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Taxonomy
Heteroptera was named by Latreille (1810). It is extant.
It was assigned to Insecta by Bode (1953) and Hong (1984); to Hemineoptera by Kukalová-Peck and Lawrence (2004); to Hemiptera by Tillyard (1918), Tillyard (1922), Cockerell (1926), Whalley (1985), Popov et al. (1994), Grimaldi and Engel (2005), Vega et al. (2006), Zhang et al. (2011) and Johnson et al. (2018); and to Heteropterodea by Cryan and Urban (2012) and Szwedo (2018).
It was assigned to Insecta by Bode (1953) and Hong (1984); to Hemineoptera by Kukalová-Peck and Lawrence (2004); to Hemiptera by Tillyard (1918), Tillyard (1922), Cockerell (1926), Whalley (1985), Popov et al. (1994), Grimaldi and Engel (2005), Vega et al. (2006), Zhang et al. (2011) and Johnson et al. (2018); and to Heteropterodea by Cryan and Urban (2012) and Szwedo (2018).
Subtaxa
Anacoloptera, Berytopsis, Cimicomorpha, Corixa pullus, Cryptocerata, Dipsocoromorpha, Duncanovelia, Enicocephalomorpha, Eocimicidae, Euheteroptera, Galerucites, Galgulidae, Geocorisae, Gerromorpha, Gymnocerata, Hadrocoridae, Hebeicoris, Hydrocorisae, Leptopodomorpha, Liasocoris, Nepa atavina, Nepomorpha, Pentatomomorpha, Sinanabis, Sinocoris, Sinovelia, Sphongophoriella, Xishanidae
Synonyms
|
Synonymy list
| Year | Name and author |
|---|---|
| 1810 | Heteroptera Latreille p. 433 |
| 1853 | Nepa atavina Heer p. 85 figs. Pl 10, fig 9 |
| 1859 | Corixa pullus Von Heyden p. 10 figs. Pl 1,fig 13 |
| 1918 | Heteroptera Tillyard |
| 1922 | Heteroptera Tillyard |
| 1926 | Heteroptera Cockerell |
| 1953 | Heteroptera Bode |
| 1984 | Heteroptera Hong p. 2 |
| 1985 | Heteroptera Whalley |
| 1994 | Heteroptera Popov et al. p. 320 |
| 2004 | Heteroptera Kukalová-Peck and Lawrence |
| 2005 | Heteroptera Grimaldi and Engel |
| 2006 | Heteroptera Vega et al. p. 327 |
| 2011 | Heteroptera Zhang et al. p. 492 |
| 2012 | Heteroptera Cryan and Urban |
| 2018 | Heteroptera Johnson et al. |
| 2018 | Heteroptera Szwedo p. 111 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subor. Heteroptera Latreille 1810
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Infraor. Cimicomorpha Leston et al. 1954
+
Fm. †Berstidae Tihelka et al. 2020
+
G. †Bersta Tihelka et al. 2020
+
†Bersta coleopteromorpha Tihelka et al. 2020
†Bersta vampirica Tihelka et al. 2020
Superfm. Cimicoidea Latreille 1802
+
Fm. Anthocoridae Fieber 1837 [minute pirate bug]
+
Subfm. Anthocorinae Fieber 1837
+
Tr. Anthocorini Fieber 1837
Tr. Cardiastethini Carayon 1972
G. †Occidoscelis Ryzhkova 2020
G. †Electrocoris Usinger 1942
+
†Electrocoris brunneus Usinger 1942
†Electrocoris pubescens Usinger 1942
Fm. Lyctocoridae Reuter 1884
+
G. †Eoanthocoris Popov 1990
+
†Eoanthocoris cretaceus Popov 1990
†Eoanthocoris ghidarinus Popov 1990
Fm. †Torrirostratidae Yao et al. 2014
+
Fm. Velocipedidae Bergroth 1891
+
Fm. †Vetanthocoridae Yao et al. 2006
+
Tr. †Crassicerini Yao et al. 2006
+
G. †Crassicerus Yao et al. 2006
G. †Curticerus Yao et al. 2006
G. †Pustulithoracalis Yao et al. 2006
Tr. †Vetanthocorini Yao et al. 2006
+
G. †Byssoidecerus Yao et al. 2006
G. †Collivetanthocoris Yao et al. 2006
G. †Curvicaudus Yao et al. 2006
G. †Liaoxia Hong 1987
G. †Longilanceolatus Tang et al. 2015
G. †Mecopodus Yao et al. 2006
G. †Pubivetanthocoris Tang et al. 2021
G. †Pumilanthocoris Hou et al. 2012
G. †Punctivetanthocoris Tang et al. 2016
G. †Vetanthocoris Yao et al. 2006
Superfm. Miroidea Hahn 1833
+
Fm. Miridae Hahn 1833 [plant bug]
+
Subfm. Cylapinae Kirkaldy 1903
+
G. †Amberofulvius Herczek 1991
G. †Ambocylapus Herczek and Popov 2000
G. †Aragocylapus Herczek et al. 2000
G. †Archeofulvius Carvalho 1966
G. †Balticofulvius Herczek and Popov 1997
Tr. Cylapini Kirkaldy 1903
Tr. Fulviini Uhler 1886
G. †Fulviocylapus Herczek and Popov 2005
G. †Germarofulvius Herczek and Popov 1999
G. †Oligocoris Jordan 1944
G. †Popoviana Wichard 1997
G. †Samlandia Herczek and Popov 2005
Subfm. Deraeocorinae Douglas and Scott 1865
+
Tr. Deraeocorini Douglas and Scott 1865
Tr. Hyaliodini Knight 1943
Invalid names: Surinamellini Carvalho and Rosas 1962 [empty]
Subfm. Isometopinae Fieber 1860
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Tr. Diphlebini Bergroth 1924
Tr. †Electromyiommini Herczek 1993
Tr. Gigantometopini Herczek 1993
G. †Isomyiomma Herczek et al. 2020
Tr. Myiommini Bergroth 1924
Subfm. Mirinae Hahn 1833
