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Harpalinae

Insecta - Coleoptera - Carabidae

Synonyms
Synonymy list
YearName and author
1810Licininae Bonelli
1861Harpalidae LeConte p. 16
1995Harpalinae Lawrence and Newton, Jr.
2011Harpalinae Bouchard et al. p. 121
2011Harpalinae Tamutis et al.
2015Licininae Liu et al. p. 428
2016Harpalinae Webster
2021Harpalinae Kirichenko-Babko et al. p. 437
2022Harpalinae Cai et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
RankNameAuthor
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderAdephaga
familyCarabidae()
subfamilyHarpalinae()
subfamilyHarpalinae()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Harpalinae Bonelli 1810
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Tr. Abacetini Chaudoir 1873
+
G. Loxandrus Leconte 1852
G. †Aethocarabus Ren 1995
+
Tr. Chlaeniini Brullé 1834
+
Subtr. Chlaeniina Brullé 1834
+
G. Chlaenius Bonelli 1810
+
Subg. Chlaenius (Agostenus) Fischer von Waldheim 1829
+
Subg. Chlaenius (Chlaeniellus) Reitter 1808
+
Chlaeniellus nigricornis Fabricius 1787
†Chlaenius electrinus Giebel 1862
†Chlaenius furvus Hong 1984
+
Invalid names: Shanwangicarabus paucilus Hong and Wang 1985 [synonym]
Chlaenius gebleri Ganglbauer 1891
†Chlaenius plicatipennis Wickham 1917
†Chlaenius primaevus Piton and Théobald 1939
†Chlaenius punctulatus Horn 1876
Invalid names: Shanwangicarabus Hong 1984 [synonym]
Tr. Galeritini Kirby 1825
+
Subtr. Galeritina Kirby 1825
+
G. Galerita Fabricius 1801
+
Galerita bicolor Drury 1770
†Galerita marshii Scudder 1900
Tr. Harpalini Bonelli 1810
+
Subtr. Anisodactylina Lacordaire 1854
+
G. Anisodactylus Dejean 1829
+
Invalid names: Eurytrichi Le Conte 1847 [synonym]
Subtr. Harpalina Bonelli 1810
+
G. Cratacanthus Dejean 1829
G. Euryderus Le Conte 1846
+
Invalid names: Nothopus Leconte 1852 [objective synonym]
G. Harpalus Latreille 1802
+
†Harpalus abolitus von Heyden and von Heyden 1866
†Harpalus bruckmanni Heer 1862
†Harpalus conditus Scudder 1900
†Harpalus constrictus Heer 1862
†Harpalus deletus Oustalet 1874
†Harpalus dilatatus Theobald 1937
†Harpalus excavatus Förster 1891
†Harpalus lheritieri Piton 1940
†Harpalus maceratus Wickham 1911
†Harpalus nero Oustalet 1874
†Harpalus nitens Heer 1870
†Harpalus nuperus Scudder 1900
†Harpalus obtusus Förster 1891
†Harpalus oustaleti Meunier 1915
†Harpalus pleistocenicus Lomnicki 1894
Harpalus pusillus Motschulsky 1850
†Harpalus redivivus Wickham 1917
†Harpalus sinis Heer 1847
†Harpalus stierlini Heer 1862
†Harpalus stygius Heer 1862
†Harpalus tabidus Heer 1847
†Harpalus tardigradus Heer 1862
†Harpalus ulomaeformis Wickham 1917
†Harpalus veterum Cockerell 1921
Harpalus vittatus Gelber 1833
†Harpalus whitfieldii Scudder 1900
G. Ophonus Dejean 1821
+
Subg. Ophonus (Metophonus) Bedel 1897
+
G. Selenophorus Dejean 1829
Invalid names: Dapti Le Conte 1847 [synonym]
Subtr. Stenolophina Kirby 1837
+
G. Bradycellus Erichson 1837
+
Bradycellus ruficollis Stephens 1828
G. Dicheirotrichus Jacquelin du Val 1857
+
G. Stenolophus Dejean 1821
+
†Stenolophus religatus Scudder 1900
Tr. Helluonini Hope 1838
+
Subtr. Omphrina Jedlicka 1941
+
G. Creagris Nietner 1857
G. Helluomorpha De Laporte 1834
+
