PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Bibionidae (march fly)

Insecta - Diptera - Bibionidae

Taxonomy

Bibionidae was named by Fleming (1821). It is extant. Its type is Bibio.

It was assigned to Diptera by Cockerell (1921), Meunier (1923), Pongrácz (1931) and Hong (1984); to Nematocera by Handlirsch (1906), Zeuner (1941), Bode (1953) and Carpenter (1992); to Bibionomorpha by Nel (1994); and to Bibionoidea by Hong and Wang (1985), Yeates et al. (2007), Skartveit (2009) and Nel et al. (2019).

Subtaxa

Bibio elongatus, Bibio incrassatus, Bibio linearis, Bibio robustus, Bibioninae, Burmahesperininae, Cretobibioninae, Dilophus magnus, Hesperininae (syn. Penthetriinae), Pleciinae (syn. Cascopleciinae), Protomyia adusta, Protomyia formicoides, Protomyia fusca

Synonyms

Bibio robustus was named by Oustalet (1870). It is a compression fossil. Its type locality is Corent (MNHN collection), which is in a Rupelian lacustrine - large marl in France.

It was considered a nomen dubium by Skartveit and Nel (2017).
Protomyia fusca was named by Oustalet (1870). It is a compression fossil. Its type locality is Corent (MNHN collection), which is in a Rupelian lacustrine - large marl in France.

It was recombined as Penthetria fusca by Evenhuis (1994); it was considered a nomen dubium by Skartveit and Nel (2017).
Protomyia adusta was named by Oustalet (1870). It is a compression fossil.

It was recombined as Penthetria adusta by Evenhuis (1994); it was considered a nomen dubium by Skartveit and Nel (2017).
Protomyia formicoides was named by Oustalet (1870). It is a compression fossil. Its type locality is Corent (MNHN collection), which is in a Rupelian lacustrine - large marl in France.

It was recombined as Penthetria formicoides by Evenhuis (1994); it was considered a nomen dubium by Skartveit and Nel (2017).
Bibio elongatus was named by Heer (1849). Its type specimen is SMNK Oe. 154 Inv. 1189, an exoskeleton, and it is a compression fossil. Its type locality is Oeningen (Carlsruhe collection), which is in a Sarmatian crater lake limestone in the Upper Freshwater-Molasse Formation of Germany.

It was considered a nomen dubium by Skartveit and Pika (2014).
Bibio linearis was named by Heer (1849). It is a compression fossil. Its type locality is Radoboj (K.K. Montanistischen Collection), which is in a Sarmatian lagoonal/restricted shallow subtidal limestone in Croatia.

It was considered a nomen dubium by Skartveit and Krizmanić (2020).
Bibio incrassatus was named by Heer (1849). It is a compression fossil. Its type locality is Radoboj (K.K. Montanistischen Collection), which is in a Sarmatian lagoonal/restricted shallow subtidal limestone in Croatia.

It was synonymized subjectively with Bibio morio by Evenhuis (1994); it was considered a nomen dubium by Skartveit and Krizmanić (2020).
Dilophus magnus was named by Dürrenfeldt (1968). Its type specimen is GZG 14836, an exoskeleton, and it is a compression fossil. Its type locality is Willershausen clay pit (GZG collection), which is in a Piacenzian pond marl in Germany.

It was considered a nomen dubium by Fitzgerald and Greenwalt (2022).

Synonymy list

Year	Name and author
1821	Bibionidae Fleming p. 55
1849	Bibio elongatus Heer p. 214 figs. Pl 16, fig 2
1849	Bibio linearis Heer p. 214 figs. Pl 16, fig 3
1849	Bibio incrassatus Heer p. 221 figs. Pl 16, fig 12
1864	Bibio elongatus Heer p. 394 fig. 311
1870	Bibio robustus Oustalet p. 128 figs. Pl 4, figs 7-9
1870	Protomyia adusta Oustalet p. 145 figs. Pl 5, fig 18
1870	Protomyia fusca Oustalet p. 145 figs. Pl 4, fig 15
1870	Protomyia formicoides Oustalet p. 150 figs. Pl 4, fig 18; pl 5, fig 19
1906	Bibionidae Handlirsch p. 488
1921	Bibionidae Cockerell
1923	Bibionidae Meunier p. 86
1931	Bibionidae Pongrácz p. 117
1931	Bibio elongatus Pongrácz p. 121 figs. Pl 7, 8; text-fig 8
1941	Bibionidae Zeuner
1953	Bibionidae Bode
1968	Dilophus magnus Dürrenfeldt p. 50 figs. Pl 1, fig 6
1984	Bibionidae Hong p. 3
1985	Bibionidae Hong and Wang p. 17
1992	Bibionidae Carpenter
1994	Penthetria adusta Evenhuis
1994	Penthetria formicoides Evenhuis
1994	Penthetria fusca Evenhuis
1994	Bibionidae Nel p. 451
2007	Bibionidae Yeates et al.
2009	Bibionidae Skartveit p. 10
2019	Bibionidae Nel et al. p. 530

Is something missing? Join the Paleobiology Database and enter the data

Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825

Rank	Name	Author
class	Insecta
order	Diptera
—	Neodiptera	Michelsen 1994
infraorder	Bibionomorpha
superfamily	Bibionoidea
family	Bibionidae
family	Bibionidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Bibionidae Fleming 1821 [march fly]

show all | hide all

Subfm. Bibioninae Fleming 1821

hide

Tr. Bibionini Fleming 1821

Subtr. Bibiodina Frey 1927

G. Bibiodes Coquillett 1904

†Bibiodes balticus Skartveit 2009

†Bibiodes confluens Cockerell 1915

†Bibiodes intermedia James 1937

†Bibiodes kishenehnensis Fitzgerald 2019

†Bibiodes nanus Skartveit 2009

†Bibiodes provincialis Skartveit and Nel 2017

Invalid names: Bibiodites Cockerell 1915 [synonym]

Subtr. Bibionina Fleming 1821

G. Bibio Geoffroy 1762

†Bibio acaptus Dürrenfeldt 1968

†Bibio aerosus Statz 1943

Invalid names: Bibio spadicea Statz 1943 [synonym]

†Bibio albangulus Dürrenfeldt 1968

†Bibio anasiformis Dürrenfeldt 1968

†Bibio anatolicus Skartveit and Nel 2012

†Bibio angustatus Heer 1849

†Bibio antiquus von Heyden 1856

†Bibio aquaesextiae Skartveit and Nel 2017

†Bibio atavus Cockerell 1909

†Bibio auripes Statz 1943

†Bibio aymardi Maneval 1936

Invalid names: Dichaneurum infossum Aymard 1856 [replaced], Dichaneurum primaevum Aymard 1856 [replaced]

