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Protozoa

Protozoa

Taxonomy
Protozoa was named by Goldfuss (1820) [also said to be 1817 or 1820]. It is extant.

It was reranked as the phylum Protozoa by von Siebold (1845) and Owen (1860).

It was assigned to Life by Owen (1860); to Animalia by von Siebold (1845), Cavalier-Smith (1993); to Eukaryota by Blackwell (2004); and to Eukaryota by Ruggiero et al. (2015).

Subtaxa
Biciliata, Entamoebidae, Eozoa, Perispira, Rhizopodea, Sarcodina, Sarcomastigota
Sister taxa
Rhizopodea
Entered
by C. Cervato on 2006-02-09; modified by M. Uhen on 2023-07-11

Synonymy list
YearName and author
1820Protozoa Goldfuss
1845Protozoa von Siebold
1860Protozoa Owen
1993Protozoa Cavalier-Smith
2004Protozoa Blackwell p. 274
2015Protozoa Ruggiero et al. p. 15

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RankNameAuthor
—Life
—Eukaryota()
RankNameAuthor
classProtozoaGoldfuss 1820
classProtozoaGoldfuss 1820

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Cl. Protozoa Goldfuss 1820
show all | hide all
Fm. Entamoebidae Chatton 1925
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G. †Entamoebites Poinar and Boucot 2006
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†Entamoebites antiquus Poinar and Boucot 2006
Unr. Eozoa
+
+
Or. Kinetoplastida Honigberg 1961
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Fm. Trypanosomatidae Doflein 1901
+
G. †Paleotrypanosoma Poinar 2008
Invalid names: Euglenoidea [empty], Symbiontida [empty]
+
+
Invalid names: Neolouka [empty]
G. Malawimonas O'Kelly and Nerad 1999
+
Invalid names: Pharyngomonada [empty], Tetramitia [empty]
Invalid names: Fornicata [empty], Heterolobosea [empty], Jakobida [empty], Metamonada [empty], Parabasalia [empty], Preaxostyla [empty]
G. Microcetella Aescht 2001
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Microcetella lappus Orpin and Mathiesen 1986
Invalid names: Microcetus Orpin and Mathiesen 1986 [objective synonym]
G. Perispira Stein 1859
Cl. Rhizopodea von Siebold 1845
+
+
†Belonechitina hirsuta Laufeld 1967
†Belonechitina robusta Eisenack 1959
G. †Euconochitina Taugourdeau 1966
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†Fungochitina pilosa Collinson and Scott 1958
Ph. †Granuloreticulosa de Saedeleer 1934
+
G. Lampas Montfort 1810
Invalid names: Athalamida [empty], Labyrinthulida [empty]
G. †Hoegisphaera Staplin 1961
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†Hoegisphaera glabra Staplin 1961
G. †Laufeldochitina Paris 1981
Subph. Sarcodina Schmarda 1871
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Cl. Reticularea Lankester 1885
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Subcl. Acantharia Haeckel 1862
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Superfm. Astrolonchacea Haeckel 1881
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Fm. Astrolonchidae Haeckel 1881
Superfm. Dorataspacea Haeckel 1862
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Fm. Dorataspidae Haeckel 1862
Subcl. Filosia Leidy 1879
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Or. Gromiida Claparède and Lachmann 1859
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Superfm. Gromiacea Reuss 1862
Invalid names: Aconchulinida [empty], Xenophyophorida [empty]
Subcl. Radiolaria Müller 1858 [radiolarian]
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G. Actinommura Haeckel 1887
G. †Adamas Afanasieva 2000
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†Adamas cathedrarius Afanasieva 2000
G. Amphicentria Ehrenberg 1862
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Amphicentria salpa Ehrenberg 1862
G. †Amphicraspedum Haeckel 1882
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†Amphicraspedum murrayanum Haeckel 1887
G. †Anthocyrtium Haeckel 1887
G. Archiphormis Haeckel 1882
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Archiphormis cancellata Ehrenberg 1854
Superfm. Archipiliacea Haeckel 1881
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Fm. Artophormidae Haeckel 1881
Fm. Aulosphaeridae Haeckel 1862
Fm. Sethophatnidae Haeckel 1881
Fm. Stichoperidae Haeckel 1881
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Invalid names: Podocampida Haeckel 1887 [synonym], Stichocyrtida Haeckel 1881 [synonym], Stichopiliidae Frizzell 1951 [synonym], Tetracyrtida Haeckel 1881 [synonym], Triacartidae Campbell 1954 [synonym]
G. †Artocyrtis Haeckel 1887