+
Tr. †Armanomirini Popov and Herczek 1998
Tr. †Cretamirini Popov and Herczek 1998
Tr. Herdoniini Distant 1904
Tr. Mecistoscelini Reuter 1910
Tr. †Scutelliferini Popov 1968
Invalid names: Phytocoris consobrinus Germar and Berendt 1856 [nomen dubium], Phytocoris involutus Germar and Berendt 1856 [nomen dubium]
Subfm. Phylinae Douglas and Scott 1865
+
G. †Gurnardinia Popov 2019
Tr. Hallodapini Van Duzee 1916
Tr. Phylini Douglas and Scott 1865
Invalid names: Phytocoris angustulus Germar and Berendt 1856 [nomen dubium]
Subfm. Psallopinae Schuh 1976
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G. †Cylapopsallops Popov and Herczek 2006
G. †Epigonomiris Herczek and Popov 1998
G. †Epigonopsallops Herczek and Popov 2009
G. †Isometopsallops Herczek and Popov 1992
G. Psallops Usinger 1946
Invalid names: Capsidae Burmeister 1835 [synonym], Fushunimiris Hong 2002 [nomen nudum], Phytocoris vetustus Germar and Berendt 1856 [nomen dubium]
Fm. Tingidae Laporte 1833 [lace bug]
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G. †Burmacader Heiss and Guilbert 2013
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†Burmacader bicoloripennis Souma et al. 2021
†Burmacader grandis Heiss and Guilbert 2021
†Burmacader lativentris Heiss and Guilbert 2018
†Burmacader multivenosus Heiss and Guilbert 2013
G. †Burmavianaida Souma et al. 2021
+
†Burmavianaida anomalocapitata Souma et al. 2021
+
Invalid names: Microtingis leptosoma Heiss and Golub 2021 [synonym]
Invalid names: Microtingis Heiss and Golub 2021 [synonym]
Subfm. Cantacaderinae Stal 1873
+
G. Cantacader Amyot and Audinet-Serville 1843
Tr. Cantacaderini Stal 1873
Tr. †Golmoniini Popov 1989
G. †Sinaldocader Popov 1989
+
†Sinaldocader drakei Popov 1989
†Sinaldocader ponomarenkoi Golub and Popov 2008
Subfm. Tinginae Laporte 1833
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G. Celantia Distant 1903
G. Cysteochila Stål 1873
G. Gitava Drake 1948
G. Hovatlas Schouteden 1957
G. Leptopharsa Stal 1873
Tr. Phatnomatini Drake and Davis 1960
Tr. Phatnomini Drake and Davis 1960
G. Phymacysta Monte 1942
Tr. Tingini Laporte 1833
+
Invalid names: Litadeini Drake and Ruhoff 1965 [synonym], Ypsotingini Drake and Ruhoff 1965 [synonym]
G. Uhlerites Drake 1927
G. †Viktorgolubia Popov 2019
Subfm. †Tingiometrinae Heiss et al. 2015
+
G. †Paleoanomala Poinar and Vega 2020
G. †Tingiometra Heiss et al. 2015
Subfm. Vianaidinae Kormilev 1955
+
G. †Vianagramma Golub and Popov 2000
G. †Vianathauma Golub and Popov 2003
Invalid names: Sinaldocaderini Popov and Golub 2019 [invalid subgroup]
Invalid names: Tingoidea Laporte 1833 [synonym]
Superfm. Reduvioidea Latreille 1807
+
Fm. †Ceresopseidae Becker-Migdisova 1958
+
G. †Ceresopsis Becker-Migdisova 1958
+
†Ceresopsis costalis Becker-Migdisova 1958
†Ceresopsis ornata Becker-Migdisova 1962
†Ceresopsis vitrea Becker-Migdisova 1962
Fm. †Palaeotanyrhinidae Poinar et al. 2022
+
Fm. Reduviidae Latreille 1807 [assassin bug]
+
Subfm. Emesinae Amyot and Audinet-Serville 1843 [thread-legged bug]
+
Tr. Collartidini Wygodzinsky 1966
Tr. Emesini Amyot and Audinet-Serville 1843
+
Invalid names: Metapterini Stal 1874 [synonym], Ploiariolini Van Duzee 1916 [synonym]
Subfm. Harpactorinae Amyot and Audinet-Serville 1843
+
Tr. Apiomerini Amyot and Audinet-Serville 1843
Tr. Harpactorini Amyot and Audinet-Serville 1843
Subfm. Holoptilinae Amyot and Audinet-Serville 1843
+
Tr. Dasycnemini Wygodzinsky and Usinger 1963
Tr. Holoptilini Amyot and Audinet-Serville 1843
Subfm. Reduviinae Latreille 1807 [assassin bug]
+
G. Platymeris Laporte 1832
G. Reduvius Fabricius 1775
G. †Simplicivenius Zhang et al. 2022
Subfm. Stenopodainae Amyot and Audinet-Serville 1843
+
G. Oncocephalus Klug 1830
G. Pygolampis Germar 1817
Superfm. Thaumastocoroidea Kirkaldy 1908
+
Fm. Thaumastocoridae Kirkaldy 1908 [palm bug]
+
Subfm. Xylastodorinae Barber 1920
+
G. Discocoris Kormilev 1955
G. †Paleodoris Poinar and Santiago-Blay 1997
G. †Protodoris Nel et al. 2004
G. †Proxylastodoris Heiss and Popov 2002
G. †Thaumastotingis Heiss and Golub 2015
G. Xylastodoris Barber 1920
Infraor. Dipsocoromorpha Miyamoto 1961
+
Superfm. Dipsocoroidea
+
Fm. Ceratocombidae Fieber 1861
+
Fm. †Cuneocoridae Handlirsch 1920
+
Fm. Dipsocoridae Dohrn 1859
+
Fm. Hypsipterygidae Drake 1961
+
Fm. Schizopteridae Reuter 1891 [jumping soil bug]
+
Subfm. Hypselosomatinae Esaki and Miyamoto 1959