†Bibio baumbergeri Theobald 1937

Invalid names: Bibio parenti Theobald 1937 [synonym]

†Bibio boulayi Riou 1999

†Bibio brachypodus Zhang 1989

†Bibio brevis Heer 1849

Invalid names: Bibio oblongus Heer 1849 [synonym]

Bibio brunettii Hardy 1973

Invalid names: Bibio abdominalis Brunetti 1912 [replaced]

†Bibio brunneifemus Zhang 1989

†Bibio brunneus Zhang 1989

†Bibio bullatus Hong 1985

†Bibio capnodes Melander 1949

†Bibio carnificus Dürrenfeldt 1968

†Bibio castaneipennis Skartveit and Wedmann 2021

†Bibio celasensis Theobald 1937

Invalid names: Bibio conformans Theobald 1937 [synonym], Bibio obtusa Theobald 1937 [synonym], Bibio tenuiapicalis Theobald 1937 [synonym]

†Bibio cockerelli James 1937

†Bibio colossoceps Zhang 1989

†Bibio compta Statz 1943

†Bibio constringtus Dürrenfeldt 1968

†Bibio contererus Dürrenfeldt 1968

†Bibio crassinervis Skartveit and Pika 2014

†Bibio cratopodus Zhang 1989

†Bibio cruciformis Dürrenfeldt 1968

†Bibio curtisi Heer 1856

†Bibio curtisii Heer 1856

Invalid names: Bibio romani Theobald 1937 [synonym], Bibio serresi Theobald 1937 [synonym], Curtisimyia eximia Giebel 1856 [objective synonym]

†Bibio deflectus Dürrenfeldt 1968

†Bibio deformatus Zhang 1989

†Bibio discerptus Dürrenfeldt 1968

†Bibio dislocatus Theobald 1937

Invalid names: Bibio consolida Theobald 1937 [synonym], Plecia macroventris Theobald 1937 [synonym]

†Bibio dormitus Dürrenfeldt 1968

†Bibio edda Skartveit et al. 2017

†Bibio elegantulus Novák 1878

†Bibio enormus Dürrenfeldt 1968

†Bibio enterodelus Unger 1841

Invalid names: Bibio lividus Heer 1849 [synonym]

†Bibio excurvatus Melander 1949

†Bibio eximius Zhang et al. 1994

†Bibio expansus Zhang 1989

†Bibio explanatus Melander 1949

†Bibio extremus Cockerell 1921

†Bibio firmus Heer 1849

Bibio flavissimus Brunetti 1925

†Bibio formosus Novák 1878

†Bibio fusiformis Heer 1849

†Bibio germari Meunier 1920

Invalid names: Bibio comosella Statz 1943 [synonym], Bibio heydeni Meunier 1920 [synonym]

†Bibio giganteus Unger 1841

Invalid names: Bibio maculatus Heer 1849 [synonym], Bibio partschii Heer 1849 [synonym], Plecia heroica von Heyden and von Heyden 1865 [synonym]

†Bibio gurnetensis Cockerell 1917

Invalid names: Bibio oblitus Cockerell 1921 [synonym]

†Bibio jamesi Melander 1949

†Bibio janus von Heyden 1870

†Bibio joursacensis Théobald 1935

†Bibio kochii Staub 1883

†Bibio latahensis Lewis 1969

†Bibio latiantennatus Dürrenfeldt 1968

†Bibio lignarius Germar 1837

†Bibio longineurus Zhang 1989

†Bibio major Oustalet 1870

Invalid names: Bibio inflexus Theobald 1937 [synonym]

†Bibio mancus Dürrenfeldt 1968

†Bibio marginatus Oustalet 1870

†Bibio medioalbus Dürrenfeldt 1968

†Bibio mimas von Heyden 1870

Invalid names: Bibio infumatus Meunier 1915 [synonym]

†Bibio miocenicus Hong and Wang 1985

†Bibio moestus Heer 1849

†Bibio nebulosus Dürrenfeldt 1968

†Bibio nigricosta Skartveit and Nel 2012

†Bibio obliteratus Cockerell 1915

†Bibio oligocenus Cockerell 1917

†Bibio pannosus von Heyden and von Heyden 1865

†Bibio penisularis Zhang 1993

†Bibio peraffinis Zhang 1989

†Bibio podager Melander 1949

†Bibio praecidus Dürrenfeldt 1968

†Bibio pulchellus Heer 1849

†Bibio roehli Statz 1943

†Bibio rottensis Statz 1943

†Bibio ruptineurus Zhang 1989

†Bibio sadoensis Fujiyama 1985

†Bibio sereri Massalongo 1856

†Bibio shanwangensis Hong 1984

†Bibio similis Dürrenfeldt 1968

†Bibio soldatus Dürrenfeldt 1968

†Bibio spectabilis Theobald 1937

†Bibio sticheli Handlirsch 1908

†Bibio styriacus Beier 1952

†Bibio succineus Skartveit 2021

Bibio tenebrosus Coquillett 1898

†Bibio tenuis Dürrenfeldt 1968

†Bibio testeus Lewis 1969

†Bibio tranopterus Zhang 1989

†Bibio tripus Dürrenfeldt 1968

†Bibio udus Zhang et al. 1994

†Bibio varineurus Zhang 1989

†Bibio ventricosus Zhang 1989

†Bibio vetus James 1937

†Bibio vixdus Dürrenfeldt 1968

†Bibio vulcanius Melander 1949

†Bibio vulpinus Statz 1943

†Bibio wickhami Cockerell 1915

†Bibio xylophilus Germar 1837

Invalid names: Bibio disjectus Theobald 1937 [nomen dubium], Bibio martinsi Heer 1861 [nomen nudum], Bibio obsoletus Heer 1849 [nomen dubium], Bibio tertiarius von Heyden 1862 [nomen dubium], Curtisimyia Giebel 1856 [synonym], Lithobibio Beier 1952 [synonym], Lithosomyia Carpenter 1986 [synonym], Mesomyia Pongrácz 1928 [synonym], Protoberis Cockerell 1915 [synonym]

Tr. Dilophini Pinto and Amorim 2000

G. Dilophus Meigen 1803

†Dilophus andrewrossi Nel et al. 2019

†Dilophus campbelli Harris 1983

†Dilophus cerestensis Nel et al. 2022

†Dilophus crassicornis Skartveit 2009

†Dilophus idanos Fitzgerald and Greenwalt 2022

†Dilophus krantzii von Heyden 1870

†Dilophus luteipennis Theobald 1937

†Dilophus matilei Waller et al. 2000

†Dilophus palaeofebrilis Skartveit 2009

†Dilophus pinguis Heer 1849

Invalid names: Bibio pinguis oeningensis Heer 1849 [synonym]

