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Phymatopleuridae
Taxonomy
Phymatopleuridae was named by Batten (1956). Its type is Phymatopleura.
It was assigned to Pleurotomariacea by Knight et al. (1960); to Pleurotomarioidea by Golikov and Starobogatov (1975), Bouchet et al. (2005), Bouchet et al. (2017), Nützel et al. (2018) and Karapunar and Nützel (2022); and to Eotomarioidea by Wagner (2023).
It was assigned to Pleurotomariacea by Knight et al. (1960); to Pleurotomarioidea by Golikov and Starobogatov (1975), Bouchet et al. (2005), Bouchet et al. (2017), Nützel et al. (2018) and Karapunar and Nützel (2022); and to Eotomarioidea by Wagner (2023).
Subtaxa
Altotomaria, Austroscalata, Borestus (syn. Platypleurotomaria), Callitomaria, Catazona, Discotomaria, Eirlysia, Euryalox (syn. Sagana), Eymarella, Fetaspira, Juvenispira, Lamellospira, Paragoniozona, Perakella, Phymatopleura (syn. Orestes) (type genus), Tapinotomaria, Termihabena, Trochotomaria, Vicnigoria, Worthenia
Synonymy list
| Year | Name and author |
|---|---|
| 1956 | Phymatopleuridae Batten |
| 1960 | Phymatopleuridae Knight et al. p. 214 |
| 1975 | Phymatopleuridae Golikov and Starobogatov p. 208 |
| 2005 | Phymatopleuridae Bouchet et al. p. 244 |
| 2017 | Phymatopleuridae Bouchet et al. p. 343 |
| 2018 | Phymatopleuridae Nützel et al. p. 480 |
| 2022 | Phymatopleuridae Karapunar and Nützel p. 36 |
| 2023 | Phymatopleuridae Wagner p. F346 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. †Phymatopleuridae Batten 1956
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G. †Austroscalata Waterhouse 1987
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†Austroscalata branxtonensis Waterhouse 1987
†Austroscalata marina Waterhouse 1987
G. †Borestus Thomas 1940
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†Borestus costatus Yoo 1988
†Borestus granihumerosus Wang 1980
†Borestus heterospira Gemmellaro 1889
†Borestus lunatus Pinilla et al. 2007
†Borestus magdalenensis Batten 1995
†Borestus pagoda Newell 1935
†Borestus planiapicata Wanner 1942
†Borestus procerus Thomas 1940
†Borestus rotundatus Batten 1972
†Borestus texanus Batten 1995
†Borestus wrighti Thomas 1940
Invalid names: Platypleurotomaria Wanner 1942 [synonym]
G. †Callitomaria Batten 1956
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†Callitomaria magna Batten 1958
†Callitomaria stanislavi Batten 1956
†Callitomaria tepuelensis Sabattini and Noirat 1969
G. †Discotomaria Batten 1956
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†Discotomaria basisulcata Batten 1956
†Discotomaria costata Batten 1958
†Discotomaria dubia Batten 1958
†Discotomaria nodosa Batten 1958
†Discotomaria xingwenensis Pan 1985
G. †Eirlysia Batten 1956
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†Eirlysia ceramicorum Peel 2016
†Eirlysia exquisita Batten 1956
†Eirlysia globosa Mazaev 2015
†Eirlysia lata Mazaev 2015
†Eirlysia lens Mazaev 2015
†Eirlysia modesta Waterhouse 1986
†Eirlysia nodata Mazaev 2015
†Eirlysia nodosa Batten 1958
†Eirlysia reticulata Batten 1958
†Eirlysia reticulatiformis Mazaev 2019
†Eirlysia robusta Branson 1930
†Eirlysia scalare Mazaev 2015
†Eirlysia seminuda Mazaev 2015
†Eirlysia spiralis Dietz 1911
†Eirlysia undata Mazaev 2015
G. †Euryalox
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†Euryalox anxianensis Pan 1982
†Euryalox applanatus Nützel et al. 2018
†Euryalox boeckhi Kittl 1900
†Euryalox juvavica Koken 1896
Invalid names: Sagana Koken 1897 [replaced]
G. †Eymarella
G. †Lamellospira Batten 1958
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†Lamellospira alveozona Kulas and Batten 1997
†Lamellospira anatola Batten 1972
†Lamellospira catherinae Gemmellaro 1889