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G. †Astractinium Haeckel 1887
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Astractinium aristotelis Ehrenberg 1847
G. †Cancellientactinia Obut and Shcherbanenko 2008
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†Cancellientactinia acifera Obut and Shcherbanenko 2008
G. †Cavasphaera Won and Iams 2015
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†Cavasphaera diversa Won and Iams 2015
†Cavasphaera inaequalis Won and Iams 2015
G. Ceratastrum Haeckel 1882
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Ceratastrum trochus Ehrenberg 1873
Superfm. Challengeriacea Murray 1876
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Fm. Lirellidae Loeblich and Tappan 1961
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Invalid names: Cadiidae Borgert 1910 [replaced]
G. Chilomma Ehrenberg 1847
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Chilomma saturnus Ehrenberg 1862
G. Clathrocircus Haeckel 1882
G. Clathrocorona Haeckel 1882
Superfm. Coccodiscacea Haeckel 1862
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Fm. Ommatodiscidae Stöhr 1880
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Invalid names: Euchitoniidae Haeckel 1882 [synonym]
Fm. Phacodiscidae Haeckel 1881
Or. Collodaria Haeckel 1881
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Fm. Collosphaeridae Müller 1858
G. Siphonosphaera Müller 1858
Fm. Sphaerozoidae Haeckel 1862
Fm. Thalassosphaeridae Haeckel 1862
G. Cornutellium Haeckel 1882
G. Corythomelissa Campbell 1951
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Corythomelissa corythium Ehrenberg 1874
G. †Cromyatractus Haeckel 1887
G. Cromyechinus Haeckel 1882
G. †Crucidiscus Rüst 1892
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Invalid names: Crucidiscus praecursor Rüst 1892 [nomen dubium]
G. Cryptoprora Ehrenberg 1847
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Cryptoprora fundicola Ehrenberg 1873
G. †Cubosphaera Deflandre and Deflandre-Rigaud 1958
G. Cyrtidosphaera Haeckel 1861
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G. †Cyrtophormis Haeckel 1887
G. Dictyocodon Haeckel 1882
G. Dictyospyrella Haeckel 1887
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Dictyospyrella tristoma Ehrenberg 1874
G. Dictyospyrissa Haeckel 1887
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Dictyospyrissa fenestra Ehrenberg 1874
G. Dipospyris Haeckel 1887
G. †Dizonium Rüst 1892
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Invalid names: Dizonium laeve Rüst 1892 [nomen dubium]
G. Druppatractona Haeckel 1887
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Druppatractona laevis Ehrenberg 1874
G. Drymyomma Jørgensen 1900
G. †Dystympanium Rüst 1892
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Invalid names: Dystympanium carbonicum Rüst 1892 [nomen dubium]
Superfm. Ethmosphaeracea Haeckel 1862
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Fm. Actinommidae Haeckel 1862
Fm. Ethmosphaeridae Haeckel 1862
+
Invalid names: Cenosphaeridae Deflandre 1952 [synonym], Liosphaeridae Campbell 1954 [synonym]
Fm. Triposphaeridae Vinassa de Regny 1898
G. Euphysetta Cleve 1898
G. †Guttacapsa Squinabol 1903
G. †Gyrosphaera Noble and Maletz 2000
+
†Gyrosphaera cavea Jones and Noble 2006
†Gyrosphaera primigena Umeda and Suzuki 2005
†Gyrosphaera raneatela Noble and Maletz 2000
†Gyrosphaera siljanensis Noble and Maletz 2000
G. †Halicapsa Rüst
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Invalid names: Halicapsa megapora Rüst 1892 [nomen dubium]
G. †Haploplus Li 1995
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†Haploplus qunii Li 1995
G. †Haplopolaris Li 1991
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G. †Hapolpolus Li 1995
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G. †Hataina Huang 1967
G. †Heliaster Hollande and Enjumet 1960
G. Heliosestilla Haeckel 1887
G. †Hexalastrum Rüst
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Invalid names: Hexalastrum infans Rüst 1892 [nomen dubium]
G. Hexaspyridium Haeckel 1887
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Hexaspyridium setigera Ehrenberg 1874
G. †Hexastylidium Aberdeen
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†Hexastylidium variatum Aberdeen 1940
G. Hymenastromma Haeckel 1887
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Hymenastromma ternarium Ehrenberg 1874
G. †Labyrinthosphaera Noble and Maletz 2000
