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Priapulida (priapulid worm)

Synonymy list
YearName and author
1906Priapulida Théel
2002Priapulida Sepkoski
2011Priapulida Edgecombe et al. p. 153
2015Priapulida Zhang et al. p. 1

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
RankNameAuthor
CycloneuraliaAhlrichs 1995
Scaldiphora
phylumPriapulidaThéel 1906
phylumPriapulidaThéel 1906

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Ph. Priapulida Théel 1906 [priapulid worm]
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G. †Acosmia Chen and Zhou 1997
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Acosmia maotiania Chen and Zhou 1997
Unr. †Archaeopriapulida Conway Morris 1977
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Fm. †Fieldiidae Conway Morris 1977
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G. †Fieldia Walcott 1912
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Fieldia lanceolata Walcott 1912
G. †Lecythioscopa Conway Morris 1977
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Lecythioscopa simplex Conway Morris 1977
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Invalid names: Canadia simplex Walcott 1931 [replaced]
Fm. †Miskoidae Walcott 1911
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G. †Louisella
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Louisella pedunculata Conway Morris 1977
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Invalid names: Miskoia pedunculata Walcott 1911 [replaced]
G. †Miskoia Walcott 1911
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Miskoia preciosa Walcott 1911
Fm. †Ottoidae Walcott 1911
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G. †Banffia Walcott 1911
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Banffia constricta Walcott 1911
Banffia episoma Conway Morris et al. 2015
G. †Ottoia Walcott 1911
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Ottoia minor Walcott 1911
Ottoia prolifica Walcott 1911
Ottoia tenuis Walcott 1911
G. †Scolecofurca Conway Morris 1977
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Scolecofurca rara Conway Morris 1977
G. †Singuuriqia Peel 2017
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Singuuriqia simoni Peel 2017
Invalid names: Ancalagonidae Conway Morris 1977 [empty]
G. †Archotuba Hou et al. 1999
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Archotuba conoidalis Hou et al. 1999
G. †Corynetis Luo and Hu 1999
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Corynetis brevis Luo and Hu 1999
Corynetis fortis Hu et al. 2012
Invalid names: Anningvermis Huang et al. 2004 [synonym]
G. †Eximipriapulus Ma et al. 2014
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Eximipriapulus globocaudatus Ma et al. 2014
G. †Omnidens Hou and Others 2006
Unr. †Palaeoscolecidomorpha Shi et al. 2022
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+
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Cricocosmia jinningensis Hou and Sun 1988
G. †Houscolex
G. †Tabelliscolex Han et al. 2003
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Tabelliscolex chengjiangensis Han et al. 2007
Tabelliscolex hexagonus Han et al. 2003
G. †Tylotites Luo and Hu 1999
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Tylotites petiolaris Luo and Hu 1999
G. †Maotianshania Sun and Huo 1987
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Maotianshania cylindrica Sun and Huo 1987
G. †Markuelia Valkov 1983
Cl. †Palaeoscolecida Conway-Morris and Robison 1986
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G. †Corallioscolex Müller and Hintz-Schallreuter 1993
G. †Cowiella Hinz 1987
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Cowiella granulata Hinz 1987
Cowiella reticulata Hinz 1987
Cowiella undulata Hinz 1987
Cowiella verrucata Hinz 1987
Or. †Cricocosmida Han et al. 2007
+
+
G. †Paramaotianshania Hu et al. 2012
Fm. †Palaeoscolecidae Whittard 1953
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G. †Guanduscolex Hu et al. 2008
G. †Hadimopanella Gedik 1977
G. †Kaimenella Märss 1988
G. †Mafangscolex Hu 2005
G. †Palaeoscolex Whittard 1953
G. †Sahascolex Ivantsov and Wrona 2004
G. †Scathascolex Smith 2015
G. †Wronascolex Ivantsov and Zhuravlev 2005
Invalid names: Tylotitidae Han and Others 2003 [empty]
G. †Floraconformis Goñi et al. 2023
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Floraconformis egiinensis Goñi et al. 2023
G. †Milaculum
Invalid names: Palaeoscolecida [invalid subgroup]
Fm. Priapulidae Gosse 1855
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G. Acanthopriapulus van der Land 1970
G. †Priapulites Schram 1973
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Priapulites konecniorum Schram 1973
G. Priapulopsis Koren and Danielssen 1875
G. Priapulus Lamarck 1816
G. †Xiaoheiqingella Hu 2002
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+
Invalid names: Yunnanpriapulus halteroformis Huang et al. 2004 [synonym]
Invalid names: Yunnanpriapulus Huang et al. 2004 [synonym]
Or. †Priapulomorpha Slavini-Plawen 1974
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G. †Ancalagon Conway Morris 1977
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Ancalagon minor Walcott 1911
Or. †Selkirkiimorpha Adrianov and Malakhov 1995
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Fm. †Selkirkiidae Conway Morris 1977
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G. †Selkirkia Walcott 1911
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Selkirkia columbia Conway Morris 1977
Selkirkia fragilis Walcott 1911
Selkirkia major Walcott 1908
Selkirkia sinica Luo and Hu 1999
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Invalid names: Paraselkirkia jinningensis Hou et al. 1999 [synonym]
Selkirkia spencei Conway Morris and Robison 1986
Selkirkia transita Wang et al. 2021
Selkirkia tsering Nanglu and Ortega-Hernández 2024
Selkirkia willoughbyi Conway Morris and Robison 1986
Invalid names: Paraselkirkia Luo and Hu 1999 [synonym]
G. †Sullulika Peel and Willman 2018
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Sicyophorus rara Luo and Hu 1999
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Invalid names: Protopriapulites haikouensis Hou et al. 1999 [synonym]
Invalid names: Palaeopriapulites Hou et al. 1999 [synonym], Protopriapulites Hou et al. 1999 [synonym]
Fm. Tubiluchidae Van der Land 1970
+
G. †Paratubiluchus Han et al. 2004
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Paratubiluchus bicaudatus Han et al. 2004
G. Tubiluchus van der Land 1968
Invalid names: Halicryptidae Salvini-Plawen 1974 [empty], Maccabeidae Por and Bromley 1974 [empty], Xiaoheiqingidae Chen 2002 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: no hard partsp
Locomotion: facultatively mobilep
Life habit: shallow infaunalp
Diet: carnivorep
Vision: blindp
Comments: 535; LIFE HABIT: shallow infauna, facultatively mobile unattached or actively slow moving, predatory, mining, or suspension feeding according to Bambach et al. (2007).p
Created: 2009-10-08 23:14:00
Modified: 2010-02-15 06:03:57
Source: p = phylum
Reference: Hendy et al. 2009

