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Dryopoidea

Insecta - Coleoptera

Synonyms
Synonymy list
YearName and author
1981Dryopoidea Crowson p. 695
2002Expansifemoria Hong p. 113
2002Expansifemoria orientalis Hong p. 114 figs. Pl 9, fig 1; text-fig 2.6.31-40
2002Brachypodites Hong p. 116
2002Brachypodites liaoningensis Hong p. 116 figs. 2.6.41-49
2002Actinocerites Hong p. 118
2002Actinocerites punctatus Hong p. 118 figs. 2.6.50-56
2021Dryopoidea Kundrata et al. p. 4
2022Dryopoidea Cai et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
RankNameAuthor
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderPolyphagaEmery 1886
infraorderElateriformiaCrowson 1960
superfamilyDryopoidea
superfamilyDryopoidea

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. Dryopoidea
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Fm. Chelonariidae Blanchard 1845
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G. Chelonarium Fabricius 1801
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†Chelonarium andabata Alekseev and Bukejs 2021
†Chelonarium dingansich Alekseev and Bukejs 2021
†Chelonarium dominicanum Poinar and Háva 2021
G. †Eochelonarium Kirejtshuk 2013
+
†Eochelonarium belle Kirejtshuk 2013
Fm. Dryopidae Billberg 1820 [long toed water beetle]
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G. Dryops Olivier 1791
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Dryops ernesti des Gozis 1886
Invalid names: Parnus Fabricius 1792 [synonym]
G. Helichus Erichson 1847
+
†Helichus eruptus Wickham 1911
†Helichus tenuior Wickham 1912
G. Helmis Latreille 1802
G. Pomatinus Sturm 1853
+
Invalid names: Parnidae Leach 1817 [replaced]
Fm. Elmidae Curtis 1830 [riffle beetle]
+
G. Cleptelmis Sanderson 1953
+
+
Invalid names: Cleptelmis ornata Schaeffer 1911 [synonym]
G. †Cretohypsilara Cai et al. 2018
+
†Cretohypsilara parva Cai et al. 2018
Subfm. Elminae Curtis 1830
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Tr. Elmini Curtis 1830
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Subtr. Elmina Curtis 1830
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G. Elmis Latreille 1802
G. Esolus Mulsant and Rey 1872
G. Limnius Illiger 1802
G. Riolus Mulsant and Rey 1872
+
Invalid names: Normandia Pic 1900 [synonym]
G. Heterlimnius Hinton 1935
+
+
Invalid names: Heterlimnius koebelei Martin 1927 [synonym]
†Heterlimnius samlandica Bollow 1940
Invalid names: Palaeoriohelmis Bollow 1940 [synonym]
G. Oulimnius Gozis 1886
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Oulimnius troglodytes Gyllenhal 1827
Subtr. Stenelmina Mulsant and Rey 1872
+
G. Stenelmis Dufour 1835
Tr. Macronychini Gistel 1848
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G. Macronychus Muller 1806
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G. Zaitzevia Champion 1923
G. Heterelmis Sharp 1882
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†Heterelmis groehni Bukejs et al. 2015
Invalid names: Larainae LeConte 1861 [empty]
Fm. Eulichadidae Crowson 1973 [forest stream beetle]
+
Subfm. Eulichadinae Crowson 1973
+
G. †Cretasyne Yan et al. 2013
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†Cretasyne lata Yan et al. 2013
†Cretasyne longa Yan et al. 2013
G. †Mesaplus Hong 1983
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†Mesaplus beipiaoensis Hong 1983
G. †Mesodascilla Martynov 1926
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†Mesodascilla jakobsoni Martynov 1926
Invalid names: Lasiosyne punctata Yan 2012 [nomen dubium], Lasiosyne shartegiensis Yan 2012 [nomen dubium]
Fm. Heteroceridae MacLeay 1825 [Variegated mud-loving beetle]
+
G. †Excavotarsus Li et al. 2020
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†Excavotarsus lini Li et al. 2020
†Excavotarsus minor Li et al. 2020
Subfm. Heterocerinae MacLeay 1825
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Tr. Heterocerini MacLeay 1825