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†Labyrinthosphaera lancia Jones and Noble 2006
†Labyrinthosphaera lenzi Jones and Noble 2006
†Labyrinthosphaera macdonaldi Noble and Maletz 2000
G. Lamprodiscus Ehrenberg 1861
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Lamprodiscus monoceros Ehrenberg 1873
G. Lithamphora Popofsky 1908
G. Lithocampium Haeckel 1882
G. Lithocorythium Ehrenberg 1847
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Lithocorythium galea Ehrenberg 1844
G. Lithomitrella Haeckel 1887
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Lithomitrella acephalum Ehrenberg 1874
G. Lithopilium Popofsky 1913
G. Lithotympanium Haeckel 1882
G. †Lynchnocanella Clark and Campbell 1942
G. Mazosphaera Ehrenberg 1861
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Mazosphaera laevis Ehrenberg 1873
G. Microlecitos Issel 1890
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Microlecitos cassis Ehrenberg 1844
G. †Neoaspiculum Won and Iams 2011
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†Neoaspiculum amplum Won and Iams 2011
†Neoaspiculum cancellium Won and Iams 2011
†Neoaspiculum echinatum Won and Iams 2015
†Neoaspiculum headense Won and Iams 2011
†Neoaspiculum laxum Won and Iams 2011
†Neoaspiculum maletziana Won and Iams 2015
†Neoaspiculum transformum Won and Iams 2011
G. †Nyfrieslandia Maletz and Bruton 2007
+
†Nyfrieslandia complicata Maletz and Bruton 2007
†Nyfrieslandia crassispinosa Won and Iams 2013
†Nyfrieslandia sphaeroidea Maletz and Bruton 2007
G. Octalacorys Haeckel 1887
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Octalacorys aculeata Ehrenberg 1874
G. †Odontosphaera Haeckel 1887
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Invalid names: Odontosphaera echinocactus Rüst 1892 [nomen dubium]
G. †Palhindeolithus Deflandre 1973
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†Palhindeolithus ambiguus Deflandre 1973
†Palhindeolithus diductus Deflandre 1973
†Palhindeolithus octostyla Dumitrica 1982
†Palhindeolithus paululus Dumitrica 1982
†Palhindeolithus pulcher Deflandre 1973
†Palhindeolithus triassicus Dumitrica 1982
G. Panartus Haeckel 1887
G. †Parafollicucullinoides Xiao et al. 2021
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†Parafollicucullinoides globosus Xiao et al. 2021
G. †Pentalacorys Haeckel 1882
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†Pentalacorys pentacantha Ehrenberg 1874
G. †Pentinastrum Haeckel 1881
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†Pentinastrum subbotinae Lipman 1960
G. Peridium Haeckel 1887
G. †Peripanartus Rüst
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Invalid names: Peripanartus deficiens Rüst 1892 [nomen dubium]
G. Peritiviator Pessagno Jr. 1976
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†Peritiviator labyrinthi Pessagno 1976
Subor. †Pheodaria
+
Superfm. †Phaeocystina
Superfm. Plagoniacea Haeckel 1881
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Fm. Plectaniidae Haeckel 1881
G. †Plagoniscus Hinde
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Invalid names: Plagoniscus colligatus Hinde 1899 [nomen dubium], Plagoniscus cristatus Hinde 1899 [nomen dubium], Plagoniscus simplex Hinde 1899 [nomen dubium], Plagoniscus vetustus Hinde 1899 [nomen dubium]
Superor. Polycystina Ehrenberg 1838
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Superfm. Actinommacea Haeckel 1862
Or. †Albaillellaria Deflandre 1953
Or. †Archaeospicularia Dumitrica et al. 2000
Or. †Entactinaria Kozur and Mostler 1982
Or. †Latentifistularia Caridroit et al. 1999
G. †Protobiramus Won and Below 1999
G. †Praespongocoelia Noble
+
G. †Priscilatentidiota Li 1995
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G. †Prismatium Rüst
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Invalid names: Prismatium paradictyum Rüst 1892 [nomen dubium]
G. Protocystis Wallich 1862
G. †Protosegmentum Wang 2008
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G. †Prunulum Rüst
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Invalid names: Prunulum armeniacum Rüst 1892 [nomen dubium], Prunulum murrayi Rüst 1892 [nomen dubium]
G. †Pseudostaurosphaera Krasheninnikov 1960
G. Psilomelissa Haeckel 1882
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Psilomelissa galeatus Ehrenberg 1873
G. Pteractis Ehrenberg 1873
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Pteractis elegans Ehrenberg 1873
