PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Byrrhoidea

Insecta - Coleoptera

Taxonomy

Byrrhoidea was named by Latreille (1804). It is extant. Its type is Byrrhus.

It was assigned to Coleoptera by Whalley (1985); and to Elateriformia by Crowson (1981), Lawrence and Newton (1995), Bouchard et al. (2011) and Cai et al. (2022).

Subtaxa

Byrrhidae, Chelonarium montanum, Serecoleus

Synonyms

Chelonarium montanum was named by Wickham (1914). Its type specimen is MCZ 2461, an exoskeleton, and it is a compression fossil. Its type locality is Florissant (MCZ Collection), which is in a Chadronian lacustrine - large shale in the Florissant Formation of Colorado.

It was considered a nomen dubium by Alekseev and Bukejs (2021).

Synonymy list

Year	Name and author
1804	Byrrhoidea Latreille p. 146
1914	Chelonarium montanum Wickham p. 434 figs. Pl 2, fig 5
1981	Byrrhoidea Crowson p. 695
1985	Byrrhoidea Whalley
1995	Byrrhoidea Lawrence and Newton, Jr.
2011	Byrrhoidea Bouchard et al. p. 48
2022	Byrrhoidea Cai et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924

Rank	Name	Author
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Neuropteriformia
—	Coleopterida
order	Coleoptera
—	Mesocoleoptera
—	Metacoleoptera
suborder	Polyphaga	Emery 1886
infraorder	Elateriformia	Crowson 1960
superfamily	Byrrhoidea
superfamily	Byrrhoidea

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. Byrrhoidea Latreille 1804

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Fm. Byrrhidae Latreille 1804 [pill beetle]

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G. †Amphicyrtella Ponomarenko 1990