†Helluomorpha protogaea Giebel 1862
Tr. Lebiini Bonelli 1810
+
G. †Agatoides Motschulsky 1856
+
†Agatoides carinulatus Motschulsky 1856
Subtr. Calleidina Chaudoir 1873
+
G. Plochionus Latreille and Dejean 1821
+
†Plochionus lesquereuxi Scudder 1900
Subtr. Cymindidina Laporte 1834
+
G. Cymindis Latreille 1806
+
Subg. Cymindis (Tarulus) Bedel 1906
+
Cymindis angularis Gyllenhal 1810
Cymindis arctica Kryzhanovskij and Emets 1979
†Cymindis aurora Horn 1876
†Cymindis extorpescens Scudder 1898
†Cymindis pulchella Heer 1847
G. †Cymindoides Motschulsky 1856
+
†Cymindoides sculptipennis Motschulsky 1856
Subtr. Dromiusina Bonelli 1810
+
G. †Antephilorhizus Kirichenko-Babko et al. 2021
+
†Antephilorhizus pripiatiensis Kirichenko-Babko et al. 2021
†Antephilorhizus zerovae Kirichenko-Babko and Perkovsky 2022
G. Dromius Bonelli 1810
+
Dromius angusticollis Sahlberg 1880
†Dromius bakeri Abdullah 1969
†Dromius resinatus Germar 1813
Invalid names: Lebina Germar 1813 [synonym]
G. Microlestes Schmidt-Göbel 1846
+
G. Paradromius Fowler 1887
+
Paradromius linearis Olivier 1795
G. Syntomus Hope 1838
+
Syntomus americanus Dejean 1831
Syntomus truncatellus Linnaeus 1761
Subtr. Lebiina Bonelli 1810
+
G. Lebia Latreille 1802 [colorful foliage ground beetle]
+
†Lebia amissa von Heyden and von Heyden 1865
†Lebia harrelli Cockerell 1936
†Lebia minuscula Piton 1936
†Lebia newberryi Scudder 1900
†Lebia protospiloptera Cockerell 1921
G. †Miolebidia Zhang 1989
+
†Miolebidia gaura Zhang 1989
Subtr. Pericalina Hope 1838
+
G. Coptodera Dejean 1825
+
†Coptodera elektra Gamboa and Ortuño 2015
G. †Protoscalidion Schaufuss 1889
+
†Protoscalidion rugiae Schaufuss 1889
Invalid names: Brachinus repressus Scudder 1900 [nomen dubium]
Tr. Licinini Bonelli 1810
+
Subtr. Dicaelina Laporte 1834
+
G. Dicaelus Bonelli 1813
+
†Dicaelus alutaceus Horn 1876
G. Diplocheila Brullé 1834
+
†Diplocheila henshawi Scudder 1890
Invalid names: Rembus Latreille 1817 [replaced]
Subtr. Licinina Bonelli 1810
+
G. Badister Clairville 1806
+
†Badister antecursor Scudder 1900
†Badister debilis Heer 1847
†Badister fragilis Heer 1862
†Badister grandis Heer 1862
†Badister prodromus Heer 1847
Tr. Odacanthini Laporte 1834
+
Invalid names: Casnoniae LeConte 1861 [synonym]
Tr. Panagaeini Bonelli 1810
+
Subtr. Panagaeina Bonelli 1810
+
G. Panagaeus Latreille 1802
+
†Panagaeus dryadum Oustalet 1874
Tr. Pseudomorphini Hope 1838
+
G. †Burmapseudomorphus Beutel et al. 2020
+
†Burmapseudomorphus planus Beutel et al. 2020
Tr. Pterostichini Bonelli 1810
+
G. Cyclotrachelus Chaudoir 1838
+
Subg. Cyclotrachelus (Evarthrus) Leconte 1853
†Cyclotrachelus tenebricus Scudder 1900
Subtr. Pterostichina Bonelli 1810
+
G. Abax Bonelli 1810
+
†Abax durhamianus Lesne 1926
Abax parallelus Duftschmid 1812
G. Myas Dejean 1828
+
†Myas rigefactus Scudder 1900
†Myas umbrarum Scudder 1900
G. Platysma Bonelli 1810
G. Plocamostethus Britton 1940
+
G. Poecilus Bonelli 1810
+
Poecilus kugelanni Panzer 1797
Poecilus nearcticus Lindroth 1966
†Poecilus virgatus Pongrácz 1935
Invalid names: Feronia Latreille 1816 [synonym]
G. Pterostichus Bonelli 1810
+
Subg. Pterostichus (Argutor) Dejean 1821
+
†Argutor antiquus Heer 1847
†Argutor pliocenicus Lesne 1920