G. Pylosphaera Ehrenberg 1858
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Pylosphaera mediterranea Ehrenberg 1858
G. †Retientactinosphaera Afanasieva 2011
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†Retientactinosphaera clavata Afanasieva 2011
†Retientactinosphaera magnifica Afanasieva 2011
Superfm. †Rotasphaeracea Noble 1994
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Invalid names: Rotasphaeridae Noble 1994 [empty]
G. †Russirad Afanasieva 2000
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†Russirad kazintsovae Afanasieva 2000
G. †Sanctipaulelum Won and Iams 2015
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†Sanctipaulelum biglobosum Won and Iams 2015
†Sanctipaulelum contractum Won and Iams 2015
†Sanctipaulelum indivisum Won and Iams 2015
†Sanctipaulelum inusitata Won and Iams 2015
†Sanctipaulelum pingue Won and Iams 2015
†Sanctipaulelum superpingue Won and Iams 2015
G. †Saturnulus Haeckel 1879
G. Schizomma Ehrenberg 1861
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Schizomma quadrilobum Ehrenberg 1862
G. Sethocorys Haeckel 1882
G. Sethodisculus Haeckel 1887
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Sethodisculus radians Ehrenberg 1841
G. Soreuma Haeckel 1882
G. Spirillina Ehrenberg 1858
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Spirillina imperforata Ehrenberg 1858
G. †Spirocapsa Rüst
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Invalid names: Spirocapsa singularis Rüst 1892 [nomen dubium]
G. †Spongocyrtis Rüst
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Invalid names: Spongocyrtis eurydictyum Rüst 1892 [nomen dubium]
G. †Spongodruppa Rüst
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Invalid names: Spongodruppa ornithopus Rüst 1892 [nomen dubium], Spongodruppa triradiata Rüst 1892 [nomen dubium]
G. †Spongopila Rüst
+
Invalid names: Spongopila juniperina Rüst 1892 [nomen dubium]
G. †Staurosphaerella Demanet
+
G. Stichopera Haeckel 1882
+
Invalid names: Stichopera aculeata Rüst 1892 [nomen dubium]
G. Stichopilidium Haeckel 1887
G. Stichopodium Haeckel 1882
G. Stichopterygium Haeckel 1882
G. †Stomatosphaera Dreyer 1889
G. Stylactis Ehrenberg 1873
+
Stylactis pacifica Ehrenberg 1873
G. †Stylotrochus Haeckel 1862
+
Invalid names: Stylotrochus dolfussi Rüst 1892 [nomen dubium]
G. †Tepka Sanfilippo et al. 1973
G. Tessarastrum Haeckel 1887
G. Tessarospyris Haeckel 1882
G. Tetracorethra Haeckel 1882
G. †Tetraedroclathrum Deflandre 1960
+
G. Thalassoplancta Haeckel
Superfm. Thallassosphaeracea Haeckel 1862
+
Fm. Lithacanthidae Popovsky 1907
G. Thamnospyris Haeckel 1882
G. Theoconus Haeckel 1887
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Invalid names: Theosyringium praecox Rüst 1892 [nomen dubium]
G. Trigonactinium Haeckel 1887
G. †Ulcundia Nazarov 1974
+
†Ulcundia incomperta Nazarov 1974
G. †Veles Sugiyama 1997
G. Xiphatractylis Haeckel 1887
+
Xiphatractylis spinulosa Ehrenberg 1874
G. Zygospyris Haeckel 1887
Invalid names: Carposphaeridium Chapman 1923 [nomen dubium], Caryosphaera [nomen dubium], Cenellipsis [nomen dubium], Cenodiscus [nomen dubium], Ceratoclathrulum Deflandre 1960 [nomen nudum], Cessipylorum [nomen nudum], Choenicosphaera Ruedemann and Wilson 1936 [nomen dubium], Clavata Feng 1992 [nomen dubium], Cornutanna Rüst 1892 [nomen dubium], Diploplegma Hinde 1890 [nomen dubium], Ectoactinia Liu 1992 [nomen dubium], Ehrenbergia Liu 1995 [nomen dubium], Guangxitrisphaera Wang and Kuang 1993 [nomen dubium], Helminentactinia Li 1991 [nomen dubium], Hemisphaera Li 1995 [nomen nudum], Litheliacea Haeckel 1862 [empty], Lithocannosphaeropsis Deflandre 1960 [nomen dubium], Palaeolithochytris Li 1995 [nomen dubium], Palamphimorphium Ters and Deflandre 1966 [nomen nudum], Paleocenosphaera Nazarov 1973 [nomen dubium], Papinochium Ters and Deflandre 1966 [nomen nudum]
Invalid names: Heliozoia [empty]
+
+
Subph. Lobosa Carpenter 1861
+
Or. Arcellinida Kent 1880
+
Superfm. Arcellacea Ehrenberg 1832
Superfm. Cryptodifflugiacea Jung 1942
G. Nebela Leidy 1875
Invalid names: Amoebida [empty], Mycetozoida [empty]
G. Pelomyxa Greef 1874
Invalid names: Acanthamoebidae [empty], Conosa [empty], Entamoebida [empty], Flabellinea [empty], Mastigamoebidae [empty], Stereomyxida [empty], Tubulinea [empty]
+
Invalid names: Apusozoa [empty], Varisulca [empty]
Invalid names: Biciliata [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
No ecological data are available

