeostiba Steel 1960
G. Phyllodrepa Thomson 1859
G. Pycnoglypta Thomson 1858 [winter rove beetle]
Subfm. Osoriinae Erichson 1839
+
Tr. Osoriini Erichson 1839
+
G. Neosorius Fagel 1959
G. Nototorchus McColl 1985
G. Osoriellus Fagel 1959
G. Osorius Guerin-Meneville 1829
Tr. Thoracophorini Reitter 1909
+
Subtr. Clavilispinina Newton and Thayer 1992
Subtr. Lispinina Bernhauer and Schubert 1910
G. †Lispinomimus Irmler 2003
G. †Paleosorius Ortega-Blanco et al. 2013
Subtr. Thoracophorina Reitter 1909
Invalid names: Lispinini Bernhauer and Schubert 1910 [synonym]
Subfm. Oxyporinae Fleming 1821
+
G. Oxyporus Fabricius 1775
+
†Oxyporus blumenbachii Gravenhorst 1806
†Oxyporus cretaceous Yamamoto 2017
†Oxyporus impressus Piton 1940
†Oxyporus seuberti Heer 1862
†Oxyporus stiriacus Scudder 1890
†Oxyporus vulcanus von Heyden and von Heyden 1866
†Oxyporus yixianus Solodovnikov and Yue 2011
Subfm. Oxytelinae Fleming 1821
+
Tr. Coprophilini Heer 1839
+
G. †Gollandia Makranczy et al. 2018
G. †Mesocoprophilus Cai and Huang 2013
G. †Sinoxytelus Yue et al. 2010
Tr. Deleasterini Reitter 1909
+
G. Deleaster Erichson 1839
G. †Protodeleaster Cai et al. 2013
G. Syntomium Curtis 1828
G. †Megalymma Tikhomirova 1980
+
†Megalymma gigantea Tikhomirova 1980
†Megalymma rohdendorfi Tikhomirova 1980
G. †Mesoxytelus Tikhomirova 1968
+
†Mesoxytelus mandibularis Tikhomirova 1968
†Mesoxytelus parvus Tikhomirova 1968
Tr. Oxytelini Fleming 1821
+
G. Anotylus Thomson 1859
G. †Dolichoxenus Engel and Chatzimanolis 2009
G. Oxytelus Gravenhorst 1802
G. Platystethus Mannerheim 1830
Tr. Syntomiini Böving and Craighead 1931
+
G. †Pseudanotylus Cai and Huang 2013
G. †Vidya Chen et al. 2023
Invalid names: Euphaniini Reitter 1909 [empty], Piestini Erichson 1839 [empty]
Subfm. Paederinae Fleming 1821
+
Tr. Lathrobiini Laporte 1835
+
Subtr. Lathrobiina Laporte 1835
Subtr. Medonina Casey 1905
G. Micrillus Erichson 1839
Subtr. Scopaeina Mulsant and Rey 1878
G. Scymbalium Erichson 1839
G. †Lithoplanes Scudder 1886
+
†Lithoplanes deleta Oustalet 1874
†Lithoplanes elongata Oustalet 1874
Invalid names: Erinnys Oustalet 1874 [replaced]
G. †Mesostaphylinus Zhang 1988
+
†Mesostaphylinus antiquus Solodovnikov and Yue 2013
†Mesostaphylinus elongatus Solodovnikov and Yue 2013
†Mesostaphylinus laiyangensis Zhang 1988
†Mesostaphylinus yixianus Solodovnikov and Yue 2013
Tr. Paederini Fleming 1821
+
Subtr. Cryptobiina Casey 1905
Subtr. Paederina Fleming 1821
Subtr. Stilicina Casey 1905
Invalid names: Lithocharina Casey 1905 [empty], Stilicopsina Casey 1905 [empty]
Tr. Pinophilini Nordmann 1837
+
Subtr. Procirrina Bernhauer and Schubert 1912
Invalid names: Pinophilina Nordmann 1837 [empty]
Subfm. Phloeocharinae Erichson 1839
+
G. Charhyphus Sharp 1887
+
†Charhyphus balticus Shavrin 2020
†Charhyphus serratus Yamamoto and Shavrin 2021
Invalid names: Triga Fauvel 1878 [replaced], Trigites Handlirsch 1907 [synonym]
G. Phloeocharis Mannerheim 1830
+
†Phloeocharis agerata Chatzimanolis et al. 2013
†Phloeocharis burmana Yamamoto and Newton 2023
Subfm. Piestinae Erichson 1839
+
G. †Paleosiagonium Yue et al. 2016
+
†Paleosiagonium adaequatum Yue et al. 2016
†Paleosiagonium brevelytratum Yue et al. 2016
G. Siagonium Kirby and Spence 1815
+
†Siagonium crassum Giebel 1856
Invalid names: Prognathus Blondel 1827 [synonym]
Subfm. †Protactinae Heer 1847
+
Subfm. Proteininae Erichson 1839
+