Age range

Maximum range based only on fossils: base of the Meishucunian to the top of the Westphalian D or 530.70000 to 307.50000 Ma
Minimum age of oldest fossil (stem group age): 525.5 Ma

Collections (171 total)


Time interval Ma Country or state Original ID and collection number
Nemakit-Daldynian538.8 - 525.5Russian Federation (Siberia) Markuelia sp. (50340)
Early/Lower Cambrian538.8 - 509.0China (Yunnan) Paraselkirkia sp., Sicyophorus sp. (185938 185939) Xiaoheiqingella sp. (228573)
Waucoban538.8 - 509.0Canada (Northwest Territories) Palaeoscolecida indet. (227949)
Caerfai - St Davids538.8 - 497.0China (Yunnan) Maotianshania cylindrica (9940 9941) Maotianshania cylindrica, Palaeoscolex sinensis (9937) Palaeoscolex sinensis, Cricocosmia jinningensis (9939)
Meishucunian530.7 - 521.0China (Shaanxi) Houscolex lepidotus, Houscolex sp. (6851)
Tommotian525.5 - 521.0China (Yunnan) Maotianshania cylindrica (6817 6818 6819) Palaeoscolex sinensis (9784 9785 9786)
Tommotian525.5 - 521.0Russian Federation (Khabarovsk Krai) Markuelia secunda (50314)
Tommotian - Caerfai525.5 - 509.0China (Yunnan) Maotianshania cylindrica (9783)
Atdabanian521.0 - 516.0China (Yunnan) Cricocosmia jiningensis, Banffia confusa, Maotianshania cylindrica, Palaeoscolex sinensis, Tylotites petiolaris, Tabelliscolex hexagonus, Sicyophorus rara, Acosmia maotiania, Corynetis brevis, Paraselkirkia sinica (68083) Cricocosmia jiningensis, Maotianshania cylindrica, Palaeoscolex sinensis (86851) Cricocosmia jiningensis, Palaeoscolex sinensis, Tabelliscolex hexagonus (86845) Cricocosmia jinningensis, Sicyophorus rara (90136) Mafangscolex sinensis (90138) Mafangscolex sinensis, Cricocosmia jinningensis (90149 90150 90164 90165) Mafangscolex sinensis, Cricocosmia jinningensis, Sicyophorus rara, Corynetis brevis, Paraselkirkia sinica (90133 90135 90137) Mafangscolex sinensis, Cricocosmia jinningensis, Sicyophorus rara, Corynetis brevis, Xiaoheiqingella peculiaris, Paraselkirkia sinica (90151) Mafangscolex sinensis, Paraselkirkia sinica (90152) Maotianshania cylindrica (90494) Maotianshania cylindrica, Mafangscolex sinensis (90161 90163 90166) Maotianshania cylindrica, Palaeoscolex sinensis (9938) Maotianshania cylindrica, Palaeoscolex sinensis, Sicyophorus rara, Palaeopriapulites parvus (86846) Maotianshania sp., Acosmia sp. (86933) Priapulida indet., Cricocosmia sp., Palaeoscolex sp., Banffia sp., Maotianshania sp., Acosmia sp., Corynetis sp., Paraselkirkia sp., Sicyophorus sp., Anningvermis sp. (86929) Priapulida indet., Palaeoscolex sp., Maotianshania sp., Corynetis sp., Paraselkirkia sp., Protopriapulites sp. (86930)
Atdabanian521.0 - 516.0China (Hubei) Maotianshania cylindrica (68086)
Atdabanian521.0 - 516.0United Kingdom (England) Cowiella granulata (79940) Cowiella reticulata, Cowiella verrucata, Cowiella undulata (79933)
Atdabanian521.0 - 516.0Mongolia (Khovsgol) Floraconformis egiinensis (235542)
Atdabanian521.0 - 516.0Mongolia Markuelia sp. (219046)
Nangaoian521.0 - 514.0China (Hubai) Selkirkia sp., Banffia sp., Archotuba sp., Corynetis sp. (219687)
