PBDB Taxon

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Trechinae

Insecta - Coleoptera - Carabidae

Taxonomy

Trechii was named by Bonelli (1810). It is extant. Its type is Trechus.

It was corrected as Trechinae by Lawrence and Newton (1995), Tamutis et al. (2011), Bouchard et al. (2011), Ortuño and Arillo (2015) and Cai et al. (2022).

It was assigned to Carabidae by Lawrence and Newton (1995), Tamutis et al. (2011), Bouchard et al. (2011), Ortuño and Arillo (2015) and Cai et al. (2022).

Subtaxa

Bembidion succini, Patrobini, Trechinites, Trechitae

Synonyms

Patrobidae was named by Kirby (1837). It is extant. Its type is Patrobus.

It was synonymized subjectively with Trechinae by Maddison et al. (2019); it was reranked as the subfamily Patrobinae by Tamutis et al. (2011), Bouchard et al. (2011) and Cai et al. (2022).

It was assigned to Carabidae by Tamutis et al. (2011), Bouchard et al. (2011) and Cai et al. (2022).
Bembidium succini was named by Giebel (1856). It is an inclusion in amber. Its type locality is Baltic amber (Leipzig University collection), which is in a Priabonian terrestrial amber in the Russian Federation.

It was recombined as Bembidion succini by Ortuño and Arillo (2015); it was considered a nomen dubium by Schmidt and Michalik (2017).

Synonymy list

Year	Name and author
1856	Bembidium succini Giebel p. 64
1995	Trechinae Lawrence and Newton, Jr.
2011	Patrobinae Bouchard et al.
2011	Trechinae Bouchard et al. p. 113
2011	Patrobinae Tamutis et al.
2011	Trechinae Tamutis et al.
2015	Bembidion succini Ortuño and Arillo p. 578
2015	Trechinae Ortuño and Arillo p. 579
2022	Patrobinae Cai et al.
2022	Trechinae Cai et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924

Rank	Name	Author
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Neuropteriformia
—	Coleopterida
order	Coleoptera
—	Mesocoleoptera
—	Metacoleoptera
suborder	Adephaga
family	Carabidae	()
subfamily	Trechinae	()
subfamily	Trechinae	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Trechinae Bonelli 1810

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Tr. Patrobini Kirby 1837

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Subtr. Patrobina Kirby 1837

G. Patrobus Dejean 1821

Patrobus assimilis Chaudoir 1844

Patrobus fossifrons Eschscholtz 1823

Patrobus foveocollis Eschscholtz 1823

†Patrobus gasiorowskii Lomnicki 1894

†Patrobus henshawi Wickham 1917

†Patrobus kudiensis Cockerell 1925

Patrobus septentrionis Dejean 1828

Patrobus stygicus Chaudoir 1872

Invalid names: Patrobus decessus Scudder 1900 [synonym], Patrobus frigidus Scudder 1900 [synonym], Patrobus gelatus Scudder 1890 [synonym]

G. †Trechinites Heer 1862

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†Trechinites clairvillii Heer 1862

†Trechinites oblongus Heer 1862

Unr. Trechitae Bonelli 1810

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G. †Balticeler Schmidt and Maddison 2021

†Balticeler kerneggeri Schmidt and Maddison 2021

Tr. Bembidiini Stephens 1827

G. Asaphidion Des Gozis 1886

Asaphidion alaskanum Wickham 1919

Asaphidion yukonense Wickham 1919

G. Bembidion Latreille 1802

Subg. Bembidion (Asioperyphus) Vysoky 1986

Bembidion (Asioperyphus) bimaculatum Kirby 1837

Bembidion (Asioperyphus) lenae Csiki 1928

Bembidion (Asioperyphus) sordidum Kirby 1837

Subg. Bembidion (Bembidion) Latreille 1802

Bembidion (Bembidion) mutatum Gemminger and Harold 1868

Bembidion (Bembidion) quadrimaculatum Linnaeus 1761

Subg. Bembidion (Bracteon) Bedel 1879

Bembidion (Bracteon) alaskense Lindroth 1962

Bembidion (Bracteon) balli Lindroth 1962

Bembidion (Bracteon) inaequale Say 1823

Bembidion (Bracteon) lapponicum Zetterstedt 1828

Bembidion (Bracteon) levettei Casey 1918

Bembidion (Bracteon) velox Linnaeus 1761

Invalid names: Bembidion (Chrysobracteon) Netolitzky 1914 [synonym]