†Amphicyrtella clavator Ponomarenko 1990

†Amphicyrtella granulum Ponomarenko 1990

Subfm. Amphicyrtinae LeConte 1861

G. Amphicyrta Erichson 1843

†Amphicyrta inhaesa Scudder 1900

Subfm. Byrrhinae Latreille 1804

Tr. Byrrhini Latreille 1804

G. Arctobyrrhus Munster 1902

Arctobyrrhus subcanus LeConte 1878

G. Byrrhus Linnaeus 1767

Byrrhus domesticus Geoffroy 1785

†Byrrhus exanimatus von Heyden and von Heyden 1866

Byrrhus eximius Le Conte 1850

†Byrrhus lucae Von Heyden 1859

†Byrrhus oeningensis Heer 1847

†Byrrhus ottawaensis Scudder 1895

Byrrhus pilula Linnaeus 1758

Byrrhus pustulatus Forster 1771

†Byrrhus romingeri Scudder 1900

G. Cytilus Erichson 1847

Cytilus alternatus Say 1825

Cytilus mimicus Casey 1912

Cytilus sericeus Forster 1771

†Cytilus tartarinus Scudder 1900

G. Porcinolus Mulsant and Rey 1869

Porcinolus murinus Fabricius 1794

Tr. Morychini El Moursy 1961

G. Morychus Erichson 1847

Subg. Morychus (Morychus) Erichson 1847

Morychus aeneus Fabricius 1775

Morychus oblongus Le Conte 1857

Morychus viridis Kuzmina and Korotyauv 1987

Tr. Pedilophorini Casey 1912

G. Pedilophorus Steffahny 1843

Tr. Simplocariini Mulsant and Rey 1869

G. †Byrrhites Kirejtshuk 2019

†Byrrhites bembridgensis Kirejtshuk 2019

G. Lioon Casey 1912

Lioon simplicipes Mannerheim 1852

G. Simplocaria Stephens 1829

Subg. Simplocaria (Simplocaria) Stephens 1829

Simplocaria arctica Poppius 1904

Simplocaria basalis Sahlberg 1903

Simplocaria elongata Sahlberg 1903

Simplocaria metallica Sturm 1807

Simplocaria tessellata Le Conte 1850

G. †Byrrhydium Heer 1864

†Byrrhydium arcuatum Heer 1864

†Byrrhydium morio Heer 1864

G. †Fangshanella Huang and Zhang 1997

†Fangshanella stolida Huang and Zhang 1997

G. Lamprobyrrhulus Ganglbauer 1902

Lamprobyrrhulus nitidus Schaller 1783

Subfm. †Lidryopinae Kirejtshuk and Nel 2016

G. †Lidryops Kirejtshuk and Nel 2016

†Lidryops occultus Kirejtshuk and Nel 2016

G. Lioligus Casey 1912

Lioligus nitidus Motschulsky 1845

G. †Mesobyrrhus Huang and Zhang 1997

†Mesobyrrhus parvus Huang and Zhang 1997

†Mesobyrrhus tanae Huang and Zhang 1997

G. †Mesosimplocaria Ponomarenko 1990

†Mesosimplocaria ovalis Ponomarenko 1990

†Mesosimplocaria parmula Ponomarenko 1990

G. †Nosotetocus Scudder 1892

†Nosotetocus debilis Scudder 1900

†Nosotetocus marcovi Scudder 1892

†Nosotetocus vespertinus Scudder 1900

G. Pelochares Mulsant and Rey 1869

Pelochares versicolor Waltl 1833

Subfm. Syncalyptinae Mulsant and Rey 1869

G. Curimopsis Ganglbauer 1902

Curimopsis albonotata Le Conte 1850

Curimopsis cyclolepidia Muenster 1902

Curimopsis moosilauke Johnson 1986

Tr. Syncalyptini Mulsant and Rey 1869

G. Syncalypta Dillwyn 1829

Invalid names: Nosodendron tritavum Scudder 1890 [nomen dubium]

G. †Serecoleus Yan et al. 2015

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†Serecoleus nadbitovae Yan et al. 2015

Invalid names: Chelonarium montanum Wickham 1914 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Angulata to the top of the Holocene or 201.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 196.5 Ma

Collections (127 total)