Subg. Pterostichus (Eosteropus) Tschitschérine 1902
Subg. Pterostichus (Melanius) Bonelli 1810
+
Invalid names: Pterostichus (Omaseus) Dejean 1821 [synonym]
Subg. Pterostichus (Metallophilus) Chaudoir 1838
+
Invalid names: Pterostichus (Lyperopherus) Motschulsky 1844 [synonym]
Subg. Pterostichus (Phonias) des Gozis 1886
+
Subg. Pterostichus (Steropus) Dejean 1821
+
Invalid names: Pterostichus (Steroderus) Motschulsky 1850 [synonym]
Pterostichus abnormis Sahlberg 1880
†Pterostichus abrogatus Scudder 1890
Pterostichus cancellatus Motschulsky 1860
Pterostichus costatus Ménétriès 1851
†Pterostichus depletus Scudder 1900
†Pterostichus destitutus Scudder 1890
†Pterostichus destructus Scudder 1890
†Pterostichus distinctus Piton and Théobald 1939
†Pterostichus dormitans Scudder 1890
†Pterostichus fernquisti Wickham 1931
†Pterostichus fractus Scudder 1890
†Pterostichus gelidus Scudder 1877
†Pterostichus gurnetensis Cockerell 1921
†Pterostichus laevigatus Scudder 1890
Pterostichus minor Gyllenhal 1827
†Pterostichus molestus Theobald 1937
Pterostichus niger Schaller 1783
Pterostichus nigrita Paykull 1790
Pterostichus parens Tschitscherine 1897
†Pterostichus pumpellyi Scudder 1900
Pterostichus sublaevis Sahlberg 1880
Pterostichus tundrae Tschitscherine 1894
†Pterostichus vetustus Heer 1862
†Pterostichus walcotti Scudder 1900
G. Stomis Clairville 1806
+
†Stomis elegans Oustalet 1874
Invalid names: Feroniens Dejean 1825 [synonym]
G. †Sinis Heer 1862
+
†Sinis brevicollis Heer 1862
Tr. Sphodrini Laporte 1834
+
Subtr. Calathina Laporte 1834
+
G. Calathus Bonelli 1810
+
Calathus fuscipes Goeze 1777
Calathus piceus Marsham 1802
G. †Quasicalathus Schmidt and Will 2022
+
†Quasicalathus agonicollis Schmidt and Will 2022
†Quasicalathus conservans Schmidt and Will 2022
†Quasicalathus elpis Ortuño and Arillo 2009
Subtr. Sphodrina Laporte 1834
+
G. Pristonychus Dejean 1828
G. Sphodrus Clairville 1806
Subtr. Synuchina Lindroth 1956
+
G. Synuchus Gyllenhal 1810
Tr. Zabrini Bonelli 1810
+
Subtr. Amarina Zimmerman 1832
+
G. Amara Bonelli 1810
+
Subg. Amara (Amara) Bonelli 1810
+
†Amara (Amara) subaenea Lomnicki 1894
Subg. Amara (Bradytus) Stephens 1827
+
Subg. Amara (Celia) Zimmerman 1832
Subg. Amara (Curtonotus) Stephens 1827
+
Curtonotus alpinus Paykull 1790
Curtonotus fodinae Mannerheim 1825
Subg. Amara (Paracelia) Bedel 1899
+
Amara aenea De Geer 1774
Amara arcticola Poppius 1906
Amara aulica Panzer 1796
+
Invalid names: Carabus ruficornis De Geer 1774 [replaced]
Amara bifrons Gyllenhal 1810
Amara brunnea Gyllenhal 1810
†Amara cockerelli Wickham 1912
Amara conflata Le Conte 1855
†Amara danae Scudder 1900
Amara erratica Duftschmid 1812
†Amara foersteri Handlirsch 1907
+
Invalid names: Amara sinuata Förster 1891 [replaced]
Amara insignis Dejean 1831
Amara municipalis Duftschmid 1812
†Amara occlusa Scudder 1900
†Amara paleomelas Scudder 1879
†Amara pinguicula Heer 1862
†Amara powellii Scudder 1900
†Amara primigenia Heer 1862
†Amara princeps Heer 1862
†Amara procera Förster 1891
Amara pseudobrunnea Lindroth 1968
†Amara revocata Scudder 1900
†Amara sterilis Scudder 1900
†Amara veterata Scudder 1900
Subtr. Zabrina Bonelli 1810
+
G. Zabrus Clairville 1806