†Dilophus priscus Loew 1850

†Dilophus pumilio Skartveit and Pika 2014

†Dilophus succineus Skartveit 2009

Invalid names: Bibio pinguis radobojanus Heer 1849 [objective synonym]

G. †Protodilophus Skartveit 2023

†Protodilophus semispinosus Skartveit 2023

Invalid names: Dilophus deletus Von Heyden 1859 [nomen dubium]

Subfm. †Burmahesperininae Ševčík et al. 2021

hide

G. †Burmahesperinus Ševčík et al. 2021

†Burmahesperinus antennatus Ševčík et al. 2021

†Burmahesperinus conicus Ševčík et al. 2021

†Burmahesperinus pedicellatus Ševčík et al. 2021

Subfm. †Cretobibioninae Skartveit and Ansorge 2020

hide

G. †Cretobibio Skartveit and Ansorge 2020

†Cretobibio burmiticus Skartveit 2023

†Cretobibio montsecensis Skartveit and Ansorge 2020

†Cretobibio rasnitsyni Lukashevich et al. 2021

Subfm. Hesperininae Walker 1848

hide

G. Hesperinus Walker 1848

†Hesperinus electrus Skartveit 2009

†Hesperinus heeri von Heyden and von Heyden 1865

Invalid names: Penthetria rottensis Statz 1943 [synonym], Penthetria scita Statz 1943 [synonym], Plecia elegantula Meunier 1915 [synonym]

†Hesperinus hyalopterus Skartveit 2009

†Hesperinus macroculatus Skartveit 2009

G. Penthetria Meigen 1803

†Penthetria abacula Lewis 1971

†Penthetria aestimata Statz 1943

†Penthetria alderensis Lewis 1971

†Penthetria amoena Heer 1849

Invalid names: Protomyia speciosa Heer 1864 [synonym]

†Penthetria anthracina Heer 1849

†Penthetria beskonakensis Skartveit and Nel 2012

†Penthetria blanchardi Oustalet 1870

†Penthetria bohemica Novák 1878

†Penthetria brevicollis Heer 1849

†Penthetria carbonum von Heyden and von Heyden 1865

†Penthetria cimicoides Heer 1849

†Penthetria claripennis Theobald 1937

Invalid names: Plecia difficilis Theobald 1937 [synonym], Plecia graciosa Theobald 1937 [synonym], Plecia varia Theobald 1937 [synonym], Plecia variegata Theobald 1937 [synonym]

†Penthetria creedensis James 1938

†Penthetria egerana Novák 1878

†Penthetria elongatipennis Theobald 1937

Invalid names: Bibio maceratus Theobald 1937 [synonym]

†Penthetria fryi Rice 1959

†Penthetria fuchsi Handlirsch 1907

Invalid names: Penthetria marinosi Bachmayer et al. 1971 [objective synonym]

†Penthetria gigantea Theobald 1937

Invalid names: Plecia crassiventris Theobald 1937 [synonym], Plecia luteipennis Theobald 1937 [synonym], Plecia miegi Theobald 1937 [synonym]

†Penthetria globularis Oustalet 1870

†Penthetria gracilis Unger 1841

†Penthetria graciliventris Theobald 1937

†Penthetria immutabilis Melander 1949

†Penthetria imperialis Novák 1878

†Penthetria incerta Oustalet 1870

†Penthetria integroneura Skartveit 2009

†Penthetria intermedia Scudder 1892

†Penthetria longa Heer 1849

†Penthetria longifurca Melander 1949

†Penthetria longiventris Theobald 1937

Invalid names: Plecia superba Theobald 1937 [synonym]

†Penthetria luberonica Skartveit and Nel 2017

†Penthetria lugens Oustalet 1870

†Penthetria montanaregis Skartveit 2009

†Penthetria murchisonii Unger 1841

†Penthetria nervisinuata Theobald 1937

†Penthetria nigra Statz 1943

†Penthetria parschlugiana Heer 1849

†Penthetria rubescens Oustalet 1870

†Penthetria rubiensis Lewis 1971

†Penthetria subterranea Theobald 1937

†Penthetria togoensis Fujiyama and Iwao 1974

†Penthetria tropica Statz 1943

Penthetria velutina Loew 1858

†Penthetria whipsawensis Rice 1959

Invalid names: Bibio alacris Oustalet 1870 [nomen dubium], Bibio gigas Oustalet 1870 [nomen dubium], Bibio macer Oustalet 1870 [nomen dubium], Bibio minor Oustalet 1870 [nomen dubium], Bibiopsis Heer 1849 [synonym], Mycetophaetus Scudder 1892 [synonym], Protomyia Heer 1849 [synonym]

Invalid names: Longicornia Hong 2002 [nomen nudum], Penthetriinae Rohdendorf 1946 [synonym]

Subfm. Pleciinae Duda 1930

hide

G. †Cretpenthetria Li et al. 2021

†Cretpenthetria burmensis Li et al. 2021

G. †Lichnoplecia Ren 1995

†Lichnoplecia kovalevi Ren 1995

G. Plecia Wiedemann 1823 [love bug]

†Plecia acourti Cockerell 1921

†Plecia aculeolata Zhang 1989

†Plecia akerionana Fitzgerald 1999

†Plecia amagua Cockerell 1925

†Plecia angustipennis Handlirsch 1910

†Plecia angustiventris Theobald 1937

†Plecia assonensis Gentilini 1993

†Plecia avus Handlirsch 1910

†Plecia axeliana Cockerell 1915

†Plecia baglii Gentilini 1991

†Plecia bivalvula Zhang 1993

†Plecia borussica Meunier 1904

†Plecia brunneipennis Piton 1939

†Plecia brunniptera Skartveit 2009

†Plecia bucklandi Heer 1849

Invalid names: Penthetria edwardsi Oustalet 1870 [synonym], Plecia dilatativentris Theobald 1937 [synonym], Protomyia elegans Heer 1856 [synonym], Protomyia gracilis Heer 1856 [synonym]

†Plecia cairnesi Rice 1959

†Plecia canadensis Handlirsch 1910

†Plecia capitata Zhang et al. 1994

†Plecia castellaroi Gentilini 1991

†Plecia chapuisii Oustalet 1872

†Plecia clavifemur Skartveit 2009

†Plecia collossea von Heyden and von Heyden 1865

Invalid names: Plecia imperialis Statz 1943 [synonym], Plecia philippi Statz 1943 [synonym], Plecia superba Meunier 1914 [synonym]

†Plecia conica Theobald 1937

Invalid names: Plecia haennii Gentilini 1993 [synonym]

†Plecia curtula Handlirsch 1910

Invalid names: Penthetria avunculus Handlirsch 1910 [synonym]

