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Behemotops proteus

Mammalia - Perissodactyla

Taxonomy
Behemotops proteus was named by Domning et al. (1986). Its type specimen is USNM 244035, a mandible, and it is a 3D body fossil. Its type locality is LACMVP 5412, which is in a Chattian offshore mudstone in the Pysht Formation of Washington.

Synonymy list
YearName and author
1986Behemotops proteus Domning et al. p. 6 figs. Fig. 4-15
1994Behemotops proteus Ray et al. p. 205
1996Behemotops proteus Domning p. 394
2001Behemotops proteus Barnes and Goedert p. 8
2008Behemotops proteus Domning p. 642
2015Behemotops proteus Beatty and Cockburn p. 2
2017Behemotops proteus Matsui p. 4
2019Behemotops proteus Matsui and Tsuihiji
2023Behemotops proteus Beatty

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
RankNameAuthor
Mammaliamorpha
Mammaliaformes
classMammalia
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Euungulata
Panperissodactyla
superorderPerissodactylamorpha
orderPerissodactyla()
orderDesmostylia
genusBehemotops
speciesproteus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Behemotops proteus Domning et al. 1986
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Diagnosis
ReferenceDiagnosis
B. L. Beatty and T. C. Cockburn 2015Dental formula is 3.1.2?.3/3? 1.2?.3. Differs from Cornwallius in: having three pairs of upper incisors; molar cusps not in longitudinal rows, not columnar, and not closely appressed to each other, but are brachydont and retain spaces between them; molars retain labial and lingual cin- gula; upper molars, specifically M3, lack styles/stylids in between the main cusps/cuspids; coronoid process of mandible is more strongly posteriorly inclined. Differs from Paleoparadoxia in: incisors not enlarged and flattened; incisor arch not flat or straight; molar cusps not in longitudinal rows, columnar, or closely appressed to each other; postorbital process of jugal pres- ent, but not articulating with a dorsally expanded zygomatic pro- cess of the squamosal. Differs from Desmostylus in: having three pairs of upper incisors; molars brachydont; postorbital process of jugal not contacting zygomatic process of squamosal because the latter is not dorsoventrally expanded.
Measurements
No measurements are available
Composition: hydroxyapatiteo
Ontogeny: modification of partso
Environment: coastalo
Locomotion: actively mobileo
Life habit: amphibiouso
Diet: herbivoreo
Reproduction: viviparouso
Created: 2005-09-11 17:03:02
Modified: 2019-05-28 17:19:55
Source: o = order
Reference: Domning et al. 1986

Age range: base of the Rupelian to the top of the Chattian or 33.90000 to 23.03000 Ma

Collections (3 total)


Time interval Ma Country or state Original ID and collection number
Rupelian33.9 - 28.1USA (Washington) Behemotops proteus (58964)
Chattian28.1 - 23.03Canada (British Columbia) Behemotops proteus (173157)
Chattian28.1 - 23.03USA (Washington) Behemotops proteus (type locality: 45809)