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G. Heterocerus Fabricius 1792
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Heterocerus fusculus Kiesenwetter 1843
Heterocerus hispidulus Kiesenwetter 1843
G. †Heterocerites Ponomarenko 1986
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†Heterocerites kobdoensis Ponomarenko 1986
†Heterocerites magnus Prokin and Ren 2011
G. Lanternarius Pacheco 1964
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Lanternarius parrotus Pacheco 1964
G. Lapsus Pacheco 1964
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Lapsus tristis Mannerheim 1853
Invalid names: Elythomerinae Pacheco 1964 [empty]
G. †Jiuquanocoleus Hong 1982
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Fm. Limnichidae Erichson 1846 [minute marsh loving beetle]
+
Subfm. Cephalobyrrhinae Champion 1925
+
G. †Anomocephalobus Li et al. 2022
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†Anomocephalobus liuhaoi Li et al. 2022
G. Erichia Reitter 1895
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†Erichia cretacea Yu et al. 2018
Subfm. Limnichinae Erichson 1846
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Tr. Limnichini Erichson 1846
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G. †Burmochares Kirejtshuk and Prokin 2022
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†Burmochares groehni Kirejtshuk and Prokin 2022
G. †Hernandochares Kirejtshuk and Prokin 2022
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†Hernandochares electricus Hernando et al. 2018
G. Limnichus Dejean 1821
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Limnichus pygmaeus Sturm 1807
G. †Palaeoersachus Pütz et al. 2004
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†Palaeoersachus bicarinatus Pütz et al. 2004
G. Platypelochares Champion 1923
Invalid names: Hyphalinae Britton 1971 [empty], Thaumastodinae Champion 1924 [empty]
Fm. Lutrochidae Kasap and Crowson 1975 [robust marsh loving beetle]
+
G. †Lutrochites Wickham 1912
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†Lutrochites lecontei Wickham 1912
Fm. †Mastigocoleidae Tihelka et al. 2022
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G. †Cretaceocoleus Tihelka and Cai 2022
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†Cretaceocoleus saetosus Tihelka et al. 2022
G. †Mastigocoleus Tihelka and Cai 2022
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†Mastigocoleus resinicola Tihelka and Cai 2022
†Mastigocoleus rhinoceros Tihelka and Cai 2022
Fm. Psephenidae Lacordaire 1854 [water penny beetle]
+
Subfm. Eubrianacinae Jacobson 1913
+
G. Eubrianax Kiesenwetter 1874
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†Eubrianax vandeli Bertrand and Laurentiaux 1963
G. †Succinumanax Bao et al. 2018
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†Succinumanax birmaniasis Bao et al. 2018
G. †Vetujinbrianax Cai and Huang 2018
+
†Vetujinbrianax cretaceus Cai and Huang 2018
Subfm. Eubriinae Lacordaire 1857
+
G. Ectopria Leconte 1853
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†Ectopria laticollis Wickham 1913
Ectopria opaca Kiesenwetter 1874
Invalid names: Grammeubria Kiesenwetter 1874 [synonym]
G. †Protacnaeus Wickham 1914
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†Protacnaeus tenuicornis Wickham 1914
G. †Psephenella Martins-Neto 1998
+
†Psephenella ferreirai Martins-Neto 1998
Subfm. Psepheninae Lacordaire 1854
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G. Psephenus Haldeman 1853
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†Psephenus lutulentus Scudder 1900
Subfm. Psephenoidinae Hinton 1939
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G. Malacopsephenoides Jeng and Sato 2006
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G. Psephenoides Gahan 1914
G. †Pseudomataeopsephus Li and Cai 2021
