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Cleroidea
Taxonomy
Cleroidea was named by Latreille (1802). It is extant. Its type is Clerus.
It was assigned to Cucujiformia by Crowson (1981), Lawrence and Newton (1995), Bouchard et al. (2011), Robertson et al. (2015) and Cai et al. (2022).
It was assigned to Cucujiformia by Crowson (1981), Lawrence and Newton (1995), Bouchard et al. (2011), Robertson et al. (2015) and Cai et al. (2022).
Subtaxa
Acanthocnemidae, Biphyllidae, Byturidae, Chaetosomatidae, Cleridae, Foveapeltis, Juraniscus, Lophocateridae, Mathesius, Melyridae, Peltidae, Phloiophilidae, Phycosecidae, Prionoceridae, Protopeltidae, Rentoniidae, Rhadalidae, Shartegopsis, Thanerocleridae, Thymalidae, Trogossitidae (syn. Ostomatidae, Temnochilidae)
Synonymy list
Year | Name and author |
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1802 | Cleroidea Latreille p. 110 |
1981 | Cleroidea Crowson p. 696 |
1995 | Cleroidea Lawrence and Newton, Jr. |
2011 | Cleroidea Bouchard et al. p. 342 |
2015 | Cleroidea Robertson et al. |
2022 | Cleroidea Cai et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Superfm. Cleroidea Latreille 1802
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Fm. Biphyllidae Leconte 1861 [false skin beetle]
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G. †Paleobiphyllus Makarov and Perkovsky 2019
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†Paleobiphyllus ponomarenkoi Makarov and Perkovsky 2019
Fm. Byturidae Gistel 1848
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Invalid names: Byturinae Gistel 1848 [empty], Platydascillinae Pic 1914 [empty]
Fm. Cleridae Latreille 1802 [checkered beetle]
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Subfm. Clerinae Latreille 1802
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Tr. Clerini Latreille 1802
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Subtr. Clerina Latreille 1802
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G. Clerus Geoffroy 1762
G. Stigmatium Gray 1832
G. Thanasimus Latreille 1829
G. Enoclerus Gahan 1910
Tr. Hydnocerini Spinola 1844
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Subtr. Hydnocerina Spinola 1844
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G. Phyllobaenus Dejean 1833
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Invalid names: Hydnocera Newman 1838 [synonym], Miohydnocera Mawdsley 1992 [synonym]
Unr. Opilonitae Gistel 1848
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Invalid names: Clerus piacentinii Piton 1940 [nomen dubium], Clerus succini Giebel 1862 [nomen dubium], Hydnocerinae Spinola 1844 [synonym], Phyllobaeninae Lacordaire 1857 [synonym]
Subfm. Epiclininae Gunter et al. 2013
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Subfm. Korynetinae Laporte 1836
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G. †Aberrokorynetes Winkler 1990
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†Aberrokorynetes abludens Winkler 1990
†Aberrokorynetes oceanojubilaei Kolibáč et al. 2021
G. Necrobia Olivier 1795
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†Necrobia divinatoria Wickham 1914
Necrobia rufipes De Geer 1775 [red-legged ham beetle]
†Necrobia sibylla Wickham 1914
Necrobia violacea Linnaeus 1758
G. Prosymnus Laporte 1836
Invalid names: Enopliinae Gistel 1856 [synonym], Epiphloeinae Kuwert 1893 [synonym], Tarsosteninae Jacquelin du Val 1860 [synonym]
Subfm. Tillinae Fischer von Waldheim 1813
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G. Pseudopallenis Kuwert 1893
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†Pseudopallenis baltica Crowson 1964
†Pseudopallenis nejistota Kolibáč 1997
†Pseudopallenis palantir Kolibáč 1997
Invalid names: Prospinoza Crowson 1964 [synonym]
Invalid names: Clerus adonis Heer 1847 [nomen dubium], Tanocleria Hong 2002 [nomen nudum], Trichodes stebingeri Cockerell 1936 [nomen dubium]
Fm. Lophocateridae Crowson 1964
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G. Antillipeltis Lawrence et al. 2014
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†Antillipeltis alleni Lawrence et al. 2014
†Antillipeltis iviei Lawrence et al. 2014
G. †Burmacateres Kolibáč and Peris 2020
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†Burmacateres longicoxa Kolibáč and Peris 2020
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Invalid names: Mesolophocateres pengweii Yu et al. 2020 [synonym]
Invalid names: Mesolophocateres Yu et al. 2020 [synonym]