Subfm. Protopselaphinae Newton and Thayer 1995
+
G. Protopselaphus Newton and Thayer 1995
+
†Protopselaphus newtoni Liu et al. 2020
†Protopselaphus thayerae Liu et al. 2020
Subfm. Pselaphinae Latreille 1802
+
Unr. Clavigeritae Leach 1815
+
Tr. Clavigerini Leach 1815
Tr. †Protoclavigerini Parker and Grimaldi 2014
Invalid names: Tiracerini Besuchet 1986 [empty]
G. †Duterotyrus Schaufuss 1890
Unr. Euplectitae Streubel 1839
+
Tr. Bythinoplectini Schaufuss 1890
Tr. Euplectini Streubel 1839
Tr. Metopiasini Raffray 1904
Tr. Trichonychini Reitter 1882
Tr. Trogastrini Brendel and Wickham 1890
Unr. Faronitae Reitter 1882
+
G. †Cretasonoma Peris et al. 2014
G. Faronus Aubé 1844
G. Sagola Sharp 1874
Invalid names: Faronini Reitter 1882 [invalid subgroup]
Unr. Goniaceritae Reitter 1882
+
Tr. Brachyglutini Raffray 1904
Tr. Bythinini Raffray 1890
Tr. Tychini Raffray 1904
Unr. Pselaphitae Latreille 1802
+
Tr. Ctenistini Blanchard 1845
Tr. Hybocephalini Raffray 1890
Tr. Pselaphini Latreille 1802
Tr. Tmesiphorini Jeannel 1949
Tr. Tyrini Reitter 1882
Invalid names: Arhytodini Raffray 1890 [empty]
Subfm. Pseudopsinae Ganglbauer 1895
+
Subfm. Scaphidiinae Latreille 1807
+
G. †Scaphidiopsis Handlirsch 1906
+
†Scaphidiopsis aequivoca Weyenbergh 1869
†Scaphidiopsis hageni Weyenbergh 1869
Subfm. Scydmaeninae Leach 1815 [antlike stone beetle]
+
Unr. †Hapsomelitae Poinar and Brown 2004
+
G. †Ektatotricha Chatzimanolis et al. 2010
G. †Electroatopos Chatzimanolis et al. 2010
G. †Hapsomela Poinar and Brown 2004
Unr. Mastigitae Fleming 1821
+
G. †Aenictosoma Schaufuss 1891
Tr. †Baltostigini Jałoszyński and Brunke 2018
Tr. Clidicini Casey 1897
Tr. Leptochromini Jałoszyński and Brunke 2018
Tr. Leptomastacini Casey 1897
Tr. Mastigini Fleming 1821
G. †Palaeomastigus Schaufuss 1890
Invalid names: Clidicus balticus Schaufuss 1896 [nomen dubium], Clidicus cretaceus Kirejtshuk et al. 2015 [nomen dubium], Papusini Jałoszyński and Brunke 2018 [empty]
Unr. Scydmaenitae Leach 1815
+
Tr. Chevrolatiini Reitter 1882
G. †Kachinus Chatzimanolis et al. 2010
Tr. Stenichnini Fauvel 1885
+
Invalid names: Cyrtoscydmini Schaufuss 1889 [synonym], Glandulariini Schaufuss 1889 [synonym]
Invalid names: Mastiginae Fleming 1821 [empty]
G. †Seniaulus von Heyden and von Heyden 1866
+
†Seniaulus scaphioides von Heyden and von Heyden 1866
Subfm. Silphinae Latreille 1807
+
G. Aclypea Reitter 1884
+
Aclypea altaica Gebler 1830
+
Invalid names: Blitophaga jacituca Rjabukhin 1990 [synonym], Blitophaga jacutica Rjabukhin 1990 [synonym]
Aclypea bituberosa Le Conte 1859
Aclypea calva Reitter 1885
Aclypea opaca Linnaeus 1758
Aclypea sericea Zubkov 1833
Invalid names: Blitophaga Reitter 1885 [synonym]
G. Dendroxena Motschulsky 1858
+
Dendroxena sexcarinata Motschulsky 1861
Invalid names: Xylodrepa Thomson 1859 [synonym]
G. Oiceoptoma Leach 1815
Tr. Silphini Latreille 1807
+
G. Necrophila Kirby and Spence 1828
G. Silpha Linnaeus 1758
G. Thanatophilus Leach 1815
Subfm. Solieriinae Newton and Thayer 1992
+
G. †Prosolierius Thayer et al. 2012
+
†Prosolierius crassicornis Thayer et al. 2012
†Prosolierius mixticornis Thayer et al. 2012
†Prosolierius parvus Peris et al. 2014
†Prosolierius tenuicornis Thayer et al. 2012
†Prosolierius thayerae Yamamoto 2023
Subfm. Staphylininae Latreille 1802
+
G. †Cretoprosopus Solodovnikov and Yue 2013
+
†Cretoprosopus problematicus Solodovnikov and Yue 2013
G. †Megolisthaerus Solodovnikov and Yue 2010