Nangaoian521.0 - 514.0China (Yunnan) Mafangscolex sinensi, Cricocosmia jinningensis, Corynetis brevis, Archotuba conoidalis, Paraselkirkia jinningensis (194160) Mafangscolex sinensis (132328) Mafangscolex sinensis, Cricocosmia jinningensis (194140) Mafangscolex sinensis, Cricocosmia jinningensis, Archotuba conoidalis (132329 194162) Mafangscolex sinensis, Cricocosmia jinningensis, Archotuba conoidalis, Paraselkirkia jinningensis (194150 194155) Mafangscolex sinensis, Cricocosmia jinningensis, Corynetis brevis, Palaeopriapulites parvus, Xiaoheiqingella peculiaris, Archotuba conoidalis, Paraselkirkia jinningensis (132330) Mafangscolex sinensis, Cricocosmia jinningensis, Palaeopriapulites parvus, Archotuba conoidalis, Paraselkirkia jinningensis (194143 194161 194163) Mafangscolex sinensis, Cricocosmia jinningensis, Xiaoheiqingella peculiaris, Archotuba conoidalis, Paraselkirkia jinningensis (194141) Mafangscolex sinensis, Cricocosmia jinningensis, Yunnanpriapulus halteroformis, Paraselkirkia jinningensis (194153) Mafangscolex sinensis, Tabelliscolex hexagonus, Cricocosmia jinningensis, Archotuba conoidalis (132327) Selkirkia sinica (234719)
Stage 3521.0 - 514.0China (Yunnan) Selkirkia transita (235078)
Atdabanian - Caerfai521.0 - 509.0Greenland Singuuriqia simoni (9780)
Branchian517.1 - 509.0United Kingdom (England) Cowiella reticulata (79935 79937) Hadimopanella apicata (79931) Milaculum sp. indet., Cowiella reticulata, Cowiella verrucata (79928)
Botomian516.0 - 512.9China (Yunnan) Guanduscolex minor (90357) Palaeoscolex sp. (90363 90367 90368 90369 90370 90371) Paramaotianshania zijunia, Corynetis fortis (139521)
Botomian516.0 - 512.9Australia (South Australia) Palaeoscolex antiquus, Wronascolex iacoborum (187498)
Botomian516.0 - 512.9Russian Federation (Siberia) Hadimopanella knappologica (174262)
Botomian516.0 - 512.9Russian Federation (Yakutia) Palaeoscolecida indet., Palaeoscolex sp., Corallioscolex sp., Palaeoscolex lubovae, Palaeoscolex spinosus, Sahascolex labyrinthus (90438)
Lenian516.0 - 510.2Antarctica (South Shetland Islands) Hadimopanella antarctica (49692)
Dyeran515.3 - 511.2USA (Nevada) Ottoia sp. (10560) Selkirkia sp. (9820)
Dyeran515.3 - 511.2USA (Pennsylvania) Ottoia sp., Selkirkia pennsylvanica (192 226194)
Dyeran515.3 - 511.2Greenland Hadimopanella apicata (49466 49467 49469 49470 49473 67782 67783 67787 67795 67797 67798 67799 67800)
Duyunian514.0 - 509.0China (Shaanxi) Palaeoscolecida indet. (90680)
Toyonian512.9 - 510.2Turkey (Konya) Hadimopanella oezgueli (93621)
Toyonian512.9 - 510.2Australia (South Australia) Kaimenella reticulata (64472 97986 97987 118171 118173)
Delamaran511.2 - 506.5USA (Utah) Selkirkia columbia, Palaeoscolex ratcliffei, Selkirkia spencei (77218) Selkirkia willoughby, Selkirkia sp. (77219)
Delamaran511.2 - 506.5USA (Idaho) Selkirkia sp. (207) Selkirkia spencei (76096)