+
G. †Buzinia Perrichot et al. 2007
G. †Hypsotsijilia Cifuentes-Ruiz and Brailovsky 2021
G. †Kachinia Chen and Wang 2018
G. †Leptosaldinea Popov and Heiss 2016
G. †Libanohypselosoma Azar and Nel 2010
G. †Tanaia Perrichot et al. 2007
G. Schizoptera Fieber 1860
+
†Schizoptera dominicana Poinar 2015
†Schizoptera hispaniolae Poinar 2015
Unr. Enicocephalomorpha
+
Fm. Aenictopecheidae Usinger 1932
+
Fm. Enicocephalidae Stal 1860 [unique-headed bug]
+
Subfm. Alienatinae Barber 1953
+
G. Alienates Barber 1953
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†Alienates brodzinskyi Grimaldi et al. 1993
†Alienates electrodominicus Grimaldi et al. 1993
G. †Paralienates Maldonado Capriles et al. 1996
+
†Paralienates hyalinus Maldonado Capriles et al. 1996
Subfm. †Allocephalocorinae Luo et al. 2020
+
Subfm. †Disphaerocephalinae Štys 1969
+
G. †Disphaerocephalus Cockerell 1917
+
†Disphaerocephalus constrictus Cockerell 1917
†Disphaerocephalus macropterus Cockerell 1917
†Disphaerocephalus swinhoei Cockerell 1917
Subfm. Enicocephalinae Stal 1860
+
G. Enicocephalus Westwood 1837
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†Enicocephalus omen Grimaldi et al. 1993
†Enicocephalus prius Grimaldi et al. 1993
†Enicocephalus seniculus Grimaldi et al. 1993
†Enicocephalus subvitreus Grimaldi et al. 1993
Invalid names: Enicocephalus fulvescens Westwood 1837 [nomen dubium]
G. †Enicocephalinus Azar et al. 1999
+
†Enicocephalinus acragrimaldii Azar et al. 1999
†Enicocephalinus ibericus Davranoglou et al. 2024
Infraor. Gerromorpha Popov 1971 [semi-aquatic bug]
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Superfm. Gerroidea Leach 1815
+
Fm. Gerridae Leach 1815 [water strider]
+
Subfm. Gerrinae Leach 1815
+
G. Aquarius Schellenberg 1800
G. †Electrogerris Andersen 2000
G. Gerris Fabricius 1794 [water strider]
G. Limnoporus Stal 1868
G. †Palaeogerris Andersen 1998
G. †Telmatrechus Scudder 1890
Subfm. Halobatinae Bianchi 1896
+
G. Halobates Eschscholtz 1829 [sea skater]
Invalid names: Limnobates prodromus Heer 1864 [nomen dubium]
G. †Succineogerris Andersen 2000
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†Succineogerris larssoni Andersen 2000
†Succineogerris nilsi Zettel and Heiss 2011
Invalid names: Gerris parabdominalis Theobald 1937 [nomen dubium], Gerris protobates Cockerell 1927 [nomen dubium], Hydrobatida Stål 1866 [synonym], Metrobates aeternalis Scudder 1890 [nomen dubium]
Invalid names: Hebroveliidae Lundblad 1939 [empty]
Superfm. Hebroidea Amyot and Audinet-Serville 1843
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Fm. Hebridae Amyot and Audinet-Serville 1843 [velvet water bug]
+
Superfm. Hydrometroidea Billberg 1820
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Fm. Hydrometridae Billberg 1820 [marsh treader]
+
Subfm. Heterocleptinae Villiers 1948
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G. †Alavametra Sánchez-García and Nel 2016
G. †Carinametra Andersen and Grimaldi 2001
Subfm. Hydrometrinae Billberg 1820
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G. †Burmametra Huang et al. 2015
G. †Christometra Pêgas et al. 2017
G. †Cretaceometra Nel and Popov 2000
G. †Eocenometra Andersen 1982
G. †Incertametra Perez Goodwyn 2002
G. †Limnacis Germar 1856
G. †Metrocephala Popov 1996
G. †Palaeometra Andersen 1998
G. †Protobacillometra Nel and Paicheler 1993
Invalid names: Limnobatodinae Esaki 1927 [empty]
Fm. Macroveliidae Mac Kinstry 1942
+
Superfm. Mesovelioidea Douglas and Scott 1867
+
Fm. Mesoveliidae Douglas and Scott 1867 [water treader]
+
G. †Glaesivelia Sánchez-García and Solórzano Kraemer 2017
+
†Glaesivelia pulcherrima Sánchez-García and Solórzano Kraemer 2017
Subfm. Mesoveliinae Douglas and Scott 1867
+
G. †Emilianovelia Solórzano Kraemer and Perrichot 2014
G. †Malenavelia Solórzano Kraemer and Perrichot 2014
G. Mesovelia Mulsant and Rey 1852
Fm. Veliidae Amyot and Audinet-Serville 1843 [riffle bug]
+
Subfm. Haloveliinae Esaki 1930
+
Subfm. Microveliinae
+
G. Microvelia Westwood 1834
+
†Microvelia electra Andersen 2001
†Microvelia grimaldii Andersen 2001
†Microvelia polhemi Andersen 2000
Subfm. Ocelloveliinae Drake and Chapman 1963
+
Subfm. Veliinae Amyot and Audinet-Serville 1843
+
G. †Arcantivelia Solórzano Kraemer and Perrichot 2014
+
†Arcantivelia petraudi Solórzano Kraemer and Perrichot 2014
G. Velia Latreille 1804 [water cricket]
