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Scarabaeinae

Insecta - Coleoptera - Scarabaeidae

Synonyms
Synonymy list
YearName and author
1802Scarabaeinae Latreille p. 144
1870Elytridium rugulosum Heer p. 78 figs. Pl 16, figs 36,37
1977Onthophagus rugulosus Birket-Smith p. 25 fig. 11
1995Scarabaeinae Lawrence and Newton, Jr.
2000Scarabaeinae Krell p. 879
2003Scarabaeinae Wappler
2007Scarabaeinae Nikolajev p. 130
2010Onthophagus spitsbergeniensis Krell
2011Scarabaeinae Bouchard et al. p. 242
2016Scarabaeinae Tarasov and Dimitrov
2016Scarabaeinae Tarasov et al. p. 7
2022Scarabaeinae Cai et al.
2023Scarabaeinae Tello et al. p. 1315

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
RankNameAuthor
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderPolyphagaEmery 1886
infraorderStaphyliniformia
superfamilyScarabaeoidea
familyScarabaeidae()
subfamilyScarabaeinae
subfamilyScarabaeinae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Scarabaeinae Latreille 1802
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G. Anachalcos Hope 1837
+
†Anachalcos mfwangani Paulian 1976
Tr. Ateuchini Perty 1830
+
Subtr. Ateuchina Perty 1830
+
G. Ateuchus Weber 1801
+
Invalid names: Choeridium Lepeletier de Saint-Fargeau and Audinet-Serville 1828 [synonym], Choeridium ebeninum Horn 1876 [nomen dubium]
G. †Lobateuchus Montreuil et al. 2010
+
†Lobateuchus parisii Montreuil et al. 2010
G. Canthochilum Chapin 1934
+
†Canthochilum alleni Tarasov and Vaz-de-Mello 2016
†Canthochilum philipsivieorum Tarasov and Vaz-de-Mello 2016
Tr. Coprini Leach 1815
+
G. Copris Geoffroy 1762
+
Subg. Copris (Copris) Geoffroy 1762
+
†Copris (Copris) kartlinus Kabakov 1988
†Copris druidum Heer 1862
Copris incertus Say 1835
†Copris leakeyorum Paulian 1976
Copris lunaris Linnaeus 1758 [horned dung beetle]
†Copris pristinus Pierce 1946
Tr. Deltochilini Lacordaire 1856
+
G. Canthon Hoffmannsegg 1817
+
G. Deltochilum Eschscholtz 1822
G. Scybalophagus Martínez 1953
+
†Scybalophagus brellenthinae Tello et al. 2023
Tr. Gymnopleurini Lacordaire 1856
+
G. Gymnopleurus Illiger 1803
+
†Gymnopleurus sisyphus Heer 1847
G. Heliocopris Hope 1837
+
†Heliocopris antiquus Fujiyama 1968
G. Metacatharsius Paulian 1939
+
†Metacatharsius rusingae Paulian 1976
Tr. Oniticellini Kolbe 1905
+
Subtr. Drepanocerina van Lansberge 1875
+
G. Eodrepanus Barbero et al. 2009
+
†Eodrepanus coopei Barbero et al. 2009
Subtr. Oniticellina Kolbe 1905
+
G. Oniticellus Dejean 1821
Tr. Onitini Laporte 1840
+
G. Onitis Fabricius 1798
Tr. Onthophagini Burmeister 1846
+
G. Onthophagus Latreille 1802
+
†Onthophagus arvernus Piton and Théobald 1939
†Onthophagus crassus Heer 1862
†Onthophagus everestae Pierce 1946
Onthophagus fracticornis Preyssler 1790
Onthophagus janus Panzer 1794
Onthophagus lenzii Harold 1874
†Onthophagus ovatulus Heer 1847
†Onthophagus pilauco Tello et al. 2021
†Onthophagus prodromus Heer 1862
†Onthophagus statzi Krell 1990
+
Invalid names: Onthophagus muelleri Statz 1952 [replaced]
Onthophagus taurus Schreber 1759
G. Phalops Erichson 1848
Tr. Phanaeini Hope 1838
+
G. Phanaeus Macleay 1819
+
†Phanaeus labreae Pierce 1946
†Phanaeus violetae Zunino 2013
Invalid names: Palaeocopris Pierce 1946 [synonym], Phanaeus antiquus Horn 1876 [nomen dubium]
Tr. Scarabaeini Latreille 1802
+
G. Scarabaeus Linnaeus 1758
Invalid names: Onthophagus spitsbergeniensis Krell 2010 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Danian to the top of the Rancholabrean or 66.00000 to 0.01200 Ma
Minimum age of oldest fossil (stem group age): 59.2 Ma