G. †Cervicatinius Tan and Ren 2007
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†Cervicatinius complanus Tan and Ren 2007
Invalid names: Sinopeltis Yu et al. 2012 [synonym]
G. Grynocharis Thomson 1859
G. Lophocateres Olliff 1883
Invalid names: Ancyronini Reitter 1876 [invalid subgroup], Lophocaterini Crowson 1964 [invalid subgroup]
Fm. Melyridae Leach 1815 [soft-winged flower beetle]
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Subfm. Dasytinae Laporte 1840
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G. Amecocerus Solier 1849
Tr. Danaceini Thomson 1859
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Tr. Amauronioidini Majer 1987
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G. †Baltamauroania Tshernyshev et al. 2023
G. †Protomauroania Tshernyshev 2021
Tr. Dasytini Laporte 1840
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Subfm. Malachiinae Fleming 1821
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Tr. Colotini Abeille de Perrin 1890
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G. Colotes Erichson 1840
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†Colotes constantini Kirejtshuk and Nel 2008
†Colotes impexus Kirejtshuk and Nel 2008
†Colotes sambicus Kubisz 2001
Tr. Malachiini Fleming 1821
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G. †Burmalachius Tshernyshev and Legalov 2023
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†Burmalachius acroantennatus Tshernyshev and Legalov 2023
G. Collops Erichson 1840
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†Collops desuetus Wickham 1914
†Collops extrusus Wickham 1914
†Collops priscus Wickham 1914
G. Malachius Fabricius 1775
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Invalid names: Malachiina Fleming 1821 [empty]
Tr. †Palpattalini Tshernyshev 2020
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G. †Palpattalus Tshernyshev 2016
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†Palpattalus baltiensis Tshernyshev 2016
†Palpattalus eocenicus Tshernyshev 2016
Invalid names: Attalini Abeille de Perrin 1891 [empty]
Subfm. Melyrinae Leach 1815
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Fm. Peltidae Kirby 1837 [shield beetle]
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G. Peltis Müller 1764
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†Peltis antehercynica Kolibáč et al. 2016
†Peltis constulata von Heyden 1862
†Peltis costulata von Heyden 1862
†Peltis laminata Wickham 1910
†Peltis minuscula Piton 1935
†Peltis tricostata Heer 1847
Invalid names: Peltinae Kirby 1837 [invalid subgroup], Peltini Latreille 1806 [invalid subgroup], Phenolia incapax Scudder 1876 [nomen dubium]
Fm. Prionoceridae Lacordaire 1857
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Fm. Rhadalidae LeConte 1861
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Subfm. Rhadalinae LeConte 1861
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Tr. Aplocnemini Mulsant and Rey 1868
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G. †Aploceble Majer 1998
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Subg. †Aploceble (Aploceble) Majer 1998
Subg. †Aploceble (Chalcaploceble) Tshernyshev 2012
G. Aplocnemus Stephens 1830
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Subg. Aplocnemus (Aplocnemus) Stephens 1830
Subg. Aplocnemus (Ischnopalpus) Schilsky 1894
Aplocnemus tarsalis Sahlberg 1822
G. Xamerpus Fairmaire 1886
Invalid names: Gietellinae Constantin and Menier 1987 [empty]
Fm. Thanerocleridae Chapin 1924
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Subfm. Zenodosinae Kolibáč 1992
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Invalid names: Thaneroclerinae Chapin 1924 [empty]
Fm. Thymalidae Leveille 1888
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Invalid names: Decamerinae Crowson 1964 [empty], Thymalinae Leveille 1888 [empty]
Fm. Trogossitidae Latreille 1802 [bark gnawing beetle]
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Subfm. Lophocaterinae Crowson 1964
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G. †Neolatitergum Zhang 2021
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†Neolatitergum glabrum Yu et al. 2014
Invalid names: Latitergum Yu et al. 2014 [replaced]
Subfm. Trogossitinae Latreille 1802
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Tr. †Lithostomatini Kolibáč and Huang 2008
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G. Temnoscheila Westwood 1830