Fm. †Hadrocoridae Handlirsch 1920
+
G. †Hebeicoris Hong 1983
+
†Hebeicoris longa Hong 1983
†Hebeicoris luanpingensis Hong 1983
†Hebeicoris xinboensis Hong 1984
Infraor. Hydrocorisae Latreille 1810
+
Invalid names: Nepaeformes Reuter 1910 [empty], Notonectaeformia [empty]
Infraor. Leptopodomorpha Popov 1971 [shore bug]
+
Superfm. Leptopodoidea Brullé 1836
+
Fm. Leptopodidae Brullé 1836 [spiny-legged bug]
+
Subfm. Leptosaldinae Cobben 1971
+
G. †Archaesalepta Grimaldi and Engel 2013
G. †Cretaceomira McKellar and Engel 2014
G. †Cretaleptus Sun and Chen 2020
G. †Leptosalda Cobben 1971
G. †Macrolepta Yu et al. 2023
G. †Parvilepta Yu et al. 2023
Invalid names: Palaeoleptinae Poinar 2009 [synonym]
Invalid names: Leptopodinae Brullé 1836 [empty]
Invalid names: Leotichiidae China 1933 [empty], Omaniidae Cobben 1970 [empty]
Superfm. Saldoidea Amyot and Audinet-Serville 1843
+
Fm. †Archegocimicidae Handlirsch 1906
+
G. †Archegocimex Handlirsch 1906
+
†Archegocimex geinitzi Handlirsch 1906
†Archegocimex liadis Handlirsch 1939
†Archegocimex primitiva Handlirsch 1920
Invalid names: Archegocoris Handlirsch 1939 [synonym], Eonabis Handlirsch 1920 [synonym]
G. †Diatillus Handlirsch 1921
G. †Entomecoris Bode 1953
+
†Entomecoris major Bode 1953
†Entomecoris minor Bode 1953
†Entomecoris morator Bode 1953
G. †Macropterocoris Bode 1953
G. †Saldomortalia Popov 1988
+
†Saldomortalia hyalina Popov 1988
†Saldomortalia zherichini Popov 1988
G. †Saldonia Popov 1973
+
†Saldonia formosa Ryzhkova 2011
†Saldonia insolita Ryzhkova 2011
†Saldonia maculata Popov 1985
†Saldonia rasnitsyni Popov 1973
†Saldonia sibirica Popov 1985
G. †Shartegocimex Ryzhkova 2011
+
†Shartegocimex distans Ryzhkova 2011
†Shartegocimex rasnitsyni Ryzhkova 2011
G. †Tyrion Ryzhkova and Coram 2016
+
†Tyrion cersei Ryzhkova and Coram 2016
†Tyrion lannister Ryzhkova and Coram 2016
Invalid names: Corynecoris semigranulatus Bode 1953 [nomen dubium], Diatillidae Handlirsch 1921 [synonym], Diatillus debilis Handlirsch 1921 [nomen dubium], Ensphingocoris praerotundatus Bode 1953 [nomen dubium], Eonabidae Handlirsch 1920 [synonym], Macropterocoris obtusus Bode 1953 [nomen dubium], Mesolygaeidae Zhang 1991 [synonym], Saldoniinae Popov 1973 [invalid subgroup], Xishaniidae Hong 1981 [synonym]
Fm. Saldidae Amyot and Audinet-Serville 1843 [shore bug]
+
Subfm. Chiloxanthinae Cobben 1959
+
G. †Brevrimatus Zhang et al. 2011
G. †Luculentsalda Zhang et al. 2013
G. †Oligosaldina Statz and Wagner 1950
G. †Olindasalda Moura-Júnior et al. 2020
G. Pentacora Reuter 1912
G. †Propentacora Polhemus 1985
G. †Venustsalda Zhang et al. 2012
Subfm. †Enicocorinae Popov 1980
+
G. †Baissotea Ryzhkova 2015
G. †Mesolygaeus Ping 1928
+
Invalid names: Enicocoris Popov 1980 [synonym], Jiaodongia Hong 1984 [synonym], Sinolygaeus Hong 1980 [synonym], Xishania Hong 1981 [synonym]
G. †Mongolocoris Ryzhkova 2012
+
Invalid names: Longianteclypea Zhang et al. 2013 [objective synonym]
G. †Propritergum Zhang et al. 2013
G. †Ulanocoris Ryzhkova 2012
Subfm. Saldinae Amyot and Audinet-Serville 1843
+
Tr. Saldini Amyot and Audinet-Serville 1843
Tr. Saldoidini Reuter 1912
Invalid names: Acanthiidae Leach 1815 [synonym]
Invalid names: Aepophilidae Puton 1879 [empty]
Invalid names: Lygaeites mysteriosus Meunier 1919 [nomen dubium]
Infraor. Nepomorpha Popov 1971
+
Superfm. Corixoidea Enderlein 1912
+
Fm. Corixidae Leach 1815 [water boatman]
+
Subfm. †Archaecorixinae Popov 1968
+
G. †Archaecorixa Popov 1968
G. †Huatingicorixa Hong 1995
G. †Liassocorixa Popov et al. 1994
G. †Mesocorixa Hong and Wang 1990
G. †Mesosigara Popov 1971
G. †Rhomboidella Mazzoni and Hunicken 1987
Subfm. †Diapherininae Popov 1988
+
Subfm. Diaprepocorinae Lunblad 1928
+
G. †Gazimuria Popov 1971
G. †Karataviella Becker-Migdisova 1949
Subfm. Micronectinae Jaczewski 1924
+
G. †Acromocoris Bode 1953
+
Invalid names: Dichaspis Bode 1953 [synonym], Strobilocoris Bode 1953 [synonym]
G. †Sigaretta Popov 1971
+
†Sigarella tenuis Hong and Wang 1990
†Sigaretta florissantella Cockerell 1906
Subfm. †Velocorixinae Popov 1986
+
G. †Bakharia Popov 1988
G. †Bumbacorixa Popov 1986
G. †Corixalia Popov 1986
G. †Coryphocorixa Popov 2014
G. †Cristocorixa Popov 1986
G. †Daohugocorixa Zhang 2010
G. †Haenbea Popov 1988
G. †Jiulongshanocorixa Zhang 2010
G. †Shelopuga Popov 1988
G. †Velocorixa Popov 1986
G. †Yanliaocorixa Hong 1983