+
†Megolisthaerus chinensis Solodovnikov and Yue 2010
†Megolisthaerus minor Cai and Huang 2013
Tr. Staphylinini Latreille 1802
+
Subtr. Acylophorina Outerelo and Gamarra 1985
Subtr. Algonina Schillhammer and Brunke 2018
Subtr. Amblyopinina Seevers 1944
Subtr. Anisolinina Hayashi 1993
Subtr. †Baltognathina Brunke et al. 2019
G. †Cretoquedius Ryvkin 1988
Subtr. Cyrtoquediina Brunke and Solodovnikov 2016
G. †Durothorax Solodovnikov and Yue 2013
Subtr. Hyptiomina Casey 1906
Subtr. Philonthina Kirby 1837
Subtr. Quediina Kraatz 1857
Subtr. Staphylinina Latreille 1802
Invalid names: Quediini Horn 1878 [synonym], Xanthopygina Sharp 1884 [empty]
Tr. Xantholinini Erichson 1839
+
G. Gyrohypnus Leach 1819
G. Leptacinus Erichson 1839
G. Neohypnus Coiffait and Saiz 1964
G. Neoxantholinus Cameron 1945
G. Xantholinus Dejean 1821
Invalid names: Platyprosopinae Lynch Arribálzaga 1884 [synonym], Platyprosopini Lynch Arribálzaga 1884 [empty], Xantholininae Erichson 1839 [synonym]
Subfm. Steninae Macleay 1825
+
G. †Festenus Żyła et al. 2017
+
†Festenus gracilis Żyła et al. 2017
†Festenus robustus Żyła et al. 2017
G. Stenus Latreille 1796
+
Subg. Stenus (Hemistenus) Motschulsky 1860
+
Invalid names: Stenus (Parastenus) Heyden 1905 [synonym]
Subg. Stenus (Nestus) Rey 1883
Subg. Stenus (Tesnus) Rey 1884
†Stenus abraham Puthz 2010
†Stenus archetypus Puthz 2010
†Stenus atavus Puthz 2010
†Stenus avus Puthz 2010
Stenus biguttatus Linnaeus 1758
Stenus brunnipes Stephens 1833
Stenus comma LeConte 1863
†Stenus groehni Puthz 2010
†Stenus gypsi Oustalet 1874
Stenus juno Paykull 1789
†Stenus ketura Puthz 2010
†Stenus methusalem Puthz 2010
†Stenus morsei Scudder 1900
†Stenus noach Puthz 2010
†Stenus ornatus Förster 1891
Stenus pinguis Casey 1884
†Stenus prodromus Heer 1856
†Stenus scribai von Heyden and von Heyden 1866
Stenus scrupeus Casey 1884
Stenus sordidus Puthz 1988
Invalid names: Areus Casey 1884 [synonym]
Subfm. Tachyporinae MacLeay 1825
+
G. †Abscondus Tikhomirova 1968
+
†Abscondus regularis Tikhomirova 1968
†Abscondus similis Tikhomirova 1968
Tr. Tachinusini Fleming 1821
+
G. †Hesterniasca Zhang et al. 1992
G. †Mesotachinus Tikhomirova 1968
G. Nitidotachinus Campbell 1993
G. †Protachinus Cai et al. 2013
Invalid names: Acylophorus immotus Scudder 1900 [nomen dubium]
Subfm. Trichophyinae Thomson 1858
+
Subfm. Trigonurinae Reiche 1821
+
Invalid names: Empelinae Newton and Thayer 1992 [empty], Habrocerinae Mulsant and Rey 1877 [empty], Leptotyphlinae Fauvel 1874 [empty], Microsilphinae Crowson 1950 [empty], Neophoninae Fauvel 1905 [empty], Silphidae Latreille 1807 [synonym], Triga coeni Scudder 1900 [nomen dubium]
Invalid names: Dasyceridae Reitter 1887 [empty], Empelidae [empty]
Invalid names: Scarabaeiformia Crowson 1960 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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Source: c = class, p = phylum | |||||
References: Bush and Bambach 2015, Kiessling 2004 |
Age range
Maximum range based only on fossils: base of the Angulata to the top of the Holocene or 201.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 196.5 Ma
Minimum age of oldest fossil (stem group age): 196.5 Ma
Collections (611 total)
Oldest occurrences | |||
Time interval | Ma | Country or state | Original ID and collection number |
---|---|---|---|
Angulata | Switzerland | Aphodiites protogaeus (131766) | |