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†Pseudomataeopsephus burmensis Li and Cai 2021
Fm. Ptilodactylidae Laporte 1836
+
Subfm. Anchytarsinae Champion 1897
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G. †Electrolichas Alekseev and Jäch 2016
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†Electrolichas circumbalticus Alekseev and Jäch 2016
G. †Aphebodactyla Chatzimanolis et al. 2012
+
†Aphebodactyla rhetine Chatzimanolis et al. 2012
Subfm. Cladotominae Pic 1914
+
G. Paralichas White 1859
+
†Paralichas striatopunctatus Kirejtshuk 2019
Subfm. Ptilodactylinae Laporte 1836
+
G. Ptilodactyla Illiger 1807
+
†Ptilodactyla eocenica Kundrata et al. 2021
†Ptilodactyla odnosum Telnov et al. 2022
G. †Ptilodactyloides Motschulsky 1856
+
†Ptilodactyloides stipulicornis Motschulsky 1856
Invalid names: Aploglossinae Champion 1897 [empty], Araeopidiinae Lawrence 1991 [empty]
Invalid names: Actinocerites Hong 2002 [nomen nudum], Brachypodites Hong 2002 [nomen nudum], Callirhipidae Emden 1924 [empty], Cneoglossidae Champion 1897 [empty], Expansifemoria Hong 2002 [nomen nudum]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Obtusum to the top of the Late/Upper Pleistocene or 196.50000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 189.6 Ma

Collections (99 total)


Time interval Ma Country or state Original ID and collection number
Obtusum196.5 - 189.6United Kingdom (England) Dryopoidea indet. (114637)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Mesodascilla jakobsoni (115792)
Callovian - Oxfordian166.1 - 157.3China (Liaoning) Mesaplus beipiaoensis (154730)
Tithonian152.1 - 145.0Mongolia Lasiosyne shartegiensis (104748 121393 125978)
Early/Lower Barremian130.0 - 125.45Lebanon Eochelonarium belle (154704)
Late/Upper Barremian130.0 - 125.45China (Nei Mongol) Cretasyne lata (106346) Cretasyne lata, Cretasyne longa (106347) Cretasyne longa (103654)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Heterocerites magnus (111716) Mastigocoleidae indet. (225721)
Aptian125.0 - 113.0China (Gansu) Jiuquanocoleus punctatus (155045)
Aptian125.0 - 113.0Mongolia (Kobdo aimak) Heterocerites kobdoensis (104054)
Aptian125.0 - 113.0Mongolia Lasiosyne punctata (103273)
Late/Upper Aptian122.46 - 113.0Brazil (Ceará) Mastigocoleus rhinoceros (122352 122557)
Early/Lower Cenomanian99.6 - 93.5Myanmar Aphebodactyla rhetine (123116) Burmochares groehni (172900) Cretaceocoleus saetosus (189783 223412) Cretohypsilara parva, Vetujinbrianax cretaceus, Succinumanax birmaniasis, Mastigocoleus resinicola, Cretaceocoleus saetosus, Anomocephalobus liuhaoi, Pseudomataeopsephus burmensis, Excavotarsus lini, Excavotarsus minor (161726) Erichia cretacea (165561) Succinumanax birmaniasis (178159 179937 200058)
Ypresian56.0 - 47.8India (Rajasthan) Elmidae indet. (198876)
Ypresian56.0 - 47.8China (Liaoning) Actinocerites punctatus (124530) Expansifemoria orientalis, Brachypodites liaoningensis (124820)
MP 1148.6 - 40.4Germany (Hessen) Eubrianacinae indet. (163150 164187) Eubrianacinae indet. 1 (120798)
Lutetian47.8 - 41.3France Eubrianax vandeli (193785)
Priabonian38.0 - 33.9United Kingdom (England) Paralichas striatopunctatus (21976) Ptilodactylidae indet. (123959) Ptilodactylidae indet., Paralichas striatopunctatus (123960)
Priabonian38.0 - 33.9Ukraine Ptilodactyla eocenica (154710) Ptilodactyla odnosum (207959)
Priabonian38.0 - 33.9Russian Federation Electrolichas circumbalticus (123419)
Priabonian38.0 - 33.9Germany Chelonariidae indet. (123769)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Chelonarium andabata (180657) Heterelmis groehni (110735) Palaeoersachus bicarinatus (124201) Palaeoriohelmis samlandica (123771) Ptilodactyla eocenica (124194)
Priabonian38.0 - 33.9Poland Platypelochares electricus, Chelonarium dingansich (110737) Ptilodactyloides stipulicornis (120811)