Invalid names: Corixa boryslavica Lomnicki 1894 [nomen dubium], Corixa elegans von Schlechtendal 1894 [nomen dubium], Corixa fasciolata Heer 1853 [nomen dubium], Corixa glacialis Lomnicki 1894 [nomen dubium], Corixa immersa Scudder 1890 [nomen dubium], Corixa rhenana Statz 1950 [nomen dubium], Corixa vanduzeei Scudder 1890 [nomen dubium], Corixa vidua Statz 1950 [nomen dubium]
Fm. †Shurabellidae Popov 1971
+
G. †Shurabella Becker-Migdisova 1949
+
†Shurabella lepyroniopsis Becker-Migdisova 1949
+
Invalid names: Coleopteropsis dolichoptera Becker-Migdisova 1949 [synonym]
Invalid names: Coleopteropsis Becker-Migdisova 1949 [synonym]
Superfm. Naucoroidea
+
Fm. Aphelocheiridae Fieber 1851
+
G. Aphelocheirus Westwood 1833
+
Subg. Aphelocheirus (Aphelocheirus) Westwood 1833
Invalid names: Atoposita Jordan 1967 [synonym], Coreoidus Jordan 1967 [synonym]
Invalid names: Atopositidae Jordan 1967 [synonym]
Fm. †Leptaphelocheiridae Polhemus 2000
+
Fm. Naucoridae Fallen 1814 [creeping water bug]
+
G. †Aphlebocoris Handlirsch 1906
+
†Aphlebocoris nana Handlirsch 1906
†Aphlebocoris punctata Handlirsch 1921
G. †Cratopelocoris López Ruf and Goodwyn 2005
+
†Cratopelocoris carpinteroi López Ruf and Goodwyn 2005
Subfm. Cryphocricinae Montandon 1897
+
Invalid names: Cataractocorini Popov 1971 [empty], Cryphocricini Montandon 1897 [empty]
Subfm. Naucorinae Fallen 1814
+
Tr. †Aidiini Popov 1971
Tr. †Angaronectini Popov 1971
Tr. †Liadonaucorini Popov 1971
Tr. Naucorini Fallen 1814
Invalid names: Ambrysinae Usinger 1941 [empty], Aphelocheirinae Fieber 1851 [empty], Aphlebocoridae Handlirsch 1906 [synonym], Apopnidae Handlirsch 1921 [synonym], Cheirochelinae Montandon 1897 [empty], Laccocorinae Stal 1876 [empty], Limnocorinae Stal 1876 [empty], Naucoris dilatatus Heer 1853 [nomen dubium], Saucrolidae Santos 1971 [synonym]
Fm. Potamocoridae Usinger 1941
+
Invalid names: Coleopterocorini Popov 1971 [empty], Potamocorini Usinger 1941 [empty]
Fm. †Triassocoridae Tillyard 1922
+
G. †Triassocoris Tillyard 1922
+
†Triassocoris grandis Tillyard 1923
†Triassocoris myersi Tillyard 1922
†Triassocoris ovalis Tillyard 1923
†Triassocoris scutulum Tillyard 1922
Superfm. Nepoidea Latreille 1802
+
Fm. Belostomatidae Leach 1815 [giant water bug]
+
G. †Belostomates Schöberlin 1888
Subfm. Belostomatinae Leach 1815
+
G. †Araripebelostomum Nel and Paicheler 1992
Tr. Belostomatini Leach 1815
Tr. Diplonychini Ribeiro et al. 2018
G. †Lethonectes Popov et al. 1994
G. †Mesonepa Handlirsch 1906
G. †Nettelstedtia Popov et al. 2000
G. †Odrowazicoris Popov 1996
G. †Propoissonia Nel and Paicheler 1992
G. †Sinobelostoma Zhao and Hong 1989
G. †Tarsabedus Popov et al. 1994
Subfm. Lethocerinae Lauck and Menke 1961
+
G. †Scarabaeides Germar 1839
+
†Scarabaeides deperditus Germar 1839
+
Invalid names: Belostomum hartingi Weyenbergh 1869 [synonym], Stanislawia ewaae Lakshiminarayana 1984 [synonym]
Invalid names: Mesobelostomum Haase 1890 [objective synonym]
Subfm. †Stygeonepinae Popov 1971
+
G. †Iberonepa Martínez-Delclòs et al. 1995
G. †Stygeonepa Popov 1971
Invalid names: Belostoma goldfussi Germar 1837 [nomen dubium], Belostomates speciosum Heer 1864 [nomen dubium], Diplonychus rotundatus Heer 1853 [nomen dubium], Paranoikidae Zamboni et al. 2002 [synonym]
Fm. Nepidae Latreille 1802
+
Invalid names: Coroides Hong 1984 [nomen dubium], Coroides xiejiaheense Hong 1984 [nomen dubium]
Invalid names: Laccotrephes incertus Popov 1971 [nomen dubium]
Superfm. Notonectoidea Latreille 1802
+
Fm. Notonectidae Latreille 1802
+
G. Notonecta Linnaeus 1758
+
Subg. Notonecta (Paranecta) Hutchinson 1929
†Notonecta cantalensis Nel and Paicheler 1992
†Notonecta mazzoniae Petrulevičius et al. 2010
†Notonecta primaeva Von Heyden 1859
†Notonecta vetula Zhang et al. 2012
†Notonecta xyphiale Popov 1964
+
Invalid names: Clemmys hydatidosa Hong 1982 [synonym], Clypostemma limna Lin 1980 [synonym], Clypostemma pelila Zhang 1985 [synonym], Clypostemma petila Zhang 1985 [synonym]
Invalid names: Clematina Popov 1964 [synonym], Clemmys Hong 1982 [synonym], Clyemmys Hong 1982 [synonym], Clypostemma Popov 1964 [synonym]
Subfm. Notonectinae Latreille 1802
+
G. †Asionecta Popov 1962
Tr. Notonectini Latreille 1802
G. †Pelonecta Popov 1971
Invalid names: Clypostemmatinae Popov 1964 [synonym]
Invalid names: Notonecta arvernica Piton 1942 [nomen dubium], Notonecta binuda Lauck 1960 [nomen dubium], Notonecta emersoni Scudder 1890 [nomen dubium], Notonecta sieblosensis Martini 1971 [nomen dubium]