Subg. Bembidion (Diplocampa) Bedel 1896

Bembidion (Diplocampa) assimile Gyllenhal 1810

Subg. †Bembidion (Eodontium) Schmidt and Michalik 2017

†Bembidion (Eodontium) bukejsi Schmidt and Michalik 2017

Subg. Bembidion (Eupetedromus) Netolitzky 1911

†Bembidion (Eupetedromus) alekseevi Schmidt and Michalik 2017

Subg. Bembidion (Furcacampa) Netolitzky 1931

Bembidion (Furcacampa) versicolor LeConte 1847

Subg. Bembidion (Hirmoplataphus) Lindroth 1963

Bembidion (Hirmoplataphus) salebratum LeConte 1848

Subg. Bembidion (Hydrium) LeConte 1847

Invalid names: Bembidion (Eurytrachelus) Motschulsky 1850 [synonym]

Subg. Bembidion (Melomalus) Casey 1918

Bembidion (Melomalus) planatum LeConte 1848

Subg. Bembidion (Metallina) Motschulsky 1850

Bembidion (Metallina) dyschirinum LeConte 1861

Bembidion (Metallina) lampros Herbst 1784

Subg. Bembidion (Neobembidion) Bousquet 2006

Bembidion (Neobembidion) constricticolle Hayward 1897

Subg. Bembidion (Nesocidium) Sharp 1903

Bembidion (Nesocidium) ignicola Blackburn 1879

Bembidion (Nesocidium) pacificum Blackburn 1878

Subg. Bembidion (Notaphus) Dejean 1821

Bembidion (Notaphus) coloradense Hayward 1897

Bembidion (Notaphus) dorsale Say 1823

Bembidion (Notaphus) nigripes Kirby 1837

Bembidion (Notaphus) varium Olivier 1795

Subg. Bembidion (Ochthedromus) LeConte 1847

†Bembidion (Ochthedromus) davidae Pierce 1944

Subg. Bembidion (Peryphanes) Jeannel 1941

Bembidion (Peryphanes) grapii Gyllenhal 1827

Bembidion (Peryphanes) yukonum Fall 1926

Subg. Bembidion (Peryphus) Dejean 1821

Bembidion (Peryphus) tetracolum Say 1823

Bembidium (Peryphus) tibiale Duftschmid 1812

Subg. Bembidion (Philochthemphanes) Netolitzky 1943

†Bembidion (Philochthemphanes) christelae Ortuño and Arillo 2010

Invalid names: Bembidion (Archaeophilochthus) Ortuño and Arillo 2010 [synonym]

Subg. Bembidion (Philochthus) Stephens 1828

Bembidion (Philochthus) biguttatum Fabricius 1779

Bembidion (Philochthus) guttula Fabricius 1792

Subg. Bembidion (Phyla) Motschulsky 1844

Bembidion (Phyla) obtusum Audinet-Serville 1821

Subg. Bembidion (Plataphus) Motschulsky 1864

Bembidion (Plataphus) arcticum Lindroth 1963

Bembidion (Plataphus) breve Motschulsky 1845

Bembidion (Plataphus) fellmanni Mannerheim 1823

Bembidion (Plataphus) gebleri Gebler 1833

Bembidion (Plataphus) haruspex Casey 1918

Bembidion (Plataphus) planiusculum Mannerheim 1843

Bembidion (Plataphus) rusticum Casey 1918

Invalid names: Bembidion (Plataphodes) Ganglbauer 1891 [synonym]