+
Tr. Zuphiini Bonelli 1810
+
Subtr. Zuphiina Bonelli 1810
+
G. Polistichus Bonelli 1810
+
†Polystichus hopei Oustalet 1874
Invalid names: Dryptini Bonelli 1810 [empty], Licininae Bonelli 1810 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Late/Upper Aptian to the top of the Holocene or 122.46000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 113.0 Ma

Collections (198 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Hauterivian - Aptian136.4 - 113.0Japan Harpalinae indet. (153093)
Late/Upper Aptian122.46 - 113.0China (Beijing Shi) Aethocarabus levigata (106965)
Early/Lower Cenomanian99.6 - 93.5Myanmar Burmapseudomorphus planus (165561) Oodes kachinensis (161726)
Turonian93.9 - 89.8Kazakhstan Harpalinae indet. (103274)
Danian - Selandian66.0 - 59.2Svalbard and Jan Mayen Carabites nitens (122011)
Selandian61.6 - 59.2France (Auvergne) Harpalus lheritieri (113685)
Late/Upper Paleocene58.7 - 55.8Argentina (Jujuy) Lebia harrelli (151868)
Ypresian56.0 - 47.8USA (Colorado) Lebia protospiloptera (105005)
Ypresian56.0 - 47.8Canada (British Columbia) Amara sp. (31677) Nebria paleomelas (122190)
Bridgerian50.3 - 46.2USA (Wyoming) Galerita marshii (122090)
Bridgerian50.3 - 46.2USA (Colorado) Harpalus veterum (114681)
Lutetian47.8 - 41.3Germany (Sachsen-Anhalt) Pristonychus sp., Poecilus virgatus (155693)
Priabonian38.0 - 33.9Poland Cymindoides sculptipennis, Agatoides carinulatus (120811) Quasicalathus elpis (110737)
Priabonian38.0 - 33.9Ukraine Antephilorhizus pripiatiensis (122997) Antephilorhizus zerovae (113621) Quasicalathus conservans (224320)
Priabonian38.0 - 33.9France Harpalus oustaleti (113709)
Priabonian38.0 - 33.9Russian Federation Quasicalathus elpis, Quasicalathus agonicollis (123419)
Priabonian38.0 - 33.9United Kingdom (England) Pterostichus gurnetensis (123960)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Calathus elpis (123462) Coptodera elektra (171610) Dromius bakeri (123766) Lebina resinata (147349) Quasicalathus elpis (110735 220533) Quasicalathus elpis, Quasicalathus agonicollis (123420)
Priabonian38.0 - 33.9Germany Protoscalidion rugiae (139931)
Chadronian37.2 - 33.9USA (Colorado) Amara cockerelli (114150) Amara danae, Pterostichus pumpellyi (106379) Amara powellii (113749) Amara powellii, Harpalus ulomaeformis, Harpalus redivivus, Pterostichus walcotti, Amara veterata (124624) Diplochila henshawi, Amara cockerelli, Amara danae, Amara revocata, Amara powellii, Pterostichus pumpellyi, Nebria occlusa, Myas rigefactus, Myas umbrarum, Pterostichus walcotti, Evarthrus tenebricus, Amara sterilis, Amara veterata, Plochionus lesquereuxi, Brachynus newberryi, Brachynus repressus, Nothopus kingii, Harpalus nuperus, Harpalus whitfieldii, Stenolophus religatus (106384) Harpalus maceratus (106381)
Late/Upper Eocene37.2 - 33.9USA (Colorado) Amara danae, Pterostichus walcotti, Amara sterilis, Harpalus whitfieldii (122093)
Early/Lower Oligocene33.9 - 28.4France Amara sinuata, Amara procera, Harpalus obtusus (121473) Harpalus excavatus (121474)
MP 2333.9 - 28.4France Harpalus nero (122535)
Rupelian33.9 - 28.1Germany Harpalus oustaleti, Pterostichus molestus, Harpalus offusus, Harpalus dilatatus (122407)