Collections (45 total)


Time interval Ma Country or state Original ID and collection number
Danian - Selandian66.0 - 59.2Svalbard and Jan Mayen Elytridium rugulosum (122011)
Ypresian56.0 - 47.8United Kingdom (England) Onthophagus sp. (126591)
Early/Lower Eocene55.8 - 48.6France Lobateuchus parisii (123459)
MP 1148.6 - 40.4Germany (Hessen) Onitini indet., Onthophagini indet. (120798)
MP 1348.6 - 40.4Germany (Eifel) Scarabaeinae indet. (24201)
Late/Upper Oligocene28.4 - 23.03France (Puy-de-Dome) Heliocopris sp., Onthophagus arvernus (155204)
Chattian28.1 - 23.03Germany (North Rhine-Westphalia) Onthophagus muelleri (122010)
Early/Lower Miocene23.03 - 15.97Japan Heliocopris antiquus (160137)
Burdigalian20.44 - 15.97China (Shandong) Onthophagus sp., Phalops sp. (128552)
Burdigalian20.44 - 15.97Kenya Anachalcos mfwangani (113556) Copris leakeyorum, Metacatharsius rusingae (113557)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Canthochilum alleni, Canthochilum philipsivieorum (146610)
Sarmatian12.7 - 11.608Germany (Baden-Württemberg) Gymnopleurus sisyphus, Onthophagus prodromus, Gymnopleurus rotundatus, Copris druidum, Onthophagus crassus, Onthophagus bisontinus (117252) Onthophagus ovatulus (115304)
Early/Lower Pliocene5.333 - 3.6Georgia Copris kartlinus (145761)
Zanclean5.333 - 3.6Japan Onthophagus sp. (165737)
Piacenzian3.6 - 2.588Germany (Niedersachsen) Coprinae indet. (109196)
Piacenzian3.6 - 2.588Japan Onthophagus sp. (165738 165742)
Early/Lower Pleistocene2.588 - 0.781Japan Onthophagus sp. (153617)
Irvingtonian1.8 - 0.3USA (Pennsylvania) Phanaeus antiquus, Choeridium ebeninum (20479)
Lujanian0.781 - 0.012Chile (Los Lagos) Onthophagus pilauco, Scybalophagus brellenthinae (195456)
Chibanian0.774 - 0.129Japan (Kumamoto) Onthophagus sp. (164901)
Chibanian0.774 - 0.129Japan Onthophagus sp. (165704)
Middle Pleistocene0.774 - 0.129United Kingdom Copris lunaris, Onthophagus taurus, Onthophagus fracticornis (168266) Onthophagus fracticornis (168267 168272) Onthophagus sp. (168269 168271 168273) Onthophagus sp., Onthophagus fracticornis (168264)
Middle Pleistocene0.774 - 0.129Japan (Kumamoto) Onthophagus sp. (165716)
Rancholabrean0.3 - 0.012USA (California) Canthon simplex, Copris pristinus (152007) Copris pristinus (152008 152009) Onthophagus sp., Copris sp., Canthon sp., Canthon praticola, Onthophagus everestae, Paleocopris labreae, Deltochilum sp. (152006)
Late/Upper Pleistocene0.129 - 0.0117Chile (Los Lagos) Scybalophagus brellenthinae (220821)
Late/Upper Pleistocene0.129 - 0.0117Ecuador Phanaeus violetae (178693)
Late/Upper Pleistocene0.129 - 0.0117USA (Virginia) Onthophagus sp., Canthon sp., Onthophagus janus (103991)
Late/Upper Pleistocene0.129 - 0.0117Japan Onthophagus sp. (166083) Onthophagus sp., Onthophagus lenzii (166084) Onthophagus spp. (166082)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom Copris lunaris (152473) Eodrepanus coopei (93819)
Late/Upper Pleistocene0.129 - 0.0117Peru Copris incertus (71346)