Subg. Bembidion (Princidium) Motschulsky 1864

Bembidion (Princidium) punctulatum Drapiez 1821

Subg. Bembidion (Semicampa) Netolitzky 1910

Bembidion (Semicampa) morulum LeConte 1863

Subg. Bembidion (Terminophanes) Müller-Motzfeld 1998

Bembidion (Terminophanes) mckinleyi Fall 1926

Subg. Bembidion (Testedium) Motschulsky 1864

Bembidion (Testedium) bipunctatum Linnaeus 1761

Subg. Bembidion (Trepanedoris) Netolitzky 1918

Bembidion (Trepanedoris) doris Panzer 1797

Bembidion (Trepanedoris) fortestriatum Motschulsky 1845

Bembidion (Trepanedoris) frontale LeConte 1847

Bembidion (Trepanedoris) pseudocautum Lindroth 1963

Subg. Bembidion (Trepanes) Motschulsky 1864

Bembidion (Trepanes) articulatum Panzer 1796

Bembidion (Trepanes) octomaculatum Goeze 1777

Bembidion aeneum Germar 1824

†Bembidion damnosum Scudder 1900

†Bembidion everestae Pierce 1944

Bembidion femoratum Sturm 1825

†Bembidion festivum Zhang et al. 1994

Bembidion gilvipes Sturm 1825

Bembidion lunatum Duftschmid 1812

Bembidion minimum Fabricius 1792

Bembidion obliquum Sturm 1825

Bembidion schueppeli Dejean 1831

Bembidion semipunctatum Donovan 1806

†Bembidion strenum Zhang 1989

Bembidion virens Gyllenhal 1827

G. Ocys Stephens 1828

Ocys harpaloides Audinet-Serville 1821

G. Sinechostictus Motschulsky 1864

Subg. Sinechostictus (Sinechostictus) Motschulsky 1864

Sinechostictus (Sinechostictus) elongatus Dejean 1831

Invalid names: Bembidiina Stephens 1827 [invalid subgroup], Tachyina Motschulsky 1862 [empty]

Tr. Pogonini Laporte 1834

G. Pogonus Dejean 1821

Subg. Pogonus (Pogonoidius) Carret 1903

Pogonus (Pogonoidius) punctulatus Dejean 1828

Tr. Tachyini Motschulsky 1862

Subtr. Tachyina Motschulsky 1862

G. Elaphropus Motschulsky 1839

Subg. Elaphropus (Barytachys) Chaudoir 1868

Elaphropus (Barytachys) incurvus Say 1830

Subg. †Elaphropus (Tarsitachys) Erwin 1971

†Elaphropus (Tarsitachys) bilobus Erwin 1971

G. Polyderis Motschulsky 1862

†Polyderis antigua Erwin 1971

G. Tachys Dejean 1821

Tachys bistriatus Duftschmid 1812

†Tachys haywardi Wickham 1913

Tachys oahuensis Blackburn 1878

Tr. Trechini Bonelli 1810

G. †Antarctotrechus Ashworth and Erwin 2016

†Antarctotrechus balli Ashworth and Erwin 2016

G. Trechiama Jeannel 1927

Subtr. Trechina Bonelli 1810

G. Trechoblemus Ganglbauer 1891

G. Trechus Clairville 1806

Subg. Trechus (Trechus) Clairville 1806

Trechus (Trechus) apicalis Motschulsky 1845

Trechus (Trechus) chalybeus Dejean 1831

Trechus (Trechus) crassiscapus Lindroth 1955

Trechus (Trechus) oregonensis Hatch 1951

Trechus amplicollis Fairmaire 1859

†Trechus balticus Schmidt and Faille 2015

†Trechus capito Förster 1891

†Trechus eoanophthalmus Schmidt et al. 2016

†Trechus exhibitorius Schmidt et al. 2016

Trechus ovipennis Motschulsky 1845

†Trechus praeglacialis Lesne 1920

Trechus quadristriatus Schrank 1781

Trechus rivularis Gyllenhal 1810

Invalid names: Bembidion succini Giebel 1856 [nomen dubium], Patrobinae Kirby 1837 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Chadronian to the top of the Holocene or 37.20000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 33.9 Ma

Collections (127 total)