G. Tenebroides Piller and Mitterpacher 1783
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Subg. Tenebroides (Polynesibroides) Kolibáč and Porch 2020
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†Tenebroides (Polynesibroides) atiu Kolibáč and Porch 2020
†Tenebroides (Polynesibroides) mihiura Kolibáč and Porch 2020
†Tenebroides (Polynesibroides) moorea Kolibáč and Porch 2020
†Tenebroides (Polynesibroides) raivavae Kolibáč and Porch 2020
†Tenebroides (Polynesibroides) rimatara Kolibáč and Porch 2020
Tenebroides (Polynesibroides) tahiti Kolibáč and Porch 2020
†Tenebroides (Polynesibroides) tubuai Kolibáč and Porch 2020
†Tenebroides corrugata Wickham 1913
†Tenebroides emortua Germar 1849
†Tenebroides eocenica Meunier 1921
†Tenebroides insignis Heer 1868
†Tenebroides tenebrioides Germar 1837
Invalid names: Trogossita Olivier 1790 [objective synonym]
Invalid names: Trogossitini Latreille 1802 [invalid subgroup]
Invalid names: Calitinae Winkler 1924 [empty], Ergoliinae Lacordaire 1854 [empty], Larinotinae Slipinski 1992 [empty], Ostomatidae Reitter 1882 [synonym], Temnochilidae [replaced]
Invalid names: Chaetosomatidae Crowson 1952 [empty], Phloiophilidae Kiesenwetter 1863 [empty], Phycosecidae Crowson 1952 [empty], Protopeltidae Crowson 1966 [empty], Rentoniidae Crowson 1966 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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Source: c = class, p = phylum | |||||
References: Kiessling 2004, Bush and Bambach 2015 |
Age range
Maximum range based only on fossils: base of the Early/Lower Toarcian to the top of the Holocene or 183.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 182.0 Ma
Minimum age of oldest fossil (stem group age): 182.0 Ma
Collections (117 total)
Time interval | Ma | Country or state | Original ID and collection number |
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Early/Lower Toarcian | Germany | Nitidulites argoviensis (123987) | |
Bajocian - Bathonian | Russian Federation (Krasnoyarsk) | Thoracotes sibiricus (118010) | |
Callovian - Oxfordian | China (Nei Mongol) | Cervicatinius complanus (113360) Idgiaites jurassicus (113366) Protoclerus korynetoides, Wangweiella calloviana, Sinomelyris praedecessor, Juraniscus majeri (113371) Sinopeltis jurassica, Sinopeltis amoena, Marginulatus venustus, Latitergum glabrum (117610) | |
Callovian - Oxfordian | Kazakhstan (Chimkent) | Juralithinus gracilidorsum (105970) Lithostoma expansum (115792) | |
Tithonian | Russian Federation (Chita) | Thoracotes glabrus (114307) | |
Tithonian | Mongolia | Shartegopsis miranda (104748) | |
Late/Upper Barremian | China (Liaoning) | Mathesius liaoningensis (111725) Paracretocateres bellus, Yixianteres beipiaoensis (124759) | |
Aptian | Russian Federation | Thoracotes glabrus (124811) | |
Aptian | Mongolia (Kobdo aimak) | Cretocateres mongolicus (104054) | |
Aptian | Russian Federation (Chita) | Thoracotes glabrus (138072) | |
Aptian | China (Shandong) | Sinosoronia longiantenna, Palaeoendomychus gymnus (106313) | |
Late/Upper Aptian | China (Beijing Shi) | Eotenebroides tumoculus (106965) | |
Late/Upper Aptian | Brazil (Ceará) | Peltinae indet. (122557) | |
Albian - Early/Lower Cenomanian | Russian Federation | Acanthocnemoides sukatshevae (35050) | |
Late/Upper Albian | Spain (Álava) | Trogossitidae indet. (108769) | |
Early/Lower Cenomanian | Myanmar | Archaeozenodosus bellus, Gracilenticrus burmiticus, Mesolophocateres pengweii, Parayixianteres parvus, Cretaidgia burmensis (165561) Burmalachius acroantennatus (182624) Cretorhadalus constantini (223712) Cretozenodosus fossilis, Burmacateres longicoxa, Microtrogossita qizhihaoi, Thanerosus antiquus, Foveapeltis rutai (161726) Foveapeltis rutai (235525) Mesolophocateres pengweii (201881) Zaiwa pankowskiorum (186684) | |
Early/Lower Cenomanian | France (Poitou-Charentes) | Cretamerus vulloi, Protodasytes cretaceus, Mesozenodosus insularis (161514) | |
Early/Lower Cenomanian | Myanmar (Kachin) | Anebomorpha cercorhampha (104053) | |
Late/Upper Santonian | Russian Federation | Paleobiphyllus ponomarenkoi (35056) | |
Paleocene - Eocene | China (Yunnan) | Platokoleos moxiensis (165275) | |
Selandian | France (Auvergne) | Protrogosita distincta (120351) | |
Selandian | Greenland | Tenebroides insignis (124000) Trogosita insignis (152291) | |
Late/Upper Paleocene | Argentina (Jujuy) | Trichodes stebingeri (151868) | |