Age range

Maximum range based only on fossils: base of the Templetonian to the top of the Holocene or 508.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 504.5 Ma

Collections (3477 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Templetonian508.0 - 504.5Australia (Queensland) Archeoentactinia angulata, Spongomassa nannosphaera, Palaeospiculum georginaensis, Palaeospiculum arcussimile, Palaeospiculum burkensis, Palaeospiculum radiatum (90974)
Floran504.5 - 503.0Australia (Queensland) Fungomacula sp. 1, Fungomacula barbatula, Archeoentactinia tetractinia, Archeoentactinia incaensis, Archeoentactinia hexactinia, Spongomassa gigantea, Palaeospiculum arcussimile (90972) Fungomacula sp. 1, Fungomacula sp. 3, Fungomacula barbatula, Archeoentactinia tetractinia, Archeoentactinia incaensis, Archeoentactinia hexactinia, Spongomassa gigantea, Palaeospiculum arcussimile (90973)
Mindyallan499.0 - 497.0Australia (Queensland) Echidnina runnegari (233863)
Franconian497.0 - 486.85Canada (Newfoundland and Labrador) Curvechidnina longispinosa, Echidnina irregularis, Subechidnina florea, Pararcheoentactinia reedae, Palaeospiculum greenpointensis (90966) Parechidnina jamesi, Curvechidnina tetractina, Curvechidnina kozuri, Curvechidnina fusa, Echidnina sp., Echidnina irregularis, Echidnina stevensi, Subechidnina gracilis, Subechidnina pressa, Subechidnina delicata, Protoentactinia gracilispinosa, Pararcheoentactinia reedae, Palaeospiculum martinpointenis, Palaeospiculum grosmornensis, Palaeospiculum tetraradiatum, Palaeospiculum greenpointensis (90967) Parechidnina jamesi, Curvechidnina tetractina, Curvechidnina longispinosa, Curvechidnina kozuri, Curvechidnina fusa, Echidnina sp., Echidnina irregularis, Echidnina curvata, Echidnina stevensi, Subechidnina gracilis, Subechidnina pressa, Subechidnina delicata, Subechidnina florea, Protoentactinia gracilispinosa, Pararcheoentactinia reedae, Palaeospiculum martinpointenis, Palaeospiculum grosmornensis, Palaeospiculum hexaradiatum (90968) Radiolaria indet. 1, Radiolaria indet. A, Parechidnina jamesi, Parechidnina aspinosa, Curvechidnina tetractina, Curvechidnina longispinosa, Curvechidnina fusa, Echidnina sp., Echidnina irregularis, Echidnina monactina, Subechidnina florea, Protoentactinia gracilispinosa, Pararcheoentactinia reedae, Palaeospiculum martinpointenis, Palaeospiculum grosmornensis, Palaeospiculum tetraradiatum, Palaeospiculum greenpointensis (90965)
Trempealeauan490.2 - 486.85Canada (Newfoundland and Labrador) Echidnina irregularis, Pararcheoentactinia reedae (90969) Echidnina sp., Echidnina irregularis, Protoentactinia gracilispinosa, Pararcheoentactinia impedia, Pararcheoentactinia reedae (90970) Parechidnina jamesi, Parechidnina aspinosa, Grosmorneus sp. A., Grosmorneus globosus, Echidnina sp., Echidnina curvata, Echidnina conjuncta, Protoentactinia gracilispinosa, Pararcheoentactinia impedia, Pararcheoentactinia reedae, Palaeospiculum tetraradiatum, Palaeospiculum hexaradiatum, Ramuspiculum trifurcatum, Ramuspiculum curvatum, Ramuspiculum multiramosum (90971)