Time interval	Ma	Country or state	Original ID and collection number
Angulata	201.3 - 196.5	Switzerland	Byrrhydium arcuatum, Byrrhydium morio (131766)
Obtusum	196.5 - 189.6	United Kingdom (England)	Byrrhoidea indet. (114637)
Callovian - Oxfordian	166.1 - 157.3	China (Nei Mongol)	Serecoleus nadbitovae (113371)
Tithonian	152.1 - 145.0	Russian Federation (Chita)	Amphicyrtella granulum, Amphicyrtella clavator (125407)
Aptian	125.0 - 113.0	Russian Federation (Buryatia)	Byrrhidae indet. (113350)
Late/Upper Aptian	122.46 - 113.0	China (Beijing Shi)	Mesobyrrhus tanae, Mesobyrrhus parvus, Fangshanella stolida (140479)
Early/Lower Aptian	121.4 - 122.46	Russian Federation (Chita)	Mesosimplocaria ovalis, Mesosimplocaria parmula (119499)
Late/Upper Albian	105.3 - 99.6	France (Poitou-Charentes)	Lidryops occultus (99114)
Bridgerian	50.3 - 46.2	USA (Wyoming)	Nosodendron tritavum (122090)
Priabonian	38.0 - 33.9	United Kingdom (England)	Byrrhites bembridgensis (123959)
Chadronian	37.2 - 33.9	USA (Colorado)	Nosotetocus debilis, Byrrhus romingeri, Amphicyrta inhaesa (124624) Nosotetocus marcovi, Nosotetocus vespertinus, Nosotetocus debilis, Byrrhus romingeri, Chelonarium montanum, Amphicyrta inhaesa, Cytilus tartarinus (106384) Nosotetocus vespertinus (106381 113749)
Late/Upper Eocene	37.2 - 33.9	USA (Colorado)	Byrrhus romingeri (122093)
MP 30	28.4 - 23.03	Germany	Byrrhus exanimatus (123945)
MP 30	28.4 - 23.03	Germany (Nordrhein-Westfalen)	Byrrhus lucae (127628)
Sarmatian	12.7 - 11.608	Germany (Baden-Württemberg)	Byrrhus oeningensis (113689 117252)
Late/Upper Miocene	11.608 - 5.333	Russian Federation	Simplocaria basalis, Morychus aeneus (113713)
Messinian	7.246 - 5.333	USA (Alaska)	Byrrhus sp., Simplocaria sp. (123352)
Gelasian	2.588 - 1.806	Greenland	Arctobyrrhus subcanus (160108) Byrrhus sp., Simplocaria spp., Simplocaria basalis, Simplocaria metallica, Simplocaria elongata, Morychus aeneus, Arctobyrrhus subcanus (160109) Simplocaria elongata (160151) Simplocaria spp., Simplocaria basalis (160153) Simplocaria spp., Simplocaria basalis, Curimopsis moosilauke, Arctobyrrhus subcanus (160144)
Early/Lower Pleistocene	2.588 - 0.781	Russian Federation	Morychus viridis (163089) Simplocaria basalis, Curimopsis cyclolepidia, Morychus viridis (163069)
Chibanian	0.774 - 0.129	Russian Federation	Morychus viridis (163090 163091 163092 163093 163094 163095 163096 163097 163098 163099) Morychus viridis, Simplocaria arctica (163100)
Middle Pleistocene	0.774 - 0.129	United Kingdom	Byrrhus sp., Pelochares versicolor (168267) Limnichus versicolor (168265) Pelochares versicolor (168264 168266 168268 168269 168270 168271 168272)
Middle Pleistocene	0.774 - 0.129	Japan	Byrrhidae indet. (165708)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Washington)	Byrrhus sp., Cytilus sp., Cytilus alternatus, Morychus oblongus, Curimopsis albonotata (167662) Cytilus mimicus (169359 169360 169361) Cytilus sp. (157092) Lioligus nitidus (169380) Lioon simplicipes (169378) Morychus sp. (169370) Morychus sp., Lioligus nitidus, Lioon simplicipes (169379)
Late/Upper Pleistocene	0.129 - 0.0117	Russian Federation (Tyumenskaya)	Morychus ostasiaticus (164864 164865 164866 164867 164868 164869) Simplocaria semistriata (164863)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Colorado)	Arctobyrrhus subcanus (169336) Byrrhus eximius (169347) Simplocaria tessellata (169333 169353)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Wisconsin)	Byrrhus sp., Cytilus sp., Curimopsis sp., Simplocaria tesselata (169406)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom (England)	Byrrhus sp., Syncalypta sp. (164906) Simplocaria semistriata (164921)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Pennsylvania)	Curimopsis moosilauke (137207)
Late/Upper Pleistocene	0.129 - 0.0117	Canada (British Columbia)	Simplocaria sp., Cytilus alternatus, Morychus oblongus, Simplocaria tessellata (167673)
Late/Upper Pleistocene	0.129 - 0.0117	Canada (Ontario)	Byrrhus ottawaensis (152573)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom (Worcestershire)	Byrrhus sp., Syncalypta sp., Morychus aeneus, Simplocaria semistriata, Cytilus sericeus, Byrrhus pilula, Byrrhus pustulatus (163070) Morychus aeneus (163075)
Late/Upper Pleistocene	0.129 - 0.0117	Russian Federation	Curimopsis cyclolepidia, Morychus viridis (153446 163102) Curimopsis cyclolepidia, Morychus viridis, Simplocaria arctica (173208) Morychus viridis (153435 153436 153437 153438 153439 153440 153441 153442 153443 153444 153445 163101 163103 163104 163105 163106 163107 163108 163109 163110 173201 173204 173205 173209 173210 173211 173212 173213 173214 173215) Morychus viridis, Simplocaria arctica (173206) Simplocaria basalis, Morychus viridis (173216)
Tarantian	0.129 - 0.0117	USA (Pennsylvania)	Cytilus alternatus (137205 137206) Cytilus alternatus, Curimopsis moosilauke (137204) Simplocaria sp., Cytilus alternatus (137201)
Holocene	0.0117 - 0.0	Russian Federation (Novosibirsk)	Cytilus sericeus (164876) Lamprobyrrhulus nitidus (164873 164874 164875) Morychus ostasiaticus (164870) Morychus ostasiaticus, Lamprobyrrhulus nitidus (164871) Morychus ostasiaticus, Porcinolus murinus (164872) Porcinolus murinus (164877 164878 164879 164880)