†Plecia decapitata Cockerell 1917

†Plecia dejecta Scudder 1890

†Plecia diatoma Zhang 1989

†Plecia dilatata Handlirsch 1910

†Plecia dubia Germar 1837

Invalid names: Helophilus primarius Germar 1837 [synonym], Plecia satyrus Statz 1943 [synonym], Protomyia abava von Heyden and von Heyden 1865 [synonym]

†Plecia dumasi Theobald 1937

†Plecia elatior Handlirsch 1910

†Plecia exigua Statz 1943

†Plecia explanata Cockerell 1917

†Plecia exposititia von Heyden 1865

†Plecia foersteri Theobald 1937

Invalid names: Plecia oviformis Theobald 1937 [synonym], Plecia splendida Theobald 1937 [synonym]

†Plecia fumosa Zhang et al. 1994

†Plecia gracilenta von Heyden and von Heyden 1865

†Plecia gracilentula Evenhuis 1994

Invalid names: Plecia gracilenta Zhang 1993 [replaced]

†Plecia gracillima Förster 1891

†Plecia gradata Melander 1949

†Plecia grossa von Heyden and von Heyden 1865

†Plecia hilaris Heer 1849

†Plecia hoffeinsorum Skartveit 2009

Invalid names: Plecia tenuicornis Skartveit 2009 [synonym]

†Plecia hypogaea von Heyden and von Heyden 1865

Invalid names: Plecia pulchella Meunier 1915 [synonym], Protomyia antenata von Heyden and von Heyden 1865 [synonym], Protomyia elongata von Heyden and von Heyden 1865 [synonym], Protomyia expositia von Heyden and von Heyden 1865 [synonym], Protomyia grandaeva von Heyden and von Heyden 1865 [synonym]

†Plecia inflata Oustalet 1870

†Plecia inflata Hull 1960

†Plecia intima Fujiyama and Iwao 1974

†Plecia jucunda Heer 1849

Invalid names: Protomyia affinis Heer 1849 [synonym]

†Plecia kanetakii Fujiyama 1970

†Plecia kelownaensis Rice 1959

†Plecia kudiella Cockerell 1925

†Plecia kuznetzovi Cockerell 1925

†Plecia larguensis Gentilini 1993

†Plecia livida Heer 1849

Invalid names: Plecia brevithoracis Theobald 1937 [synonym], Plecia cerestensis Theobald 1937 [synonym], Plecia constricta Theobald 1937 [synonym], Plecia flichei Theobald 1937 [synonym], Plecia fusciventris Theobald 1937 [synonym], Plecia goreti Theobald 1937 [synonym], Plecia maculitarsis Theobald 1937 [synonym], Plecia maculiventris Theobald 1937 [synonym], Protomyia brevipennis Heer 1856 [synonym]

†Plecia luctuosa von Heyden and von Heyden 1865

†Plecia ludongensis Hong and Wang 1985

†Plecia lugubris Heer 1849

†Plecia luteola von Heyden and von Heyden 1865

†Plecia lygaeoides Heer 1849

Invalid names: Bibio detractus Theobald 1937 [synonym], Plecia acutiventris Theobald 1937 [synonym], Plecia guinieri Theobald 1937 [synonym], Protomyia latipennis Heer 1849 [synonym]

†Plecia macilenta Skartveit and Pika 2014

†Plecia macrocephala von Heyden and von Heyden 1865

Invalid names: Protomyia schineri von Heyden and von Heyden 1865 [synonym]

†Plecia maimensis Gentilini 1993

†Plecia melanderi Cockerell 1911

†Plecia minutula Rice 1959

†Plecia morio Heer 1849

†Plecia myersi Peterson 1975

†Plecia nana Handlirsch 1910

†Plecia obsitula Cockerell 1925

†Plecia orycta Melander 1949

†Plecia oustaleti Brongniart 1876

†Plecia parisiensis Gee et al. 2001

†Plecia pealei Scudder 1890

†Plecia pennata Statz 1943

†Plecia pictipennis Handlirsch 1910

Invalid names: Penthetria lambei Handlirsch 1910 [synonym], Penthetria ovalis Handlirsch 1910 [synonym], Penthetria separanda Handlirsch 1910 [synonym]

†Plecia pinguis von Heyden and von Heyden 1865

Invalid names: Bibio brachypteroides Meunier 1915 [synonym], Plecia vulpina Statz 1943 [synonym], Protomyia lapidaria von Heyden and von Heyden 1865 [synonym]

†Plecia pisaurensis Gentilini 1991

†Plecia platoptera Zhang 1993

†Plecia platyptera Handlirsch 1910

†Plecia prisca Meunier 1899

†Plecia pristina Hardy 1971

†Plecia proserpina von Heyden and von Heyden 1865

†Plecia pulchra Handlirsch 1910

†Plecia pulla Handlirsch 1910

Invalid names: Penthetria brevipes Handlirsch 1910 [synonym]

†Plecia quaesita Novák 1878

†Plecia rectivenia Zhang 1989

†Plecia redempta Cockerell 1925

†Plecia reducta Handlirsch 1910

†Plecia refracta Cockerell 1925

†Plecia rhenana von Heyden and von Heyden 1865

†Plecia rhodopterina Cockerell 1925

†Plecia similkameena Scudder 1879

†Plecia sluiteri Meunier 1917

†Plecia solaris Zhang 1989

†Plecia spinula Zhang 1989

†Plecia sturmi Statz 1943

†Plecia stygia von Heyden and von Heyden 1865

Invalid names: Protomyia veterana von Heyden and von Heyden 1865 [synonym]

†Plecia tessella Melander 1949

†Plecia theobaldi Skartveit and Nel 2017

Invalid names: Bibio nigripennis Theobald 1937 [replaced]

†Plecia transitoria Handlirsch 1910

Invalid names: Penthetria falcatula Handlirsch 1910 [synonym], Penthetria fragmentum Handlirsch 1910 [synonym]

†Plecia tulameenensis Rice 1959

†Plecia undans Zeuner 1941

†Plecia vergnei Piton 1935

†Plecia villosa Zhang 1989

†Plecia volgeri Von Heyden 1859

†Plecia vulcania Zhang 1989

†Plecia winchesteri Cockerell 1917

†Plecia winnertzi von Heyden and von Heyden 1865

†Plecia woodruffi Cockerell 1916

Invalid names: Bibio cylindratus Oustalet 1870 [nomen dubium], Clothonopsis Hong and Wang 1987 [nomen dubium], Clothonopsis miocenica Hong and Wang 1987 [nomen dubium], Epiplecia Giard 1879 [synonym], Plecia joannis Oustalet 1870 [nomen dubium], Plecia larteti Oustalet 1870 [nomen dubium], Plecia longipennis Oustalet 1870 [nomen dubium], Plecia painvini Meunier 1915 [nomen dubium], Plecia pallida Oustalet 1870 [nomen dubium], Plecia retracta Theobald 1937 [nomen dubium], Plecia rubescens Oustalet 1870 [nomen dubium], Plecia sauvagei Oustalet 1870 [nomen dubium]