Superfm. Ochteroidea Kirkaldy 1906
+
Fm. Ochteridae Kirkaldy 1906 [velvety shore bug]
+
G. †Parvochterus Zhang et al. 2022
+
†Parvochterus lanceolarus Zhang et al. 2022
†Parvochterus reticulatus Zhang et al. 2022
G. †Pristinochterus Yao et al. 2007
+
†Pristinochterus ovatus Yao et al. 2011
†Pristinochterus zhangi Yao et al. 2007
G. †Yuripachys Kment et al. 2020
+
†Yuripachys dubius Popov 1986
Invalid names: Meropachys Popov 1986 [replaced]
Fm. †Propreocoridae Popov et al. 1994
+
Fm. †Pseudonerthridae Martins-Neto and Goodwyn 2005
+
Superfm. Pleoidea Fieber 1851
+
Fm. Helotrephidae Esaki and China 1927
+
Invalid names: Helotrephinae Esaki and China 1927 [empty], Idiocorinae Esaki and China 1927 [empty], Neotrephinae China 1940 [empty]
Invalid names: Pleidae Fieber 1851 [empty]
Fm. †Pterocimicidae Popov et al. 1994
+
Infraor. Pentatomomorpha Leston et al. 1954
+
Superfm. Aradoidea Brullé 1836
+
Fm. Aradidae Brullé 1836 [flat bug]
+
Subfm. †Archearadinae Heiss and Grimaldi 2002
+
G. †Archearadus Heiss and Grimaldi 2001
G. †Archemezira Heiss and Chen 2023
G. †Cretopiesma Grimaldi and Engel 2008
+
Invalid names: Microaradus Heiss and Poinar 2012 [synonym]
G. †Cretozemira Heiss 2023
Subfm. Calisiinae Stal 1873
+
G. †Allocalisius Heiss 2002
G. †Archecalisius Heiss 2019
G. Calisiopsis Champion 1898
G. Calisius Stal 1860
G. †Calisiomorpha Heiss 2016
+
†Calisiomorpha herczeki Heiss 2023
†Calisiomorpha yuripopovi Heiss 2016
Subfm. Carventinae Usinger and Matsuda 1959
+
G. Acaricoris Harris and Drake 1948
G. Carventus Stal 1865
G. Neoproxius Usinger and Matsuda 1959
G. †Proneoproxius Heiss and Poinar 2012
Subfm. Mezirinae Oshanin 1908
+
G. †Brevisensoria Poinar 2011
G. Mezira Amyot and Audinet-Serville 1843
G. †Myanmezira Heiss and Poinar 2012
G. Neuroctenus Fieber 1860
Fm. Termitaphididae Meyers 1924 [termite bug]
+
G. Termitaradus Meyers 1924
+
†Termitaradus avitinquilinus Grimaldi and Engel 2008
†Termitaradus dominicanus Poinar 2011
†Termitaradus mitnicki Engel 2009
†Termitaradus protera Poinar and Doyen 1992
Superfm. Coreoidea Reuter 1910
+
Fm. Alydidae Amyot and Audinet-Serville 1843 [broad-headed bug]
+
G. Alydus Fabricius 1803
G. Daclera Signoret 1863
Subfm. Micrelytrinae Stal 1867
+
Tr. Micrelytrini Stal 1867
Tr. Noliphini Ahmad 1965
G. †Willershausenia Popov 2007
Invalid names: Alydus pulchellus Heer 1853 [nomen dubium], Daclera naturalis Theobald 1937 [nomen dubium], Leptocorisinae Stal 1886 [empty]
Fm. Coreidae Leach 1815 [leaf-footed bug]
+
Subfm. Coreinae Leach 1815
+
G. †Achrestocoris Scudder 1890
Tr. Anisoscelini Laporte 1833
Tr. Cloresmini Stål 1873
Tr. Coreini Leach 1815
Tr. Dasynini Bergroth 1913
G. †Ferriantenna Cumming and Le Tirant 2021
Tr. Gonocerini Mulsant and Rey 1870
Tr. Homoeocerini Amyot and Audinet-Serville 1843
G. †Magnusantenna Du and Chen 2021
G. †Phthinocoris Scudder 1890
G. †Piezocoris Scudder 1890
G. †Pseudodasynus Zhang 1989
Tr. Spartocerini Amyot and Audinet-Serville 1843
G. †Coreiopsis Hong 1984
+
†Coreiopsis miocenica Hong and Wang 1987
†Coreiopsis parva Hong and Wang 1987
†Coreiopsis punctata Hong 1984
G. †Coreites Heer 1853
+
†Coreites crassus Heer 1853
†Coreites dubius Piton 1935
†Coreites oblongus Heer 1853
†Coreites redemtus Heer 1853
Fm. Rhopalidae Amyot and Audinet-Serville 1843 [scentless plant bug]
+
Invalid names: Corizidae Costa 1853 [synonym]
Fm. †Trisegmentatidae Zhang et al. 1994
+
Fm. †Yuripopovinidae Azar et al. 2011
+
G. †Caulisoculus Zhang and Chen 2020
+
†Caulisoculus electrus Zhang and Chen 2020
†Caulisoculus minutus Shang et al. 2020
†Caulisoculus monlyae Zhuo and Chen 2022
G. †Megaoptocoris Zhou et al. 2022
+
†Megaoptocoris punctatus Zhou et al. 2022
†Megaoptocoris similis Zhou et al. 2022
G. †Miropictopallium Fabrikant and Novoselska 2024
+
†Miropictopallium coloradmonens Fabrikant and Novoselska 2024
G. †Pseudocaulisoculus Kóbor and Roca-Cusachs 2021
+
†Pseudocaulisoculus longicornis Kóbor and Roca-Cusachs 2021
Invalid names: Dehiscensicoridae Du et al. 2016 [synonym]
Superfm. Lygaeoidea Schilling 1829
+
Fm. Berytidae Fieber 1851 [stilt bug]
+
Fm. Blissidae Stal 1862
+
Fm. Geocoridae Dahlbom 1851
+
Subfm. Geocorinae Dahlbom 1851
+
G. †Eocenogeocoris Kóbor 2023
G. Geocoris Fallén 1814
G. †Protogeocoris Kóbor et al. 2023
Fm. Heterogastridae Stal 1872
+
G. Heterogaster Schilling 1829