Tremadocian486.85 - 477.1Canada (Newfoundland and Labrador) Echidnina sp., Echidnina bengtsoni, Westernbrookia diversa, Westernbrookia ovata, Protoentactinia transformis, Protoentactinia kozuriana, Protoentactinia gracilispinosa, Protoentactinia primigena, Protospongentactinia spongiosa, Pararcheoentactinia stilla, Pararcheoentactinia cowheadensis, Pararcheoentactinia reedae, Palaeospiculum neofurcatum, Neopalaeospiculum sp., Neopalaeospiculum transformum, Protoproventocitum aitchisoni, Archeoproventocitum nudiformum, Archeoproventocitum retiformum (234141) Echidnina sp., Echidnina bengtsoni, Westernbrookia ovata, Westernbrookia cancella, Protoentactinia transformis, Protoentactinia kozuriana, Protoentactinia gracilispinosa, Protoentactinia primigena, Protospongentactinia sp., Protospongentactinia spongiosa, Pararcheoentactinia stilla, Pararcheoentactinia cowheadensis, Pararcheoentactinia reedae, Palaeospiculum neofurcatum, Neopalaeospiculum laxum, Neopalaeospiculum transformum, Neopalaeospiculum densum, Protoproventocitum aitchisoni, Cowheadia duplextesta, Archeoproventocitum nudiformum, Archeoproventocitum retiformum (234138) Protoentactinia gracilispinosa, Pararcheoentactinia reedae, Protoproventocitum aitchisoni, Protoproventocitum nazarovi (234142) Protoentactinia transformis, Protoentactinia gracilispinosa, Pararcheoentactinia reedae, Protoproventocitum aitchisoni, Protoproventocitum nazarovi (234143)
Tremadocian486.85 - 477.1Mexico Radiolaria indet. (236919)
Tremadocian486.85 - 477.1Russian Federation (Siberia) Desmochitina minor, Desmochitina sp. (205623)
Tremadoc486.4 - 477.1Russian Federation (Gorny Altai) Desmochitina minor, Desmochitina erinacea (203936 203939)
Tremadoc486.4 - 477.1USA (Nevada) Parechidnina nevadensis, Echidnina bengtsoni, Protoentactinia sp. A, Protoentactinia sp. B, Protoentactinia sp. C, Protoentactinia minuta, Noblella tremadociensis, Protoentactinia gracilispinosa, Protoentactinia latospinosa (90927)
Moridunian477.1 - 473.8Kazakhstan Proventocitum procerulum, Inanihella bakanasensis, Inanihella akzhala (38015)
Floian477.1 - 471.3Russian Federation (Siberia) Lagenochitina deunffi (205381)
Floian477.1 - 471.3China (Hubei) Belonechitina henryi (203695)
Dapingian - Darriwilian471.3 - 458.2Oman Belonechitina micracantha, Laufeldochitina baculiformis, Lagenochitina obeligis (90376)
Darriwilian469.4 - 458.2Oman Belonechitina micracantha, Euconochitina vulgaris (90377)
Kunda469.1 - 464.3Estonia (Ida-Viru) Lagenochitina esthonica (195558)
Llanvirn468.0 - 458.2Argentina Cyathochitina sp., Desmochitina sp., Rhabdochitina sp., Lagenochitina sp. (8888)
Llanvirn468.0 - 458.2Norway Radiolaria indet., Archaeospicularia indet., Protoentactinia sp., Sphaeroentactinia hexaspinosa, Palaeospiculum sp., Proventocitum procerulum, Haplentactinia armillata, Inaniguttidae indet., Inanigutta penrosei, Inanigutta sp. A, Inanigutta sp. B, Inanigutta magnifica (90922) Radiolaria indet., Protoentactinia sp., Sphaeroentactinia hexaspinosa, Proventocitum procerulum, Haplentactinia armillata, Inaniguttidae indet., Inanigutta penrosei, Inanigutta sp. B (90921)
Lasnamagi461.4 - 459.7Latvia Cyathochitina sebyensis, Desmochitina minor, Tanuchitina tallinnensis, Rhabdochitina gracilis (172964)
Lasnamagi461.4 - 459.7Estonia Cyathochitina campanulaeformis, Cyathochitina sebyensis, Belonechitina crinita, Belonechitina micracantha, Laufeldochitina striata, Tanuchitina tallinnensis, Pterochitina retracta, Euconochitina primitiva (210664)