G. †Protopenthetria Li et al. 2021

†Protopenthetria skartveiti Li et al. 2021

Invalid names: Cascopleciinae Poinar 2010 [synonym], Fushunoplecia Hong 2002 [nomen nudum]

Invalid names: Bibio elongatus Heer 1849 [nomen dubium], Bibio incrassatus Heer 1849 [nomen dubium], Bibio linearis Heer 1849 [nomen dubium], Bibio robustus Oustalet 1870 [nomen dubium], Dilophus magnus Dürrenfeldt 1968 [nomen dubium], Protomyia adusta Oustalet 1870 [nomen dubium], Protomyia formicoides Oustalet 1870 [nomen dubium], Protomyia fusca Oustalet 1870 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Late/Upper Barremian to the top of the Late/Upper Pleistocene or 130.00000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 125.45 Ma

Collections (192 total)

Time interval	Ma	Country or state	Original ID and collection number
Early/Lower Barremian	130.0 - 125.45	Spain	Cretobibio montsecensis (118913)
Late/Upper Barremian	130.0 - 125.45	China (Hebei)	Lichnoplecia kovalevi (107001)
Late/Upper Aptian	122.46 - 113.0	Brazil (Ceará)	Bibionidae indet. (122352) Cretobibio rasnitsyni (217013)
Albian	113.0 - 100.5	Jordan	Bibionidae indet. (124847)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar	Burmahesperinus sp., Burmahesperinus antennatus, Burmahesperinus conicus, Burmahesperinus pedicellatus (187929) Cretobibio burmiticus, Protodilophus semispinosus (133102) Protopenthetria skartveiti, Cretpenthetria burmensis (161726)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar (Kachin)	Cascoplecia insolitis (104053)
Turonian	93.9 - 89.8	Botswana	Bibionidae indet., Bibiodes sp. (123430)
Late/Upper Campanian	83.5 - 70.6	Canada (Manitoba)	Plecia myersi (122354)
Selandian	61.6 - 59.2	France (Auvergne)	Bibio sp. (113685 123463)
Thanetian	59.2 - 56.0	United Kingdom (Scotland)	Plecia undans (113632)
Ypresian	56.0 - 47.8	USA (Colorado)	Plecia winchesteri (105004)
Ypresian	56.0 - 47.8	France (Oise)	Plecia parisiensis (123850) Plecia sp. (68174)
Ypresian	56.0 - 47.8	Italy	Bibio sereri (152649)
Ypresian	56.0 - 47.8	Canada (British Columbia)	Penthetria angustipennis, Penthetria reducta, Penthetria fragmentum, Penthetria curtula, Penthetria dilatata, Penthetria platyptera, Plecia transitoria (119857) Penthetria fryi, Plecia intermedia, Plecia curtula, Plecia reducta, Plecia canadensis, Plecia avus (120981) Penthetria fryi, Plecia kelownaensis, Plecia avus, Plecia transitoria, Plecia pictipennis (120934) Penthetria fryi, Plecia minutula, Plecia curtula, Plecia tulameenensis, Plecia reducta, Plecia similkameena, Plecia canadensis, Plecia transitoria, Plecia pictipennis (120943) Penthetria fryi, Plecia tulameenensis, Plecia reducta, Plecia avus, Plecia pictipennis (120940) Penthetria pictipennis (119896) Penthetria pictipennis, Penthetria pulchra, Plecia curtula (119863) Penthetria pulla, Penthetria brevipes, Penthetria pictipennis, Penthetria elatior, Penthetria falcatula, Penthetria nana, Penthetria separanda, Penthetria avunculus, Penthetria avus, Penthetria lambei, Penthetria ovalis, Penthetria transitoria, Penthetria canadensis, Plecia reducta (119864) Penthetria similkameena, Plecia canadensis, Plecia transitoria (112781) Penthetria whipsawensis (120624) Plecia angustipennis, Plecia canadensis (120988) Plecia avus (121037) Plecia cairnesi (120992) Plecia cairnesi, Plecia reducta, Plecia avus, Plecia pictipennis (120991) Plecia curtula (120989 120990) Plecia dilatata (121041) Plecia kelownaensis (119464) Plecia pulchra (121046) Plecia reducta, Plecia pictipennis (121009) Plecia transitoria (121040) Plecia tulameenensis, Plecia similkameena, Plecia canadensis, Plecia avus, Plecia transitoria (119895) Pleciinae indet. (31677)
Ypresian	56.0 - 47.8	China (Liaoning)	Longicornia tenuis, Fushunoplecia eocenica (124820)
Wasatchian	55.8 - 50.3	USA (Wyoming)	Plecia pealei (122174 122192 122193 187759)
Waipawan - Bortonian	55.8 - 39.1	New Zealand	Dilophus campbelli (119708)
Bridgerian	50.3 - 46.2	USA (Wyoming)	Plecia dejecta (122090) Plecia sp. (21851)
Bridgerian	50.3 - 46.2	USA (Utah)	Plecia woodruffi (124691)
Bridgerian	50.3 - 46.2	USA (Colorado)	Plecia rhodopterina (122746) Plecia winchesteri (124625)
Lutetian	47.8 - 41.3	USA (Montana)	Bibiodes kishenehnensis (117614) Dilophus idanos (135666)
Uintan	46.2 - 39.7	USA (Colorado)	Plecia akerionana (138790)
Priabonian	38.0 - 33.9	Russian Federation (Kaliningrad)	Bibio sp. (124201) Dilophus sp., Penthetria montanaregis, Plecia clavifemur, Plecia hoffeinsorum, Dilophus crassicornis, Bibio succineus (217198) Plecia borussica, Plecia prisca (123927) Plecia sp. 1, Plecia sp. 2, Plecia sp. 3, Dilophus sp., Penthetria montanaregis, Plecia brunniptera, Plecia clavifemur, Plecia prisca, Bibiodes nanus, Dilophus crassicornis, Dilophus succineus (124517)
Priabonian	38.0 - 33.9	Poland	Dilophus priscus (109625)
Priabonian	38.0 - 33.9	France (Gard)	Bibio celasensis, Bibio obtusa (122405) Dilophus sp., Bibio celasensis, Penthetria longiventris, Penthetria graciliventris (144529)