Time interval	Ma	Country or state	Original ID and collection number
Priabonian	38.0 - 33.9	Russian Federation	Elaphropus bilobus, Trechus balticus (123419)
Priabonian	38.0 - 33.9	Denmark	Tarsitachys bilobus (123762)
Priabonian	38.0 - 33.9	Russian Federation (Kaliningrad)	Balticeler kerneggeri (110735 138800 147612 220533) Bembidion alekseevi (139928) Bembidion bukejsi (152022) Bembidion christelae (123462) Bembidium succini (147344) Trechus exhibitorius (123420)
Priabonian	38.0 - 33.9	Germany	Trechus eoanophthalmus (123769)
Chadronian	37.2 - 33.9	USA (Colorado)	Tachys haywardi (113749)
Early/Lower Oligocene	33.9 - 28.4	France	Trechus capito (121473)
Rupelian	33.9 - 28.1	Germany	Trechus capito (122407)
Early/Lower Miocene	23.03 - 15.97	Mexico (Chiapas)	Polyderis antigua (140260)
Burdigalian	20.44 - 15.97	USA (California)	Bembidion davidae (161515) Bembidion everestae (161516)
Burdigalian	20.44 - 15.97	Russian Federation	Patrobus kudiensis (113705)
Burdigalian	20.44 - 15.97	China (Shandong)	Bembidion festivum (138946) Bembidion strenum (128552)
Middle Miocene	15.97 - 11.608	Antarctica	Antarctotrechus balli (137622)
Sarmatian	12.7 - 11.608	Germany (Baden-Württemberg)	Trechinites clairvillii, Trechinites oblongus (117252)
Late/Upper Miocene	11.608 - 5.333	Russian Federation	Bembidion sp. 1 (153434) Bembidion sp. 4, Bembidion sp. 5, Bembidion sp. 1, Bembidion sp. 2, Bembidion sp. 3, Bembidion sp. (113713)
Messinian	7.246 - 5.333	USA (Alaska)	Bembidion sp., Bembidion sp., Patrobus septentrionis, Bembidion spp. (123352)
Piacenzian	3.6 - 2.588	USA (Alaska)	Asaphidion yukonense (162793)
Gelasian	2.588 - 1.806	Greenland	Bembidion mckinleyi (160159) Bembidion spp., Bembidion dyschirinum, Bembidion sordidum (160108) Bembidion spp., Bembidion fellmanni (160152) Bembidion spp., Bembidion fellmanni, Bembidion mckinleyi, Bembidion sordidum (160151) Bembidion spp., Bembidion levettei, Bembidion velox, Bembidion lampros, Bembidion yukonum (160153) Bembidion spp., Bembidion sordidum (160147) Bembidion spp., Bembidion sp., Bembidion spp., Bembidion velox, Bembidion dyschirinum, Bembidion fellmanni, Bembidion arcticum, Bembidion planiusculum, Bembidion gebleri, Bembidion lenae, Bembidion yukonum, Bembidion sordidum (160146) Bembidion spp., Bembidion sp., Patrobus stygicus, Asaphidion alaskanum, Bembidion spp., Bembidion levettei, Bembidion lapponicum, Bembidion balli, Bembidion vitiosum, Bembidion dyschirinum, Bembidion fellmanni, Bembidion difficile, Bembidion arcticum, Bembidion planatum, Bembidion planiusculum, Bembidion gebleri, Bembidion salebratum, Bembidion mckinleyi, Bembidion grapii, Bembidion yukonum, Bembidion bimaculatum, Bembidion sordidum (160109) Bembidion spp., Patrobus stygicus, Bembidion spp., Bembidion levettei, Bembidion alaskense, Bembidion dyschirinum, Bembidion fellmanni, Bembidion planatum, Bembidion planiusculum, Bembidion gebleri, Bembidion mckinleyi, Bembidion yukonum, Bembidion bimaculatum, Bembidion sordidum (160144)
Early/Lower Pleistocene	2.588 - 0.781	Russian Federation	Bembidion spp. (163069 163089)
Calabrian - Ionian	1.8 - 0.129	United Kingdom (Durham)	Trechus praeglacialis (140489)
Chibanian	0.774 - 0.129	Russian Federation	Bembidion sp. (163091 163095)
Chibanian	0.774 - 0.129	Japan (Kumamoto)	Bembidion sp. (164901)
Chibanian	0.774 - 0.129	Japan	Bembidion sp. (165704 165705)
Middle Pleistocene	0.774 - 0.129	United Kingdom	Bembidion doris, Tachys bistriatus, Bembidion punctulatum, Bembidion tetracolum, Bembidion elongatum, Bembidion assimile (168264) Bembidion doris, Trechus quadristriatus, Bembidion articulatum, Bembidion octomaculatum (168266) Bembidion lampros, Bembidion doris, Tachys bistriatus, Bembidion varium, Bembidion semipunctatum, Bembidion tetracolum, Bembidion elongatum, Bembidion gilvipes, Bembidion biguttatum (168270) Bembidion lampros, Bembidion doris, Trechus quadristriatus, Bembidion elongatum, Bembidion assimile, Bembidion minimum, Bembidion articulatum, Bembidion obtusum (168265) Bembidion lampros, Bembidion doris, Trechus quadristriatus, Bembidion punctulatum, Bembidion tetracolum, Bembidion elongatum, Bembidion quadrimaculatum (168267) Bembidion lampros, Bembidion gilvipes, Bembidion assimile, Bembidion minimum, Bembidion articulatum, Bembidium harpaloides (168268) Bembidion lampros, Bembidion semipunctatum, Bembidion assimile, Bembidion articulatum (168269) Bembidion lampros, Bembidion varium, Bembidion elongatum, Bembidion assimile (168272) Bembidion lunatum, Trechus quadristriatus, Bembidion elongatum, Bembidion gilvipes, Bembidion assimile (168271)
