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Hybosoridae (scavenger scarab beetle)

Insecta - Coleoptera - Hybosoridae

Synonymy list
YearName and author
1847Hybosoridae Erichson
1981Hybosoridae Crowson p. 695
1995Hybosoridae Lawrence and Newton, Jr.
2000Hybosoridae Krell p. 878
2007Hybosoridae Nikolajev p. 78
2011Hybosoridae Bouchard et al.
2016Hybosoridae Bai and Zhang p. 235
2022Hybosoridae Cai et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
RankNameAuthor
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderPolyphagaEmery 1886
infraorderStaphyliniformia
superfamilyScarabaeoidea
familyHybosoridaeErichson 1847
familyHybosoridaeErichson 1847

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Hybosoridae Erichson 1847 [scavenger scarab beetle]
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Subfm. Anaidinae Nikolajev 1996
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G. Anaides Westwood 1842
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†Anaides howdeni Ocampo 2006
G. †Crassisorus Nikolajev et al. 2012
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†Crassisorus fractus Nikolajev et al. 2012
G. †Cretanaides Nikolajev 1996
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†Cretanaides trogopterus Nikolajev 1996
G. †Protanaides Nikolajev 2010
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†Protanaides sibiricus Nikolajev 2010
Subfm. Ceratocanthinae Martinez 1968
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Tr. Ceratocanthini Martinez 1968
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G. Ceratocanthus White 1842
+
†Ceratocanthus huarongii Lu et al. 2022
G. Germarostes Paulian 1982
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†Germarostes emarginatus Poinar 2014
G. Nesopalla Paulian and Howden 1982
+
†Nesopalla succini Lu et al. 2022
Invalid names: Acanthocerini [replaced]
G. †Palaeopycnus Lu et al. 2022
+
†Palaeopycnus circus Lu et al. 2022
†Palaeopycnus fushengii Lu et al. 2022
Invalid names: Ivieolini Howden and Gill 2000 [empty]
G. †Coprologus Heer 1847
+
†Coprologus gracilis Heer 1847
Subfm. Hybosorinae Erichson 1847
+
G. †Cretohybosorus Nikolajev 1999
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†Cretohybosorus buryaticus Nikolajev 1999
†Cretohybosorus striatulus Nikolajev 1999
G. †Fortishybosorus Yan et al. 2013
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†Fortishybosorus ericeusicus Yan et al. 2013
G. Hybosorus MacLeay 1819
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†Hybosorus lividus Heer 1862
†Hybosorus ocampoi Bai and Zhang 2016
G. †Jurahybosorus Nikolajev 2005
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†Jurahybosorus kazakhstanicus Nikolajev 2008
†Jurahybosorus mongolicus Nikolajev 2005
G. †Leptosorus Nikolajev 2006
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†Leptosorus fortus Ren et al. 1995
†Leptosorus zherikhini Nikolajev 2006
G. Phaeochrous Laporte 1840
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†Phaeochrous tertiarium Deichmüller 1881
G. †Protohybosorus Nikolajev 2010
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†Protohybosorus grandissimus Nikolajev 2010
†Protohybosorus karatavicus Nikolajev 2010
†Protohybosorus mesoasiaticus Nikolajev 2010
Invalid names: Anaidini Nikolajev 1996 [empty], Hybosorini Erichson 1847 [empty]
Subfm. Liparochrinae Ocampo 2006
+
G. †Libanochrus Kirejtshuk et al. 2011
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†Libanochrus calvus Kirejtshuk et al. 2011
Subfm. †Mimaphodiinae Nikolajev 2007
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G. †Mimaphodius Nikolajev 2007
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†Mimaphodius pusillus Nikolajev 2007
G. †Procoilodes Ocampo 2002
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†Procoilodes adrastus Ocampo 2002
G. †Pulcherhybosorus Yan et al. 2012
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†Pulcherhybosorus tridentatus Yan et al. 2012
G. †Sinochaetodus Lu et al. 2018
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†Sinochaetodus tridentatus Lu et al. 2018
G. †Sinohybosorus Nie et al. 2018
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†Sinohybosorus cheni Nie et al. 2018
G. †Tyrannasorus Ratcliffe and Ocampo 2001
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†Tyrannasorus rex Ratcliffe and Ocampo 2001
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Bajocian to the top of the Sarmatian or 170.30000 to 11.60800 Ma
Minimum age of oldest fossil (stem group age): 166.1 Ma

Collections (28 total)


Time interval Ma Country or state Original ID and collection number
Bajocian - Bathonian170.3 - 166.1Mongolia (Uver-Khangay Aymag) Jurahybosorus mongolicus (118011)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Jurahybosorus kazakhstanicus (105971) Protohybosorus grandissimus (112846) Protohybosorus karatavicus (113439) Protohybosorus mesoasiaticus (106102)
Early/Lower Barremian130.0 - 125.45Lebanon Libanochrus calvus (123416)
Late/Upper Barremian130.0 - 125.45China (Nei Mongol) Crassisorus fractus (125655) Pulcherhybosorus tridentatus (125656)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Fortishybosorus ericeusicus (124759) Sinohybosorus cheni, Sinochaetodus tridentatus (182836)
Aptian125.0 - 113.0Russian Federation (Buryatia) Cretanaides trogopterus (113354) Cretohybosorus buryaticus, Mimaphodius pusillus (122549) Cretohybosorus striatulus (106103) Leptosorus zherikhini (125791) Protanaides sibiricus (105969)
Aptian125.0 - 113.0China (Nei Monggol) Geotrupoides fortus (152245)
Early/Lower Cenomanian99.6 - 93.5Myanmar Hybosorus ocampoi (178121) Palaeopycnus circus (161726) Palaeopycnus fushengii (212006)
Late/Upper Eocene37.2 - 33.9Czech Republic Bolboceras tertiarium (138883)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Anaides howdeni (207031) Ceratocanthus huarongii (226931) Ceratocanthus sp., Ceratocanthus emarginatus, Germarostes sp. A (122472) Germarostes sp. A, Germarostes sp. B (186640) Nesopalla succini (226930) Procoilodes adrastus, Tyrannasorus rex (122982)
Sarmatian12.7 - 11.608Germany (Baden-Württemberg) Coprologus gracilis (115304) Hybosorus lividus (117252)