Delamaran511.2 - 506.5Canada (British Columbia) Ottoia prolifica, Selkirkia columbia (84414 84416 84417 84434 84437) Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84423 84436) Priapulida indet. A, Selkirkia sp. A, Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84425) Priapulida indet. B, Fieldia lanceolata, Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84418) Selkirkia columbia (84413 84415 84420) Selkirkia sp. A, Ottoia prolifica, Selkirkia columbia (84421) Selkirkia sp. A, Ancalagon minor, Fieldia lanceolata, Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84424) Selkirkia sp. A, Fieldia lanceolata, Ottoia prolifica, Selkirkia columbia (84427) Selkirkia sp. A, Fieldia lanceolata, Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84422) Selkirkia sp. A, Ottoia prolifica, Selkirkia columbia (84412 84419 84426 84430 84431 84433) Selkirkia sp. A, Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84428 84429 84432 84435)
Amgan510.2 - 504.5Russian Federation (Siberia) Hadimopanella knappologica (110619)
Amgan510.2 - 504.5Kyrgyzstan Hadimopanella oezgueli (156378)
Solvan510.2 - 504.5Russian Federation (Siberia) Hadimopanella knappologica (49429)
Solvan510.2 - 504.5USA (Nevada) Ottoia sp. (9215)
Taijangian509.0 - 504.5China (Guizhou) Ottoia sp. (79770)
Middle Cambrian509.0 - 497.0USA (Utah) Banffia episoma (80122)
Middle Cambrian509.0 - 497.0Canada (British Columbia) Priapulida indet., Selkirkia sp., Banffia sp. (171308)
Middle Cambrian509.0 - 497.0Canada (Brithish Columbia) Selkirkia sp. (188664)
St Davids509.0 - 497.0Canada (British Columbia) Ottoia sp. (122017 122072) Ottoia sp., Ottoia prolifica (122094)
Drumian504.5 - 500.5USA (Utah) Scathascolex minor, Ottoia prolifica, Selkirkia willoughbyi (230679)
Menevian504.5 - 499.5USA (Utah) Ottoia sp., Selkirkia sp., Palaeoscolecida gen. et sp. nov. A (230681)
Marjumian504.5 - 497.0USA (Utah) Ottoia prolifica (77206)
Guzhangian500.5 - 497.0China (Yunnan) Palaeoscolecida indet. (219647)
Merioneth497.0 - 485.4Russian Federation (Siberia) Protoscolex tchopkoensis (10009)
Tremadocian485.4 - 477.7Morocco (Zagora) Selkirkia tsering (235072)
Floian477.7 - 470.0China (Hunan) Ottoia sp., Palaeoscolecida indet. (226639)
Darriwilian467.3 - 458.4United Kingdom (Wales) Priapulida NMW.2021.3 G.97, Palaeoscolecida NMW.2021.3 G.89, Archaeopriapulida NMW.2021.3 G.19 (230215)
Llandeilo461.1 - 458.4United Kingdom (Wales) Palaeoscolecidae gen. et sp. nov. Botting et al. in press (216511)
Caradoc458.4 - 449.5USA (New York) Protoscolex sp. (340)
Turinian457.3 - 453.8Canada (British Columbia) Milaculum sp. (192195)
Rocklandian457.3 - 449.6USA (Wisconsin) Milaculum sp. (90362)
Rocklandian - Shermanian457.3 - 449.6USA (Wisconsin) Milaculum sp. (90360)
Shermanian457.3 - 449.6USA (Wisconsin) Milaculum sp. (90366)
Edenian451.0 - 450.2USA (Ohio) Protoscolex sp. (383 384)
Maysvillian450.2 - 449.6Canada (Ontario) Protoscolex sp. (26302)
Richmondian449.6 - 445.5Canada (Nunavut) Milaculum ethinclarki (360)
Rhuddanian - Aeronian443.4 - 438.5USA (Iowa) Protoscolex sp. (26860)
Telychian438.5 - 433.4USA (Wisconsin) Palaeoscolecida indet. (91415)
Ludlow427.4 - 423.0USA (Illinois) Protoscolex ruedemanni (76450) Protoscolex sp. (26861)
Westphalian D309.8 - 307.5USA (Illinois) Priapulites konecniorum (122493 122538 151852 168004)