†Lamellospira cincta Batten 1958
†Lamellospira conica Batten 1958
†Lamellospira spinosa Batten 1989
G. †Paragoniozona Nelson 1947
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†Paragoniozona granulostriata Meek and Worthen 1860
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Invalid names: Pleurotomaria aspera Girty 1934 [replaced]
†Paragoniozona millegranosa Girty 1934
†Paragoniozona multilirata Nelson 1947
†Paragoniozona nodolirata Nelson 1947
†Paragoniozona ornata Karapunar and Nützel 2022
†Paragoniozona paucinodosa Nelson 1947
†Paragoniozona venustiformis Licharew 1967
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Invalid names: Rhineoderma ferganica Licharew 1967 [synonym], Yunnania semigranulosa Licharew 1967 [synonym]
†Paragoniozona yanceyi Karapunar and Nützel 2022
†Paragoniozona yunnania Batten 1972
G. †Phymatopleura Girty 1939
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†Phymatopleura baccata Reed 1944
†Phymatopleura brazoensis Shumard 1860
†Phymatopleura conica Karapunar and Nützel 2022
†Phymatopleura girtyi Karapunar and Nützel 2022
†Phymatopleura heteropleura Elias 1958
†Phymatopleura kattaensis Waagen 1880
†Phymatopleura linkiana King 1850
†Phymatopleura missouriensis Bandel 2009
†Phymatopleura nodosa Girty 1912
†Phymatopleura paraliensis Reed 1944
†Phymatopleura preclara Hoare et al. 1997
†Phymatopleura thyrrena Gemmellaro 1889
†Phymatopleura uchhaliensis Reed 1944
Invalid names: Orestes Girty 1912 [replaced]
G. †Tapinotomaria Batten 1956
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†Tapinotomaria cancellata Mazaev 2019
†Tapinotomaria coronata Batten 1958
†Tapinotomaria costata Batten 1958
†Tapinotomaria crassa Batten 1958
†Tapinotomaria duplicostata Batten 1958
†Tapinotomaria globosa Batten 1958
†Tapinotomaria mirabilis Batten 1958
†Tapinotomaria psiche Gemmellaro 1889
†Tapinotomaria pyramidalis Batten 1958
†Tapinotomaria rugosa Batten 1956
†Tapinotomaria spinosa Batten 1966
†Tapinotomaria submirabilis Batten 1958
G. †Worthenia de Koninck 1883
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†Pleurotomaria (Worthenia) constantini Mansuy 1912
Subg. †Worthenia (Humiliworthenia) Yin and Yochelson 1983
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†Worthenia (Humiliworthenia) anisica Nützel et al. 2018
†Worthenia (Humiliworthenia) hausmanni Goldfuss 1841
†Worthenia (Humiliworthenia) nuda Koken 1900
Subg. †Worthenia (Worthenia) de Koninck 1883
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†Worthenia (Worthenia) conica Assmann 1937
†Worthenia (Worthenia) dobrogeana Nützel et al. 2018
†Worthenia (Worthenia) ecki Assmann 1924
†Worthenia (Worthenia) extensus Yin and Yochelson 1983
†Worthenia (Worthenia) xui Yin and Yochelson 1983
†Worthenia alticarinata Batten 1989
†Worthenia angulata Münster 1841
†Worthenia basifalcata Korner 1937
†Worthenia bialveozona Batten 1989
†Worthenia bicarinata Batten 1989
†Worthenia bieberi Kittl 1891
†Worthenia bilineata Klipstein 1843
†Worthenia cainallensis Kittl 1899
†Worthenia castlemanensis Lintz 1958
†Worthenia cirriformis Laube 1868
†Worthenia constantini Mansuy 1912
†Worthenia contabulata Costa 1964
†Worthenia corrugata Chronic 1952
†Worthenia crenilunula Yoo 1994
†Worthenia crenulata Batten 1989
†Worthenia curvistriata Koken and Perner 1925
†Worthenia dharmaensis Blaschke 1907
†Worthenia dieneri Blaschke 1907
†Worthenia distincta Kittl 1894
†Worthenia dregeri Kittl 1891
†Worthenia duplicata Kittl 1891
†Worthenia dyadica Kittl 1903
†Worthenia eremita Koken 1897
†Worthenia escheri Stoppani 1865
†Worthenia extendia Tong and Erwin 2001
†Worthenia furcata Kittl 1891
†Worthenia gepidorum Kittl 1900