Rupelian33.9 - 28.1USA (Montana) Harpalini indet. (132747)
Late/Upper Oligocene28.4 - 23.03France (Puy-de-Dome) Harpalus nero (155205) Harpalus nero, Chlaenius primaevus, Pterostichus nov. sp. (155204)
MP 3028.4 - 23.03Germany Harpalus abolitus (123945)
Chattian28.1 - 23.03Czech Republic Harpalinae indet. (139944)
Chattian28.1 - 23.03Kazakhstan (Saisan) Calathus sp. (120122)
Chattian28.1 - 23.03France (Provence-Alpes-Côte d'Azur) Harpalus nero (122488) Harpalus oustaleti, Feronia minax, Feronia provincialis, Harpalus deletus, Stomis elegans (122487)
Chattian28.1 - 23.03France (Bouches-Du-Rhone) Panagaeus dryadum (123944) Polystichus hopei (122501)
Burdigalian20.44 - 15.97China (Shandong) Miolebidia gaura, Chlaenius furvus, Anisodactylus giganteus (128551) Shanwangicarabus furvus, Shanwangicarabus paucilus (128549) Shanwangicarabus furvus, Shanwangicarabus paucinus (128550)
Langhian15.97 - 13.82USA (Washington) Pterostichus fernquisti (140651)
Middle Miocene15.97 - 11.608Germany Lebia amissa (152936)
Middle Miocene15.97 - 11.608Japan Harpalinae indet. (153619)
Badenian13.65 - 12.7Austria Harpalinae indet. (131964)
Sarmatian12.7 - 11.608Germany (Baden-Württemberg) Argutor antiquus, Harpalus sinis (115305) Badister debilis (122210) Badister macrocephalus, Badister grandis, Badister fragilis, Pterostichus minutulus, Pterostichus vetustus, Amara princeps, Amara primigenia, Amara pinguicula, Harpalus stierlini, Harpalus tardigradus, Harpalus stygius, Harpalus constrictus, Dichirotrichus lividus, Sinis brevicollis (117252) Cymindis pulchella, Badister prodromus (115304) Harpalus bruckmanni, Dichirotrichus lividus (122868)
Sarmatian12.7 - 11.608Croatia Harpalus tabidus (117250)
Late/Upper Miocene11.608 - 5.333Russian Federation Amara sp., Pterostichus sp., Pterostichus sp., Pterostichus spp., Pterostichus sp., Amara quenseli, Dicheirotrichus cognatus, Pterostichus costatus (113713) Pterostichus sp., Pterostichus sp., Pterostichus sp. (153434)
Messinian7.246 - 5.333USA (Alaska) Amara sp., Pterostichus corvinus, Pterostichus vermiculosus (123352)
Early/Lower Pliocene5.333 - 3.6Russian Federation Pterostichus spp. (153127)
Zanclean5.333 - 3.6Japan Oodini indet. (165737)
Pliocene5.333 - 2.588Japan Oodini indet. (165743) Oodini indet., Pterostichinae indet. (165744)
Blancan4.9 - 1.8USA (Texas) Selenophorus sp. (154909)
Piacenzian3.6 - 2.588Germany (Niedersachsen) Lebia sp., Harpalinae indet. (109196)
Piacenzian3.6 - 2.588Japan Oodini indet. (165742) Pterostichus planicollis, Oodini indet., Pterostichinae indet. (165738)
Gelasian2.588 - 1.806Canada (Nunavut) Pterostichus ventricosus (162754)
Gelasian2.588 - 1.806France Lebia minuscula (113996)
Gelasian2.588 - 1.806Greenland Amara glacialis (160160) Amara spp., Pterostichus spp., Amara brunnea (160143) Amara spp., Pterostichus spp., Pterostichus brevicornis, Pterostichus planus (160151) Amara spp., Pterostichus spp., Pterostichus brevicornis, Pterostichus stygicus, Pterostichus nigrita, Pterostichus corvinus, Pterostichus caudicalis, Pterostichus arcticola, Pterostichus riparius, Pterostichus vermiculosus, Pterostichus agonus, Amara quenseli, Chlaenius interruptus, Amara pseudobrunnea, Amara brunnea, Pterostichus