Pliensbachian | Russian Federation (Kemerovo) | Ochtebiites minor (125341) | |
Early/Lower Toarcian | Germany (Niedersachsen) | Aposphinctus conservatus, Streblocardioides striatus (107343) Opiselleipon gravis, Hexameristus inflatus (105002) Spalacoides simplex (107555) | |
Toarcian - Aalenian | Russian Federation (Kemerovo) | Hydrophilidae indet. (117989) | |
Bajocian - Bathonian | Russian Federation (Krasnoyarsk) | Morsum abdominale (118010) | |
Bajocian - Bathonian | China (Anhui) | Mesagyrtes lintouensis (106695) | |
Bajocian - Bathonian | Mongolia (Uver-Khangay Aymag) | Jurahybosorus mongolicus (118011) | |
Bathonian | Mongolia | Hydrophilidae indet. (119621) | |
Callovian - Oxfordian | China (Nei Mongol) | Alloioscarabaeus cheni (113366) Mesochodaeus daohugouensis (113365) Silphidae indet., Mesapatetica aenigmatica, Sinanthobium daohugouense, Juroglypholoma antiquum, Protolisthaerus jurassicus, Juraesalus atavus (113371) | |
Callovian - Oxfordian | China (Liaoning) | Protostaphylinus mirus (151526) | |
Callovian - Oxfordian | Kazakhstan (Chimkent) | Mesotachinus major, Archodromus brachypterus, Archodromus comptus, Globoides microps, Abolescus glabratus, Mesoxytelus mandibularis, Tunicopterus sigara (105970) Mesotachinus major, Archodromus comptus, Abolescus glabratus, Mesoxytelus mandibularis, Protohybosorus mesoasiaticus (106102) Mesotachinus major, Mesotachinus minor, Abscondus similis, Archodromus brachypterus, Porrhodromus communis, Abolescus glabratus, Mesoxytelus parvus, Jurahybosorus kazakhstanicus (105971) Ochtebiites incertus, Paraspercheus asiaticus (105426) Protohybosorus grandissimus (112846) Protohybosorus karatavicus (113439) Tachyporoides villosus, Mesotachinus medius, Mesotachinus major, Abscondus regularis, Archodromus brachypterus, Globoides oculatus, Sulcelytrinus antiquus, Abolescus glabratus, Paraspercheus asiaticus (106101) | |
Callovian - Oxfordian | China (Hebei) | Prostaphylinus mirus (154824) | |
Oxfordian | Russian Federation | Mesagyrtes communis (123219) | |
Oxfordian | Russian Federation (Buryat) | Mesagyrtes communis, Ochtebiites altus, Eophyllodrepa propinqua, Anicula inferna (105017) | |
Oxfordian - Kimmeridgian | China (Yunnan) | Hydrophilidae indet. (164859) | |
Tithonian | Australia (New South Wales) | Protachinus minor, Juroglypholoma talbragarense, Olisthaerinae indet., Protochares brevipalpis (128121) | |
Tithonian | Russian Federation | Mesosperchus tarsalis (125408 138150) | |
Tithonian | Mongolia | Hydraenidae indet., Paralucanus mesozoicus, Helophorus spp., Helophorus costalis (104748) Hydraenidae indet., Prospercheus cristatus, Helophorus spp., Helophorus inceptivus, Helophorus costalis (121394) Mesagyrtoides fulvus (152133) Prospercheus cristatus, Helophorus inceptivus (131803) | |
Tithonian | Russian Federation (Chita) | Aposphinctus sibiricus (152172) Aposphinctus sibiricus, Mesohelophorus elongatus (125407) Hydrophilopsia longitarsalis, Mesosperchus tarsalis, Mesohelophorus elongatus (114307) Mesosperchus tarsalis (119424 152174) Staphylinidae indet., Cuneocharis elongatus, Mesoporus gracilis, Prodaia inflata, Daiodromus umbrosus, Morda mora (119397) Undiatina pilosa, Trigunda lata, Mesosperchus tarsalis (123197) | |
Early/Lower Tithonian | Germany (Bayern) | Cassida aequivoca, Scaphidium hageni (123906) Geotrupes lithographicus (133578) Mesosperchus schultzi (123144) |