+
†Heterogaster antiquus Heer 1853
†Heterogaster breviscutatus Theobald 1937
†Heterogaster famosus Förster 1891
†Heterogaster pumilio Heer 1853
†Heterogaster radobojanus Heer 1853
†Heterogaster redivivus Heer 1853
†Heterogaster sundgoviensis Theobald 1937
†Heterogaster troglodytes Heer 1853
Invalid names: Heterogaster tristis Heer 1864 [nomen nudum]
Fm. Lygaeidae Schilling 1829 [milkweed bug]
+
Subfm. Cyminae Stal 1862
+
G. †Cephalocoris Heer 1853
G. †Cymocoris Popov 1986
G. †Procymus Usinger 1940
G. †Lithochromus Scudder 1890
+
†Lithochromus extraneus Scudder 1890
†Lithochromus gardneri Scudder 1890
†Lithochromus mortuarius Scudder 1890
†Lithochromus obstrictus Scudder 1890
Subfm. Lygaeinae Schilling 1829
+
Tr. †Gurnardobayini Popov 2019
G. Lygaeosoma Spinola 1837
G. Lygaeus Fabricius 1794
Invalid names: Astacopinae Kirkaldy 1899 [synonym]
G. †Lygaeites Heer 1853
+
†Lygaeites acutus Heer 1853
†Lygaeites amabilis Cockerell 1921
†Lygaeites lividus Heer 1853
†Lygaeites obsoletus Heer 1853
†Lygaeites ovalis Heer 1853
†Lygaeites pusillus Heer 1853
G. †Necrochromus Scudder 1890
+
†Necrochromus cockerelli Scudder 1890
†Necrochromus labatus Scudder 1890
†Necrochromus saxificus Scudder 1890
G. †Procrophius Scudder 1890
+
†Procrophius communis Scudder 1890
†Procrophius costalis Scudder 1890
†Procrophius languens Scudder 1890
Invalid names: Nysius stratus Scudder 1890 [nomen dubium], Nysius terrae Scudder 1890 [nomen dubium], Nysius tritus Scudder 1890 [nomen dubium], Nysius vecula Scudder 1890 [nomen dubium], Nysius vinctus Scudder 1890 [nomen dubium]
Fm. Rhyparochromidae Amyot and Audinet-Serville 1843 [seed bug]
+
G. †Linnaea Scudder 1890
+
†Linnaea abolita Scudder 1890
†Linnaea carcerata Scudder 1890
†Linnaea evoluta Scudder 1890
†Linnaea gravida Scudder 1890
†Linnaea holmesii Scudder 1890
†Linnaea putnami Scudder 1890
Subfm. Rhyparochrominae Amyot and Audinet-Serville 1843
+
Tr. Drymini Stål 1872
Tr. Gonianotini Stål 1872
Tr. Lethaeini Stål 1872
Tr. Myodochini Still 1874
Tr. Rhyparochromini Amyot and Audinet-Serville 1843
Invalid names: Aphaninae Puton 1887 [synonym]
Invalid names: Pyrrhocoris rottensis Kment 2021 [nomen dubium]
Fm. †Pachymeridiidae Handlirsch 1906
+
G. †Aphanocoris Popov 1990
+
†Aphanocoris femoratus Popov 1990
†Aphanocoris parallelus Popov 1990
†Aphanocoris porrectus Popov 1990
†Aphanocoris turgensis Popov 1990
G. †Positocoris Popov 1990
+
†Positocoris angustus Popov 1990
†Positocoris intermedius Popov 1990
†Positocoris longirostris Popov 1990
†Positocoris minimus Popov 1990
†Positocoris sibiricus Popov 1990
G. †Takshania Popov 1990
+
†Takshania implicata Popov 1990
†Takshania modesta Popov 1990
†Takshania transversalis Popov 1990
Invalid names: Hypocimicidae Handlirsch 1921 [synonym], Pachymerus zucholdi Giebel 1856 [nomen dubium], Psychrocoridae Handlirsch 1920 [synonym], Sisyrocoridae Handlirsch 1921 [synonym]
Superfm. Pentatomoidea Leach 1815
+
Fm. Acanthosomatidae Signoret 1863 [shield bug]
+
Fm. Cydnidae Billberg 1820 [burrowing bug]
+
Subfm. Amnestinae Hart 1919
+
G. †Acanthamnestus Du et al. 2022
G. Amnestus Dallas 1851
G. †Brevipronotum Azar and Huang 2023
G. †Chilamnestocoris Lis et al. 2018
G. †Cilicydnus Yao et al. 2007
G. †Laevigemma Du et al. 2022
G. †Latiscutella Pinto and de Ornellas 1974
G. †Orienicydnus Yao et al. 2007
G. †Pricecoris Pinto and de Ornellas 1974
G. †Pullneyocoris Lis and Roca-Cusachs 2020
G. †Punctacorona Wang et al. 2019
G. †Quinalveus Du et al. 2022
Invalid names: Latiscutellidae Pinto and de Ornellas 1974 [synonym], Pricecoridae Pinto and de Ornellas 1974 [synonym]
G. †Cydnopsis Heer 1853
+
†Cydnopsis coleopteroides Heer 1853
†Cydnopsis dangeardi Piton 1936
†Cydnopsis deleta Heer 1853
†Cydnopsis exilis Heer 1853
†Cydnopsis haidingeri Heer 1853
†Cydnopsis handlirschi Cockerell 1909
†Cydnopsis meunieri Kinzelbach 1970
†Cydnopsis nana Kinzelbach 1970
†Cydnopsis pygmaea Heer 1853
†Cydnopsis scutellaris Heer 1853
G. †Necrocydnus Scudder 1890
+
†Necrocydnus amyzonus Scudder 1890
†Necrocydnus gosiutensis Scudder 1890
†Necrocydnus revectus Scudder 1890
†Necrocydnus senior Scudder 1890
†Necrocydnus solidatus Scudder 1890
†Necrocydnus stygius Scudder 1890
†Necrocydnus torpens Scudder 1890
†Necrocydnus vulcanius Scudder 1890
G. †Procydnus Scudder 1890
+
†Procydnus devictus Scudder 1890
†Procydnus divexus Scudder 1890
†Procydnus eatoni Scudder 1890