pinguedineus (160109) Amara spp., Pterostichus spp., Pterostichus brevicornis, Pterostichus stygicus, Pterostichus vermiculosus (160153) Amara spp., Pterostichus spp., Pterostichus spp., Pterostichus brevicornis (160146) Amara spp., Pterostichus spp., Pterostichus spp., Pterostichus brevicornis, Pterostichus agonus, Dromius angusticollis, Pterostichus pinguedineus (160144) Pterostichus spp., Pterostichus spp., Pterostichus vermiculosus (160147) Pterostichus spp., Pterostichus vermiculosus (160152)
Early/Lower Pleistocene2.588 - 0.781Japan Chlaenius sp. (165711) Panagaeinae indet. (153617)
Early/Lower Pleistocene2.588 - 0.781Russian Federation Pterostichus sp., Pterostichus spp., Amara sp., Dicheirotrichus cognatus, Poecilus nearcticus, Pterostichus sublaevis, Pterostichus tundrae, Curtonotus alpinus, Harpalus vittatus, Dicheirotrichus mannerheimi, Amara interstitialis, Pterostichus abnormis (163069) Pterostichus spp., Pterostichus spp., Poecilus nearcticus, Curtonotus alpinus, Harpalus amputatus, Dicheirotrichus mannerheimi, Pterostichus abnormis (163089)
Calabrian1.8 - 0.774Japan Chlaenius spp., Pterostichus planicollis, Chlaenius gebleri (160539)
Irvingtonian1.8 - 0.3USA (Pennsylvania) Pterostichus sp., Chlaenius punctulatus, Cymindis aurora, Dicaelus sp., Dicaelus alutaceus, Pterostichus sp. (20479)
Calabrian - Ionian1.8 - 0.129United Kingdom (Durham) Argutor pliocenicus, Pterostichini indet. (140489)
Chibanian0.774 - 0.129Japan Chlaenius sp., Pterostichus sp., Pterostichinae indet. (165704) Oodini indet., Pterostichinae indet., Synuchus sp. (165700) Pterostichinae indet. (165705)
Chibanian0.774 - 0.129Russian Federation Curtonotus alpinus, Amara glacialis, Harpalus vittatus (163098) Poecilus nearcticus, Amara glacialis, Harpalus amputatus (163096) Pterostichus sp., Curtonotus alpinus, Harpalus vittatus, Harpalus amputatus (163094) Pterostichus sp., Curtonotus alpinus, Pterostichus vermiculosus (163099) Pterostichus sp., Poecilus nearcticus, Curtonotus alpinus, Harpalus amputatus (163093) Pterostichus sp., Poecilus nearcticus, Curtonotus alpinus, Harpalus amputatus, Curtonotus fodinae (163097) Pterostichus sp., Poecilus nearcticus, Curtonotus alpinus, Harpalus vittatus, Cymindis arcticus (163090) Pterostichus sp., Poecilus nearcticus, Curtonotus alpinus, Harpalus vittatus, Harpalus amputatus, Curtonotus fodinae (163095) Pterostichus sp., Poecilus nearcticus, Curtonotus alpinus, Harpalus vittatus, Harpalus amputatus, Cymindis arcticus (163092) Pterostichus sp., Pterostichus costatus, Pterostichus sublaevis, Curtonotus alpinus, Harpalus vittatus, Pterostichus vermiculosus (163100) Pterostichus spp., Poecilus nearcticus, Curtonotus alpinus, Harpalus vittatus, Harpalus amputatus (163091)
Middle Pleistocene0.774 - 0.129Japan (Kumamoto) Chlaenius sp. (165715 165716)
Middle Pleistocene0.774 - 0.129Japan Pterostichinae indet. (165707 165708) Synuchus sp. (165709)
Middle Pleistocene0.774 - 0.129United Kingdom (Warwickshire) Dromius longiceps (169388) Pterostichus strenuus, Pterostichus vernalis (169391) Pterostichus vernalis (169394)