Priabonian	38.0 - 33.9	United Kingdom (England)	Bibio gurnetensis, Bibio oligocenus, Bibiodites confluens, Protoberis obliteratus, Bibio oblitus, Bibio extremus, Dilophus andrewrossi (123959) Penthetria sp., Bibio oligocenus, Bibiodes confluens, Dilophus andrewrossi (123960) Plecia acourti (21976)
Priabonian	38.0 - 33.9	Russian Federation	Penthetria sp., Hesperinus electrus, Hesperinus hyalopterus, Hesperinus macroculatus, Penthetria integroneura, Plecia brunniptera, Plecia clavifemur, Plecia hoffeinsorum, Plecia prisca, Plecia tenuicornis, Bibiodes balticus, Dilophus crassicornis, Dilophus palaeofebrilis (123419)
Priabonian	38.0 - 33.9	France	Bibionidae indet., Plecia longiventris, Plecia superba, Plecia angustiventris, Plecia foersteri, Plecia longa, Plecia splendida, Plecia graciliventris, Bibio celasensis, Bibio tenuiapacalis, Bibio elongatipennis, Bibio macerata, Plecia varia (113709)
Priabonian	38.0 - 33.9	Ukraine	Plecia hoffeinsorum (217199)
Chadronian	37.2 - 33.9	USA (Colorado)	Bibio cockerelli, Bibio jamesi, Bibio podager (124681) Bibio cockerelli, Bibiodes intermedia (106382) Bibio wickhami (124680) Bibio wickhami, Bibio cockerelli, Bibio explanatus, Bibio vulcanius (122975) Bibio wickhami, Bibio cockerelli, Bibio vetus (106381) Bibio wickhami, Bibiodes intermedia, Plecia gradata (106379) Bibiodes intermedia (106380) Mycetophaetus intermedius (106384) Plecia axeliana, Bibio wickhami (113749) Plecia decapitata (122977) Plecia explanata, Bibio wickhami, Bibio cockerelli, Bibio vetus, Hesperinus axelianus, Hesperinus immutabilis, Hesperinus melanderi, Penthetria longifurca, Plecia orycta, Plecia tessella, Bibio capnodes, Bibio excurvatus, Bibio jamesi, Bibio vulcanius (124624) Plecia melanderi (144782)
Late/Upper Eocene	37.2 - 33.9	France	Bibio celasensis (184102)
Late/Upper Eocene	37.2 - 33.9	France (Gard)	Plecia longiventris, Plecia angustiventris, Plecia foersteri, Plecia splendida, Plecia dumasi, Bibio conformans (122406)
Early/Lower Oligocene	33.9 - 28.4	France (Auvergne Region)	Plecia splendida, Dichaneurum infossum, Dichaneurum primaevum (31767)
Early/Lower Oligocene	33.9 - 28.4	France	Plecia bucklandi, Plecia grossa, Plecia exposititia (121474) Plecia bucklandi, Plecia rhenana, Plecia luctuosa (122408) Plecia rhenana, Plecia grossa, Plecia lapidaria, Plecia gracillima, Plecia lygaeoides (121476) Plecia rubescens (121475) Plecia sp. 1, Plecia sp. 2, Plecia rhenana, Plecia pallida, Plecia stygia, Plecia grossa (121473)
MP 23	33.9 - 28.4	France	Bibionidae indet., Bibio sp., Plecia sp., Plecia bucklandi, Plecia splendida, Bibio painvini, Plecia livida, Plecia acutiventris, Bibio inflexus, Plecia flichei, Plecia fusciventris, Plecia guinieri, Plecia goreti, Plecia brevithoracis, Plecia cerestensis, Plecia maculitarsis, Plecia maculiventris, Bibio detractus (122535) Bibionidae indet., Plecia sp., Plecia lugens, Plecia nervisinuata, Plecia constricta, Plecia goreti, Plecia maculiventris (122536) Plecia livida, Penthetria nervisinuata, Bibio major, Dilophus cerestensis (157327)
Rupelian	33.9 - 28.1	France	Dilophus sp., Protomyia oustaleti (139949) Plecia conica (184088) Plecia livida (184089) Plecia lygaeoides, Plecia livida (184090) Plecia maimensis, Plecia assonensis, Plecia larguensis (156997)
Rupelian	33.9 - 28.1	France (Auvergne)	Bibio gigas, Protomyia adusta, Protomyia globularis (152488) Bibionidae indet., Bibio edwardsii, Plecia major, Bibio gigas, Protomyia joannis (152487) Bibionidae indet., Plecia major, Bibio gigas, Bibio robustus, Bibio lartetii, Protomyia incerta (152491) Bibionidae indet., Plecia pallida, Protomyia longa, Protomyia lugens, Protomyia inflata, Bibio edwardsii, Plecia major, Bibio marginatus, Bibio macer, Bibio alacris, Bibio robustus, Bibio cylindratus, Bibio minor, Bibio lartetii, Protomyia longipennis, Protomyia joannis, Protomyia fusca, Protomyia adusta, Protomyia sauvagei, Protomyia blanchardi, Protomyia rubescens, Protomyia formicoides, Protomyia incerta, Bibio obsoletus (127630)
Rupelian	33.9 - 28.1	USA (Montana)	Plecia sp., Plecia inflata (132747)
Rupelian	33.9 - 28.1	Germany	Bibionidae indet., Plecia sp., Bibiodes sp., Dilophus sp., Plecia lapidaria, Plecia foersteri, Plecia splendida, Bibio baumbergeri, Bibio parenti, Bibio dislocata, Bibio consolida, Plecia gigantea, Plecia crassiventris, Plecia lugens, Plecia inflata, Plecia claripennis, Plecia macroventris, Plecia oviformis, Plecia conica, Plecia miegi, Plecia luteipennis, Plecia subterranea, Plecia variegata, Plecia varia, Plecia difficilis, Plecia graciosa, Plecia rubescens (122407)
Rupelian	33.9 - 28.1	France (Alpes-de-Haute)	Dilophus sp., Hesperinus sp., Plecia livida, Plecia maimensis, Plecia assonensis, Plecia larguensis (164936) Plecia conica, Plecia maimensis, Plecia assonensis, Plecia larguensis (184087)
Oligocene	33.9 - 23.03	France	Plecia haennii (157308)
Whitneyan	33.3 - 30.8	USA (Montana)	Bibionidae indet., Bibio sp., Plecia sp., Hesperinus sp., Penthetria sp. (120623) Penthetria rubiensis, Penthetria abacula, Penthetria alderensis (120622)
Late/Upper Oligocene	28.4 - 23.03	USA (Colorado)	Plecia creedensis (117882 121977)
Late/Upper Oligocene	28.4 - 23.03	France (Bouches-du-Rhone)	Bibio celasensis (113989) Bibio nigripennis (122464)