†Worthenia gubleri Delpey 1941
†Worthenia hoettingensis Alma 1926
†Worthenia humiligrada Ketwetsuriya et al. 2020
†Worthenia humilis Böhm 1895
†Worthenia indifferens Kittl 1894
†Worthenia kingi Batten 1989
†Worthenia klamathensis Smith 1927
†Worthenia latialveozona Batten 1989
†Worthenia legrandi Kues and Batten 2001
†Worthenia liebeneri Laube 1868
†Worthenia loczyi Kittl 1900
†Worthenia margaritacea Laube 1868
†Worthenia morovi Mazaev 2020
†Worthenia multicarinata Mansuy 1912
†Worthenia nana Grabau 1931
†Worthenia pagoda Mansuy 1914
†Worthenia parvula Hoare et al. 1997
†Worthenia pilula Batten 1989
†Worthenia planalveozona Batten 1989
†Worthenia plutonis Kittl 1894
†Worthenia praerupta Böhm 1895
†Worthenia punctata Wanner 1922
†Worthenia rhombifera Korner 1937
†Worthenia schirjaevensis Stuckenberg 1905
†Worthenia sculpta Phillips 1836
†Worthenia sigaretoides Kittl 1894
†Worthenia speciosa Meek and Worthen 1860
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Invalid names: Worthenia perizomata White 1881 [synonym]
†Worthenia strigata Böhm 1895
†Worthenia strigillata Herrick 1888
†Worthenia subplicata Klipstein 1843
†Worthenia substriata Klipstein 1843
†Worthenia subtilis Kittl 1891
†Worthenia superstes Szabo 1980
†Worthenia tabulata Conrad 1835
†Worthenia tongxinensis Guo 1983
†Worthenia triton d'Orbigny 1849
†Worthenia vamosensis Kittl 1900
†Worthenia whitehorsensis Saunders and Yochelson 1964
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Invalid names: Worthenia beedei Knight 1940 [replaced]
†Worthenia windowblindensis Batten and Stokes 1986
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: o = order | |||||
| Reference: Bambach et al. 2007 | |||||
Age range: base of the Late/Upper Famennian to the top of the Bajocian or 364.70000 to 168.30000 Ma
Collections (523 total)
Oldest occurrences | |||
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Late/Upper Famennian | China | Phymatopleura sp. (162315) | |
| Hastarian | Australia (New South Wales) | Worthenia crenilunula, Borestus costatus (43901) | |
| Kinderhookian | Australia (New South Wales) | Worthenia sp. (75614) | |
| Kinderhookian | USA (Iowa) | Phymatopleura sp. (49824) | |
| Tournaisian | USA (Iowa) | Worthenia sp. (3309) | |
| Tournaisian | Australia (New South Wales) | Borestus costatus (43949 44159 44161) Paragoniozona sp. (44158) Worthenia sp., Borestus costatus (43950 44080 44088) Worthenia sp., Worthenia crenilunula (44079) Worthenia sp., Worthenia crenilunula, Borestus costatus (43904) | |
| Kinderhookian - Osagean | Mexico (Sonora) | Borestus sp. (5616) | |
| Carboniferous | USA (Oklahoma) | Phymatopleura sp. (157 158 159 172) Phymatopleura sp., Worthenia sp. (117 163) | |
| Carboniferous | USA (Missouri) | Phymatopleura sp. (115) | |
| Osagean | USA (Ohio) | Worthenia strigillata (93067 155894) | |
Youngest occurrences |
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| Time interval | Ma | Country or state | PBDB collection number |
| Sinemurian | Austria | Trochotomaria lobitzeri (173631 173632) | |
| Raricostatum - Davoei | Hungary | Worthenia superstes (78690) | |
| Pliensbachian | Hungary | Worthenia superstes (91406) | |
| Davoei | Hungary | Worthenia superstes (108954 119137) | |
| Ibex | Hungary | Worthenia superstes (108945 108949) | |
| Margaritatus | Spain (Alicante ) | Worthenia superstes (130881) | |
| Margaritatus | Hungary | Worthenia superstes (108960) | |
| Early/Lower Pliensbachian - Middle Toarcian | Hungary | Worthenia superstes (108969) | |
| Late/Upper Aalenian | Chile (Atacama) | Trochotomaria sp. (221797) | |
| Bajocian | Italy (Central Italy ) | Trochotomaria problematica (123682) | |