†Procydnus mamillanus Scudder 1878
†Procydnus pronus Scudder 1890
†Procydnus quietus Scudder 1890
†Procydnus reliquus Scudder 1890
†Procydnus vesperus Scudder 1890
G. Prolobodes Amyot and Audinet-Serville 1843
+
Invalid names: Discostoma Scudder 1890 [objective synonym], Lobostoma Amyot and Audinet-Serville 1843 [replaced]
G. †Thlibomenus Scudder 1890
+
†Thlibomenus limosus Scudder 1890
†Thlibomenus macer Scudder 1890
†Thlibomenus parvus Scudder 1890
†Thlibomenus perennatus Scudder 1890
†Thlibomenus petreus Scudder 1890
Fm. Cyrtocoridae Distant 1880
+
Fm. Dinidoridae Stål 1867 [dinidorid bug]
+
Invalid names: Dinidorinae Stal 1867 [empty], Megymeninae [empty]
Fm. †Kobdocoridae Popov 1986
+
G. †Extralapis Ryzhkova 2023
+
†Extralapis breviscutum Ryzhkova 2023
†Extralapis carens Ryzhkova 2023
Invalid names: Venicoridae Yao et al. 2012 [synonym]
Fm. Megarididae McAtee and Malloch 1928
+
Fm. †Mesopentacoridae Popov 1968
+
G. †Mesopentacoris Popov 1968
+
†Mesopentacoris costalis Popov 1968
†Mesopentacoris orientalis Popov 1989
Fm. Pentatomidae Leach 1815 [stink bug]
+
G. †Eospinosus Wedmann et al. 2021
+
†Eospinosus greenriverensis Wedmann et al. 2021
†Eospinosus peterkulkai Wedmann et al. 2021
Subfm. Pentatominae Leach 1815
+
Tr. Aeliini Douglas and Scott 1865
G. †Cacoschistus Scudder 1890
Tr. Carpocorini Mulsant and Rey 1866
Tr. Eysarcorini Mulsant and Rey 1866
Tr. Halyini Amyot and Audinet-Serville 1843
G. †Mataeoschistus Scudder 1890
G. †Necanicarbula Zhang 1989
Tr. Pentatomini Leach 1815
G. †Pentatomites Scudder 1890
G. †Polioschistus Scudder 1890
G. †Poteschistus Scudder 1890
Tr. Sciocorini Amyot and Audinet-Serville 1843
Tr. Strachiini Mulsant and Rey 1866
G. †Teleocoris Kirkaldy 1910
G. †Thlimmoschistus Scudder 1890
G. †Thnetoschistus Scudder 1890
G. †Tiroschistus Scudder 1890
G. †Pentatomoides Jordan 1967
+
†Pentatomoides atratus Jordan 1967
+
Invalid names: Apodiphus robustus Jordan 1967 [synonym], Pentatomoides echinodes Jordan 1967 [synonym]
G. †Teleoschistus Scudder 1890
+
†Teleoschistus antiquus Scudder 1878
†Teleoschistus multinervosus Henriksen 1922
†Teleoschistus placatus Scudder 1890
†Teleoschistus rigoratus Scudder 1890 [stink bug]
Fm. Phloeidae Amyot and Audinet-Serville 1843
+
G. Phloea Lepeletier de Saint-Fargeau and Audinet-Serville 1825
+
Invalid names: Phloeocoris Burmeister 1835 [objective synonym]
Fm. †Primipentatomidae Yao et al. 2013
+
G. †Primipentatoma Yao et al. 2013
+
†Primipentatoma fangi Yao et al. 2013
†Primipentatoma peregrina Yao et al. 2013
Fm. †Probascanionidae Handlirsch 1921
+
Fm. †Protocoridae Handlirsch 1906
+
G. †Protocoris Heer 1852
+
†Protocoris indistinctus Popov et al. 1994
†Protocoris insignis Heer 1864
†Protocoris planus Heer 1852
Fm. Scutelleridae Leach 1815 [metallic shield bug]
+
Subfm. Pachycorinae Amyot and Audinet-Serville 1843
+
G. Pachycoris Burmeister 1835
G. Tetyra Fabricius 1803
G. †Teleocydnus Henriksen 1922
+
†Teleocydnus auriculatus Piton 1940
†Teleocydnus transitorius Henriksen 1922
Subfm. Tessaratomidae Stal 1865 [giant shield bug]
+
Superfm. Piesmatoidea Amyot and Audinet-Serville 1843
+
Fm. Piesmatidae Amyot and Audinet-Serville 1843 [ash grey leaf bug]
+
Subfm. Piesmatinae Amyot and Audinet-Serville 1843
+
G. Piesma Lepeletier de Saint-Fargeau and Audinet-Serville 1828
Superfm. Pyrrhocoroidea Amyot and Audinet-Serville 1843
+
Fm. Pyrrhocoridae Amyot and Audinet-Serville 1843
+
G. Dysdercus Amyot and Audinet-Serville 1843 [cotton stainer bug]
+
†Dysdercus cinctus Scudder 1890
†Dysdercus unicolor Scudder 1890
G. Pyrrhocoris Fallen 1814 [firebug]
G. †Rhepocoris Scudder 1890
+
†Rhepocoris abscissus Scudder 1890
†Rhepocoris caducus Scudder 1890
†Rhepocoris collisus Scudder 1890
†Rhepocoris defectus Scudder 1890
†Rhepocoris exanimatus Scudder 1890
†Rhepocoris inhibitus Scudder 1890
†Rhepocoris macrescens Scudder 1890
†Rhepocoris minimus Scudder 1890
†Rhepocoris praetectus Scudder 1890
†Rhepocoris praevalens Scudder 1890
†Rhepocoris propinquans Scudder 1890
Invalid names: Parodarmistus Scudder 1890 [synonym]
Invalid names: Largidae Amyot and Audinet-Serville 1843 [empty]
Invalid names: Corixa pullus Von Heyden 1859 [nomen dubium], Cryptocerata [empty], Eocimicidae Handlirsch 1906 [empty], Galgulidae Westwood 1840 [empty], Nepa atavina Heer 1853 [nomen dubium], Xishanidae Hong 1981 [empty]
Diagnosis
No diagnoses are available