Late/Upper Oligocene	28.4 - 23.03	Germany	Bibio tertiarius (152951)
MP 28	28.4 - 23.03	Germany (Rheinland-Pfalz)	Bibio xylophilus, Penthetria nigra, Plecia dubia, Bibio castaneipennis (71942)
MP 30	28.4 - 23.03	Germany (Nordrhein-Westfalen)	Bibiopsis volgeri, Bibio lignarius (127628) Plecia superba, Bibio infumatus, Bibio brachypteroides, Bibio janus, Plecia pulchella, Protomyia hypogaea, Protomyia antenata, Protomyia sluiteri, Protomyia veterana, Protomyia elongata, Bibio germari, Bibio heydeni, Protomyia expositia, Protomyia elegantula, Plecia carbonum (138696)
MP 30	28.4 - 23.03	Germany	Helophilus primarius, Bibio lignarius, Bibio xylophilus, Phthiria dubia (115802) Plecia rhenana, Protomyia luctuosa, Protomyia lapidaria, Bibio deletus, Protomyia hypogaea, Protomyia antenata, Bibio pannosus, Protomyia abava, Protomyia collossea, Protomyia winnertzi, Protomyia grossa, Protomyia proserpina, Protomyia macrocephala, Protomyia exposititia, Protomyia stygia, Protomyia pinguis, Protomyia veterana, Protomyia grandaeva, Protomyia luteola, Protomyia schineri, Protomyia elongata, Protomyia gracilenta, Protomyia heeri, Plecia heroica, Dilophus krantzii, Bibio mimas (123945) Protomyia luctuosa, Protomyia lapidaria, Bibio janus, Protomyia winnertzi, Protomyia macrocephala, Protomyia exposititia, Protomyia stygia, Protomyia grandaeva, Dilophus krantzii (154946)
Chattian	28.1 - 23.03	Germany (North Rhine-Westphalia)	Dilophus sp. 1, Penthetria sp., Penthetria sp 2, Plecia grossa, Plecia hypogaea, Plecia grandaeva, Plecia philippi, Plecia satyrus, Plecia imperialis, Plecia pennata, Plecia sturmi, Plecia exigua, Plecia vulpina, Penthetria nigra, Penthetria tropica, Penthetria aestimata, Penthetria rottensis, Penthetria scita, Bibio roehli, Bibio comosella, Bibio germari, Bibio rottensis, Bibio vulpina, Bibio aerosa, Bibio spadicea, Bibio auripes, Bibio compta, Plecia sp. 4, Bibio sp 1, Plecia sp. 7, Plecia sp. 12, Plecia sp. 1, Plecia sp. 2, Plecia rhenana, Plecia sp. 10, Plecia sp. 11, Plecia sp. 3, Plecia sp. 5, Plecia sp. 6, Plecia sp. 13, Plecia sp. 14, Penthetria sp 1, Bibio sp 4, Bibio sp 2, Bibio sp 6, Bibio sp 3, Bibio sp 5, Plecia sp. 8, Plecia sp. 15, Plecia lapidaria (122010)
Chattian	28.1 - 23.03	France (Bouches-Du-Rhone)	Bibio sp., Dilophus sp., Plecia bucklandi, Plecia splendida, Plecia lugens, Bibio spectabilis, Bibio romani, Bibio disjectus, Bibio serresi, Plecia dilatativentris, Plecia livida, Plecia gracilis, Plecia elegans, Bibio curtisi, Plecia guinieri, Plecia brevipennis, Bibio curtisii, Penthetria nervisinuata, Bibio aquaesextiae, Bibiodes provincialis (122501) Bibio sp., Plecia bucklandi, Plecia splendida, Bibio conformans, Plecia lugens, Bibio morio, Bibio painvini, Bibio spectabilis, Bibio romani, Bibio disjectus, Bibio fusiformis, Bibio detractus, Bibio serresi, Plecia dilatativentris, Plecia nervisinuata, Plecia livida, Plecia gracilis, Plecia elegans, Plecia constricta, Bibio curtisi, Plecia guinieri, Plecia brevipennis, Plecia lygaeoides (138838) Bibionidae indet. (153412) Plecia bucklandi, Plecia splendida, Bibio morio, Bibio detractus, Plecia livida, Plecia gracilis, Plecia elegans, Plecia retracta, Plecia constricta, Bibio curtisi, Plecia brevipennis (122509) Plecia livida, Plecia elegans (122511) Plecia rhenana, Plecia lapidaria, Bibio baumbergeri, Bibio painvini, Bibio spectabilis, Penthetria subterranea (113712) Protomyia bucklandi (122878) Protomyia gracilis, Protomyia elegans, Bibio curtisii, Plecia macilenta, Penthetria nervisinuata (122876) Protomyia lygaeoides, Protomyia bucklandi, Bibio morio, Protomyia gracilis, Protomyia brevipennis, Bibio fusiformis, Empis sp., Penthetria sp., Diptera indet., Bibio moestus (122877)
Chattian	28.1 - 23.03	France	Plecia livida, Penthetria luberonica (183981)
Chattian	28.1 - 23.03	France (Provence-Alpes-Côte d'Azur)	Bibionidae indet., Plecia splendida, Bibio conformans, Plecia lugens, Bibio painvini, Bibio spectabilis, Bibio luteipennis, Bibio romani, Bibio disjectus, Bibio detractus, Bibio serresi, Plecia dilatativentris, Plecia nervisinuata, Plecia livida, Plecia gracilis, Plecia elegans, Plecia retracta, Plecia acutiventris, Bibio curtisi, Plecia brevipennis, Bibio fusiformis, Plecia lygaeoides (122488) Plecia splendida, Bibio conformans, Bibio serresi, Plecia livida, Plecia gracilis, Plecia elegans, Plecia constricta, Bibio curtisi (122487)
Aquitanian	23.03 - 20.44	Germany	Penthetria sp. (190126) Plecia sp., Bibio sticheli (123872)
Aquitanian	23.03 - 20.44	Turkey	Bibio anatolicus, Bibio nigricosta, Penthetria beskonakensis (123307)
Early/Lower Miocene	23.03 - 15.97	Slovenia	Bibio sticheli (152681)
Early/Lower Miocene	23.03 - 15.97	Mexico (Chiapas)	Plecia pristina (140210)
Early/Lower Miocene	23.03 - 15.97	Germany	Bibio antiquus (152937)
Early/Lower Miocene	23.03 - 15.97	Japan	Bibio sadoensis (114003)
Early/Lower Miocene	23.03 - 15.97	Spain (Castellon)	Bibio sp. (225781)
Burdigalian	20.44 - 15.97	Russian Federation	Plecia amagua, Plecia kudiella, Plecia refracta, Plecia redempta, Plecia kuznetzovi, Plecia obsitula (113705)