Middle Pleistocene	0.774 - 0.129	United Kingdom (Warwickshire)	Bembidion assimile (169392) Bembidion assimile, Trechus rivularis (169391 169393) Bembidion sp., Bembidion assimile, Trechus rivularis, Bembidion schueppeli (169388) Trechus rivularis (169390)
Middle Pleistocene	0.774 - 0.129	Japan	Bembidion sp. (165707 165709)
Middle Pleistocene	0.774 - 0.129	Japan (Kumamoto)	Bembidiinae indet. (165716)
Late/Upper Pleistocene	0.129 - 0.0117	Canada (Ontario)	Bembidion balli (174230) Bembidion sp. 1, Bembidion sp. 2, Patrobus septentrionis, Bembidion balli (174229) Bembidium damnosum, Patrobus decessus (152577) Patrobus decessus, Patrobus frigidus (152575) Patrobus gelatus (122182)
Late/Upper Pleistocene	0.129 - 0.0117	Russian Federation (Astrakhan)	Bembidion sp., Patrobus sp., Pogonus punctulatus, Bembidion minimum (125794)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom (Worcestershire)	Bembidion bipunctatum (163075) Bembidion spp., Patrobus septentrionis, Bembidion lampros, Bembidion bipunctatum, Bembidion obliquum, Bembidion virens, Bembidion femoratum, Bembidion lunatum, Bembidion aeneum, Patrobus assimilis (163070)
Late/Upper Pleistocene	0.129 - 0.0117	Russian Federation	Bembidion sp. (153443 153444 153446 163101 163102 163104 163106 163110 173201 173202 173205 173206 173208 173216)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Wisconsin)	Bembidion sp., Bembidion mutatum, Asaphidion yukonense, Trechus apicalis, Bembidion grapei (169406)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Illinois)	Patrobus henshawi (164922)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Colorado)	Bembidion bimaculatum, Bembidion coloradense, Bembidion constricticolle (169336) Bembidion sp. (169333) Bembidion sp., Bembidion breve (169350) Bembidion sp., Bembidion breve, Elaphropus incurvus (169347)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom (England)	Bembidion doris (164906 164909) Bembidion doris, Bembidion guttula (164905) Bembidion guttula (164904) Patrobus assimilis (164908) Patrobus septentrionis (164917)
Late/Upper Pleistocene	0.129 - 0.0117	Japan	Bembidiinae indet. (166082)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Pennsylvania)	Bembidion fortestriatum (137212) Bembidion mutatum (137210) Patrobus septentrionis, Trechus crassiscapus, Bembidion fortestriatum (137209) Patrobus sp., Patrobus septentrionis, Trechus crassiscapus, Patrobus stygicus (137207) Trechus crassiscapus, Bembidion versicolor (137211) Trechus crassiscapus, Patrobus stygicus (137208)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Washington)	Bembidion haruspex, Bembidion planiusculum (169374) Bembidion rusticum (169359) Bembidion sp. (169370) Bembidion sp., Trechus chalybeus (169360) Bembidion sp., Trechus oregonensis (169376 169378) Bembidion spp., Patrobus stygicus, Bembidion bimaculatum, Trechus ovipennis, Trechus oregonensis, Bembidion inaequale, Bembidion nigripes, Patrobus fossifrons (167662) Bembidion spp., Trechus sp., Patrobus stygicus, Bembidion bimaculatum, Trechus ovipennis, Bembidion inaequale, Bembidion nigripes, Patrobus fossifrons (157092) Patrobus fossifrons (169366) Trechini indet. (169373) Trechus chalybeus (169361) Trechus oregonensis (169371 169372 169375 169377 169380) Trechus sp., Bembidion fortestriatum, Trechus oregonensis (169379) Trechus sp., Bembidion planiusculum (169365)
Late/Upper Pleistocene	0.129 - 0.0117	Ukraine	Patrobus gasiorowskii, Bembidium tibiale, Bembidium bipunctatum (153476)
Late/Upper Pleistocene	0.129 - 0.0117	Canada (British Columbia)	Bembidion spp., Trechus spp., Bembidion dorsale (167673)
Tarantian	0.129 - 0.0117	USA (Pennsylvania)	Bembidion anguliferum group, Patrobus sp., Patrobus septentrionis, Trechus crassiscapus, Patrobus stygicus (137201) Bembidion anguliferum group, Patrobus sp., Patrobus septentrionis, Trechus crassiscapus, Patrobus stygicus, Patrobus foveocollis (137204) Bembidion anguliferum group, Trechus crassiscapus, Patrobus foveocollis, Bembidion versicolor, Bembidion pseudocautum, Bembidion frontale (137206) Bembidion incertum group, Patrobus sp., Patrobus septentrionis, Trechus crassiscapus, Patrobus stygicus, Bembidion mutatum (137205)
Holocene	0.0117 - 0.0	USA (Hawaii)	Tachys oahuensis, Bembidion ignicola, Bembidion pacificum (224098)