Chadronian37.2 - 33.9USA (Colorado) Dryops eruptus (106379) Dryops tenuior, Lutrochites lecontei (106381) Dryops tenuior, Psephenus lutulentus, Dryops eruptus, Protacnaeus tenuicornis (106384) Ectopria laticollis (113749)
Chattian28.1 - 23.03Germany (North Rhine-Westphalia) Helmis decoratus (122010)
Chattian28.1 - 23.03Czech Republic Dryopidae indet. (139944)
Chattian28.1 - 23.03Brazil Psephenella ferreirai (153054 156004)
Early/Lower Miocene23.03 - 15.97Mexico (Chiapas) Ptilodactylinae indet. (125757)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Chelonarium dominicanum (122472)
MN 515.97 - 13.65Germany (Baden-Württemberg) Dryopidae indet. (109058)
Messinian7.246 - 5.333Japan Eubrianax pellucidus (153919)
Gelasian2.588 - 1.806France Parnus sericeus (113996)
Chibanian0.774 - 0.129Japan Zaitzevia sp. (165706)
Middle Pleistocene0.774 - 0.129Japan Malacopsephenoides japonicus (194482) Psepheninae indet., Ectopria opaca (211426)
Middle Pleistocene0.774 - 0.129United Kingdom Dryops sp., Helichus substriatus, Esolus parallelepipedus, Oulimnius tuberculatus, Elmis aenea, Oulimnius troglodytes, Limnius volckmari, Normandia nitens, Macronychus quadrituberculatus, Limnichus pygmaeus (168265 168270) Dryops sp., Heterocerus sp., Helichus substriatus, Esolus parallelepipedus, Oulimnius tuberculatus, Elmis aenea, Oulimnius troglodytes, Limnius volckmari, Normandia nitens, Macronychus quadrituberculatus, Limnichus pygmaeus (168266) Dryops sp., Limnichus pygmaeus (168268) Helichus substriatus, Esolus parallelepipedus, Oulimnius tuberculatus, Oulimnius troglodytes, Limnius volckmari, Normandia nitens, Macronychus quadrituberculatus, Limnichus pygmaeus (168267) Helichus substriatus, Stenelmis canaliculatus, Esolus parallelepipedus, Oulimnius tuberculatus, Elmis aenea, Oulimnius troglodytes, Limnius volckmari, Normandia nitens, Macronychus quadrituberculatus (168269) Heterocerus sp., Riolus cupreus, Helichus substriatus, Stenelmis canaliculatus, Oulimnius tuberculatus, Oulimnius troglodytes, Limnius volckmari, Normandia nitens, Limnichus pygmaeus (168264) Limnius volckmari (168273) Riolus cupreus, Helichus substriatus, Stenelmis canaliculatus, Esolus parallelepipedus, Oulimnius tuberculatus, Elmis aenea, Oulimnius troglodytes, Limnius volckmari, Normandia nitens, Macronychus quadrituberculatus, Limnichus pygmaeus (168271 168272)
Middle Pleistocene0.774 - 0.129United Kingdom (Warwickshire) Dryops sp. (169388) Dryops sp., Limnius volkmari, Esolus parallelepipedus, Oulimnius troglodytes (169392) Dryops sp., Limnius volkmari, Esolus parallelepipedus, Oulimnius tuberculatus, Elmis aenea, Oulimnius troglodytes (169389) Dryops sp., Limnius volkmari, Esolus parallelepipedus, Oulimnius tuberculatus, Elmis aenea, Oulimnius troglodytes, Limnius opacus (169391) Dryops sp., Riolus sp., Esolus parallelepipedus, Oulimnius tuberculatus, Oulimnius troglodytes (169390) Oulimnius troglodytes (169393 169398 169399 169400) Oulimnius tuberculatus (169394 169397 169401) Oulimnius tuberculatus, Elmis aenea, Oulimnius troglodytes (169396) Oulimnius tuberculatus, Oulimnius troglodytes, Normandia nitens (169395)
Late/Upper Pleistocene0.129 - 0.0117USA (Washington) Cleptelmis ornata, Heterlimnius koebelei, Lapsus tristis (167662) Elmidae indet., Lanternarius parrotus (157092)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Worcestershire) Dryops ernesti, Heterocerus hispidulus, Heterocerus fusculus, Heterocerus fenestratus (163070)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation (Astrakhan) Dryops sp. (125794)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (England) Dryops sp. (164904 164906 164919) Dryops sp., Heterocerus sp., Riolus cupreus (164905) Limnius volkmari (164903)
Late/Upper Pleistocene0.129 - 0.0117Canada (British Columbia) Heterocerus sp. (167673)
Late/Upper Pleistocene0.129 - 0.0117Ukraine Parnus prolifericornis (153476)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation Heterocerus sp. (153436)