Burdigalian	20.44 - 15.97	Greece	Penthetria fuchsi (144526) Penthetria marinosi (130761)
Burdigalian	20.44 - 15.97	Austria	Bibionidae indet. (139943)
Burdigalian	20.44 - 15.97	China (Shandong)	Bibio shanwangensis, Bibio flavissimus (130412) Bibio shanwangensis, Bibio miocenica, Plecia ludongensis, Plecia solaris, Bibio ventricosus, Bibio expansus, Bibio deformatus, Fungivora bullata, Bibio brunneus, Bibio cratopodus, Bibio ruptineurus, Clothonopsis miocenica (128550) Bibio shanwangensis, Plecia diatoma, Plecia vulcania, Plecia villosa, Plecia rectivenia, Bibio brunneifemus, Bibio peraffinis, Bibio brachypodus, Bibio tranopterus, Bibio cratopodus, Bibio varineurus, Bibio longineurus, Bibio eximius (128551) Plecia capitata (138944) Plecia fumosa, Plecia bivalvula, Plecia gracilenta (138946) Plecia platoptera, Bibio penisularis, Bibio udus (138945) Plecia spinula, Plecia aculeolata, Plecia villosa, Bibio ventricosus, Bibio expansus, Bibio colossoceps, Bibio abdominalis, Bibio tranopterus, Bibio varineurus (128552)
Burdigalian - Langhian	20.44 - 13.82	Dominican Republic	Dilophus matilei (156229) Plecia pristina (122474)
MN 4	16.9 - 15.97	Czech Republic	Bibiopsis egerana, Bibiopsis imperialis, Bibio formosus, Bibio elegantulus, Protomyia bohemica, Plecia quaesita (117612)
Langhian	15.97 - 13.82	Japan	Bibio sp., Plecia kanetakii (103101)
Langhian	15.97 - 13.82	USA (Idaho)	Bibio latahensis, Bibio testeus (97616)
Langhian	15.97 - 13.82	USA (Washington)	Bibio sp. (97619)
MN 5 - MN 6	15.97 - 12.75	Bosnia and Herzegovina	Bibio sp., Plecia sp. (211309)
Middle Miocene	15.97 - 11.608	Germany	Bibiopsis carbonum (152936)
Middle Miocene	15.97 - 11.608	Bulgaria	Plecia sp. (181730)
Middle Miocene - Late/Upper Miocene	15.97 - 5.333	Thailand	Plecia sp. (140983)
Badenian	13.65 - 12.7	Austria (Steiermark)	Lithobibio styriacus (139945) Protomyia parschlugiana (115306)
Sarmatian	12.7 - 11.608	Croatia	Bibio gracilis, Protomyia latipennis (156758) Bibio lividus, Protomyia anthracina (113693) Bibio moestus, Bibio maculatus, Mesomyia brevis (156759) Bibio sp., Penthetria longa, Penthetria obsoleta, Bibio partschii, Bibio maculatus, Bibiopsis murchisonii, Penthetria anthracina (117247) Plecia lygaeoides, Plecia lugubris, Bibio giganteus, Bibio enterodelus, Penthetria anthracina, Bibio crassinervis, Penthetria cimicoides, Dilophus pinguis (224110) Protomyia bucklandi, Protomyia longa, Bibio morio, Plecia lugubris, Bibio linearis, Bibio gracilis, Bibio pinguis, Bibio incrassatus, Bibio lividus, Bibiopsis murchisonii, Bibiopsis brevicollis, Protomyia anthracina (117761) Protomyia lygaeoides, Protomyia longa, Bibio giganteus, Bibio gracilis, Bibio maculatus, Bibio pinguis, Bibio enterodelus, Bibio firmus, Bibiopsis cimicoides, Protomyia latipennis, Bibio murchisonis (117246)
Sarmatian	12.7 - 11.608	Germany (Baden-Württemberg)	Bibio fusiformis, Bibio brevis, Bibio moestus, Bibio pulchellus, Protomyia jucunda, Protomyia affinis (113689) Bibio fusiformis, Bibio brevis, Bibio moestus, Plecia hilaris, Bibio elongatus, Bibio angustatus, Bibio pulchellus, Protomyia jucunda, Protomyia speciosa, Bibio crassinervis, Dilophus pumilio, Penthetria brevicollis, Plecia macilenta (117252) Bibio fusiformis, Bibio brevis, Bibio moestus, Plecia hilaris, Bibio oblongus (115304) Bibio fusiformis, Bibio brevis, Bibio obsoletus, Bibio moestus, Bibio elongatus, Bibio angustatus, Bibio oblongus, Protomyia jucunda, Protomyia amoena (115305) Bibio moestus (117338) Bibio oeningensis (117260)
Tortonian	11.62 - 7.246	Iceland	Bibio sp., Dilophus sp., Penthetria sp., Bibio edda (154940)
Tortonian	11.62 - 7.246	France	Bibio joursacencis (138766)
Late/Upper Miocene	11.608 - 5.333	Romania	Bibio kochii (155724)
Turolian	8.7 - 5.333	France (Ardèche)	Bibio boulayi (156164) Bibio joursacensis, Plecia vergnei (172761)
Messinian	7.246 - 5.333	Italy	Bibio sp., Plecia spp., Penthetria sp., Plecia castellaroi, Plecia pisaurensis, Plecia baglii (157306)
Messinian	7.246 - 5.333	USA (Alaska)	Bibio sp. (103102)
Messinian	7.246 - 5.333	Italy (Tuscany)	Bibio sp., Penthetria sp. (113751)
Piacenzian	3.6 - 2.588	Germany (Niedersachsen)	Bibio sp., Plecia sp., Dilophus magnus, Bibio enormus, Bibio latiantennatus, Bibio medioalbus, Bibio contererus, Bibio mancus, Bibio dormitus, Bibio soldatus, Bibio tripus, Bibio acaptus, Bibio carnificus, Bibio constringtus, Bibio praecidus, Bibio deflectus, Bibio similis, Bibio discerptus, Bibio nebulosus, Bibio vixdus, Bibio cruciformis, Bibio albangulus, Bibio anasiformis, Bibio tenuis (109196)
Piacenzian	3.6 - 2.588	Japan	Bibionidae indet., Bibio sp., Penthetria togoensis, Plecia intima (160140) Bibionidae indet., Plecia sp., Penthetria velutina, Bibio tenebrosus (188976)
Gelasian	2.588 - 1.806	France	Bibionidae indet., Plecia sp., Plecia brunneipennis (113996) Plecia lugubris, Plecia vergnei (113994) Plecia sp. (138769)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Washington)	Bibionidae indet. (167662)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom (Worcestershire)	Dilophus sp. (163070 163075)