Ypresian | France (Oise) | Colotes constantini, Colotes impexus (123850) | |
Ypresian | China (Liaoning) | Tanocleria tenuis (124820) | |
Ypresian | Canada (British Columbia) | Prionocerites tattriei (123795) | |
Bridgerian | USA (Wyoming) | Phenolia incapax (121306) | |
MP 11 | Germany (Hessen) | Trogossita eocaenica (120036) | |
MP 13 | Germany (Eifel) | Cleridae indet. (135640 135643) Cleridae indet., Cassidinae indet. 1 (138912) | |
Priabonian | Ukraine | Protocephaloncus perkovskyi (113621) Protomauroania mikhailovi (216854) | |
Priabonian | Poland | Aploceble berendti, Xamerpus sp., Aberrokorynetes abludens, Cymatodera helcaraxe, Smudlotillus bubik, Orthrius gandalf (109625) Colotes sambicus, Aberrokorynetes oceanojubilaei (110737) Promanodes serafini (123215) | |
Priabonian | Russian Federation | Palpattalus eocenicus, Seidlitzella hoffeinsorum, Aplocnemus hoffeinsorum, Aploceble lateantennata, Dasytes ochraceus, Premalachius obscurus, Protomauroania chaetophorata (123419) | |
Priabonian | Russian Federation (Kaliningrad) | Aploceble viridiaeneus, Palpattalus eocenicus, Palpattalus baltiensis, Aplocnemus baltiensis, Palpattalusinus transitivus, Aliattalus intercalaris (134456) Baltamauroania mirabilicornis (139928) Dasytidae indet., Aploceble kunowi, Aploceble berendti, Aploceble fuscipes, Xamerpus sp., Aplocnemus tarsalis, Cymatodera jetotak, Bilbotillus glum, Mitrandiria sauron, Phyllobaenus thingol (123962) Diplocoelus probiphyllus (138000) Eurymetopum wachteli (167590) Orthrius succini (123936) Promanodes alleni (153734) Prospinoza baltica (123766) Pseudopallenis baltica, Pseudopallenis nejistota, Thanasimodes jantar (147612) Strotocera marshalli (124205) Trogossitidae indet., Temnochilidae indet., Peltis sp., Lophocateres sp., Grynocharis sp., Calitys sp. (124073) Visokorynetes priscus, Pseudopallenis palantir (123221) Zahradnikius intercedens (124088) | |
Priabonian | Germany | Cleridae indet., Palpattalus baltiensis (123769) | |
Priabonian | United Kingdom (England) | Attalus flexus (123960) | |
Chadronian | USA (Colorado) | Collops priscus, Collops desuetus, Collops extrusus, Enoclerus florissantensis, Enoclerus pristinus, Hydnocera wolcotti, Necrobia divinatoria (106384) Eudasytites listriformis (106381) Eudasytites listriformis, Hydnocera wolcotti, Malachius pristinus, Necrobia sibylla (113749) Peltis laminata (124681) Tenebroides corrugata (124624) Trichochrous miocenus (106382) | |
Rupelian | Germany | Thanasimus ostenderus (122407) | |
Late/Upper Oligocene | France (Puy-de-Dome) | Troglops sp. (155204) | |
MP 30 | Germany | Peltis costulata (123945) Trogosita emortua (144974) Trogosita tenebrioides (115802) | |
Burdigalian | China (Shandong) | Cleridae indet. (128552) Pyrrhoclerus platyocephalus, Plastophagus pellous, Thanasimus taenianus (128551) Titanoclerus shanwangianus (138944) | |
Burdigalian - Langhian | Dominican Republic | Antillipeltis alleni (122982) Antillipeltis iviei (140453) Arawakis poinari (122472) | |
Sarmatian | Germany (Baden-Württemberg) | Gymnochila obesa, Trogosita sculpturata, Trogosita longicollis, Trogosita amissa, Trogosita bella, Trogosita assimilis, Trogosita koellikeri (117252) Peltis tricostata, Clerus adonis (115304) Trogossita koellikeri (115305) | |
Messinian | France | Peltis minuscula (138771) | |
Piacenzian | Germany (Niedersachsen) | Peltis antehercynica (182675) | |
Gelasian | France | Clerus piacentinii (113994) | |
Gelasian | Greenland | Amecocerus sp. (160109) | |
Early/Lower Pleistocene | Russian Federation | Cleridae indet., Troglocollops arcticus (163069) | |
Chibanian | Russian Federation | Troglocollops arcticus (163097 163098) | |
Middle Pleistocene | United Kingdom | Dasytes sp. (168264) | |
Rancholabrean | USA (California) | Necrobia violacea (196762) | |
Late/Upper Pleistocene | Russian Federation | Troglocollops arcticus (153435 153436 153437 153438 153439 153440 153442 153443 153445 153446 163103 173212 173213 173216) | |
Holocene | Argentina | Necrobia rufipes (162173) | |
Holocene | French Polynesia (Austral Islands) | Tenebroides mihiura, Tenebroides tubuai (219257) Tenebroides raivavae (219259) Tenebroides tubuai (219258) | |
Holocene | Cook Islands | Tenebroides atiu (143888) | |
Holocene | Tanzania | Clerus succini (129861) Tillus nigripes (152160) Tillus novemmaculatus, Stigmatium bifasciatum (165051) | |
Holocene | French Polynesia | Tenebroides moorea, Tenebroides tahiti (219255) Tenebroides rimatara (219256) |