Middle Pleistocene0.774 - 0.129United Kingdom Calathus fuscipes (168271) Calathus piceus, Oodes helopioides (168269) Harpalus sp., Amara aenea, Pterostichus gracilis, Pterostichus minor, Pterostichus niger, Pterostichus melanarius, Chlaenius sulcicollis (168266) Pterostichus anthracinus, Pterostichus melanarius, Amara bifrons, Dromius linearis (168265) Pterostichus gracilis, Pterostichus minor, Dromius linearis, Microlestes minutulus (168272) Pterostichus strenuus, Amara aenea, Poecilus kugelanni, Pterostichus niger, Pterostichus melanarius, Abax parallelus, Calathus fuscipes, Zabrus tenebricosus (168267) Pterostichus vernalis, Pterostichus gracilis, Pterostichus minor (168270) Pterostichus vernalis, Trichocellus placidus, Bradycellus ruficollis, Pterostichus minor, Chlaenius sulcicollis, Syntomus truncatellus (168264)
Rancholabrean0.3 - 0.012USA (California) Pterostichus sp., Amara insignis (150336)
Late/Upper Pleistocene0.129 - 0.0117Japan Oodini indet. (164902) Pterostichus sp., Oodini indet., Synuchus sp. (166082)
Late/Upper Pleistocene0.129 - 0.0117USA (Wisconsin) Cryobius sp., Calathus ingratus (169406)
Late/Upper Pleistocene0.129 - 0.0117USA (Colorado) Pterostichus sp. (169333 169347) Pterostichus sp., Harpalus sp. (169353)
Late/Upper Pleistocene0.129 - 0.0117Ukraine Pterostichus anthracinus, Pterostichus blanduloides, Pterostichus aethiops, Amara subaenea, Amara boryslavica, Harpalus pleistocenicus (153476)
Late/Upper Pleistocene0.129 - 0.0117USA (Massachusetts) Cymindis extorpescens (152284)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom Harpalus sp. (152473)
Late/Upper Pleistocene0.129 - 0.0117USA (Virginia) Dicaelus sp., Galerita bicolor (103991)
Late/Upper Pleistocene0.129 - 0.0117USA (Washington) Pterostichus castaneus (169377 169378) Pterostichus pumilis (169375 169379 169380) Pterostichus sp., Pterostichus adstrictus, Syntomus americanus, Microlestes sp. (157092) Pterostichus sp., Pterostichus riparius, Chlaenius interruptus, Pterostichus adstrictus, Amara conflata, Calathus ruficollis (167662)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation Amara glacialis, Curtonotus fodinae (173203) Curtonotus alpinus (173202) Poecilus nearcticus, Curtonotus alpinus (173210 173212 173213) Poecilus nearcticus, Curtonotus alpinus, Harpalus vittatus, Harpalus amputatus, Cymindis arcticus (153440) Pterostichus sp. (163106 173204 173209) Pterostichus sp., Curtonotus alpinus, Amara glacialis, Harpalus vittatus, Harpalus amputatus, Curtonotus fodinae, Amara arcticola (173201) Pterostichus sp., Curtonotus alpinus, Dicheirotrichus mannerheimi (163108) Pterostichus sp., Curtonotus alpinus, Harpalus amputatus (173215) Pterostichus sp., Poecilus nearcticus (163105 163107) Pterostichus sp., Poecilus nearcticus, Curtonotus alpinus (163103 163109 173214) Pterostichus sp., Poecilus nearcticus, Curtonotus alpinus, Dicheirotrichus mannerheimi (163101) Pterostichus sp., Poecilus nearcticus, Curtonotus alpinus, Harpalus pusillus, Harpalus amputatus, Dicheirotrichus mannerheimi, Cymindis arcticus, Amara quenseli, Pterostichus abnormis, Pterostichus parens, Pterostichus cancellatus (173216) Pterostichus sp., Poecilus nearcticus, Curtonotus alpinus, Harpalus vittatus, Harpalus pusillus (163104) Pterostichus sp., Poecilus nearcticus, Pterostichus sublaevis, Curtonotus alpinus (163110 173211) Pterostichus sp., Poecilus nearcticus, Pterostichus tundrae, Curtonotus alpinus, Amara glacialis, Harpalus vittatus, Harpalus pusillus, Harpalus amputatus, Dicheirotrichus mannerheimi (153446) Pterostichus sp., Pterostichus agonus (173205 173206 173207) Pterostichus sp., Pterostichus costatus, Poecilus nearcticus, Pterostichus sublaevis, Curtonotus alpinus, Amara glacialis (173208) Pterostichus sp., Pterostichus costatus, Pterostichus sublaevis, Curtonotus alpinus, Harpalus amputatus, Dicheirotrichus mannerheimi (163102) Pterostichus spp., Poecilus nearcticus, Curtonotus alpinus (153435 153436 153437) Pterostichus spp., Poecilus nearcticus, Curtonotus alpinus, Harpalus vittatus, Cymindis arcticus (153439) Pterostichus spp., Poecilus nearcticus, Curtonotus alpinus, Harpalus vittatus, Harpalus amputatus, Cymindis arcticus (153438) Pterostichus spp., Poecilus nearcticus, Pterostichus sublaevis, Curtonotus alpinus, Harpalus amputatus, Dicheirotrichus mannerheimi (153441) Pterostichus spp., Poecilus nearcticus, Pterostichus sublaevis, Pterostichus tundrae, Curtonotus alpinus, Harpalus vittatus (153442) Pterostichus spp., Poecilus nearcticus, Pterostichus sublaevis, Pterostichus tundrae, Curtonotus alpinus, Harpalus vittatus, Harpalus pusillus (153443) Pterostichus spp., Pterostichus tundrae, Curtonotus alpinus (153444 153445)
Late/Upper Pleistocene0.129 - 0.0117USA (Illinois) Chlaenius plicatipennis (164922)
Late/Upper Pleistocene0.129 - 0.0117USA (Alaska) Pterostichus sp. (169385)
Late/Upper Pleistocene0.129 - 0.0117Canada (Ontario) Badister antecursor (152576) Pterostichus abrogatus, Pterostichus destitutus, Pterostichus fractus, Pterostichus destructus, Loxandrus gelidus (122182) Pterostichus abrogatus, Pterostichus destructus (152575) Pterostichus abrogatus, Pterostichus dormitans, Pterostichus fractus, Pterostichus depletus, Harpalus conditus (152577)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Worcestershire) Amara spp., Harpalus sp., Amara quenseli, Amara interstitialis, Harpalus cordatus, Amara municipalis, Amara erratica, Amara torrida, Feronia adstricta, Feronia nigrita, Feronia diligens, Calathus melanocephalus, Cymindis angularis (163070) Amara torrida (163075)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation (Astrakhan) Amara sp., Pterostichus sp., Chlaenius nigricornis, Amara aenea, Pterostichus vernalis (125794)
Late/Upper Pleistocene0.129 - 0.0117USA (Ohio) Pterostichus dormitans (152574)
Late/Upper Pleistocene0.129 - 0.0117USA (Pennsylvania) Harpalus sp., Pterostichus patruelis, Cymindis cribricollis, Pterostichus punctatissimus (137207) Pterostichus patruelis (137209 137212)
Late/Upper Pleistocene0.129 - 0.0117Canada (British Columbia) Cymindis sp., Cymindis cribricollis (167673)
Tarantian0.129 - 0.0117USA (Pennsylvania) Amara sp., Pterostichus patruelis, Cymindis cribricollis (137204) Pterostichus patruelis, Cymindis cribricollis (137206) Pterostichus sp., Pterostichus patruelis (137201)
Holocene0.0117 - 0.0Tanzania Helluomorpha protogaea, Chlaenius electrinus (129861)
Holocene0.0117 - 0.0New Zealand Plocamostethus planiusculus (175901)