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Mellivorinae (ratel)

Mammalia - Carnivora - Mustelidae

Taxonomy
Mellivorinae was named by Gray (1865) [also attributed to Gill 1872]. It is extant.

It was assigned to Mustelidae by Bryant (1968), Bjork (1970), Hendey (1974), Edwards (1976), Ginsburg (1977), Kurten and Anderson (1980), Muizon (1982) and Valenciano et al. (2015).

Synonymy list
YearName and author
1865Mellivorinae Gray
1968Mellivorinae Bryant p. 2
1970Mellivorinae Bjork p. 19
1974Mellivorinae Hendey p. 68
1976Mellivorinae Edwards p. 2
1977Mellivorinae Ginsburg p. 226
1980Mellivorinae Kurten and Anderson p. 155
1982Mellivorinae Muizon p. 262 figs. Fig. 1
2015Mellivorinae Valenciano et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
infraorderCynodontia()
—Mammaliamorpha
—Mammaliaformes
classMammalia
RankNameAuthor
—Theriamorpha(Rowe 1993)
—Theriiformes()
—Trechnotheria
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—Laurasiatheria
—Scrotifera
—Ferae()
—CarnivoramorphaWyss and Flynn 1993
—CarnivoraformesFlynn et al.
orderCarnivora
suborderCaniformiaKretzoi 1943
infraorderCanoidea(Simpson 1931)
superfamilyArctoideaFlower 1869
—Mustelida
familyMustelidaeFischer 1817
subfamilyMellivorinaeGray 1865
subfamilyMellivorinaeGray 1865

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Mellivorinae Gray 1865 [ratel]
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Tr. †Eomellivorini Valenciano and Govender 2020
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G. †Ekorus Werdelin 2003
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†Ekorus ekakeran Werdelin 2003
G. †Eomellivora Zdansky 1924
+
†Eomellivora fricki Pia 1939
†Eomellivora hungarica Kretzoi 1942
†Eomellivora piveteaui Ozansoy 1965
+
Invalid names: Eomellivora liguritor Crusafont-Pairo and Ginsburg 1973 [synonym]
†Eomellivora tugenensis Morales and Pickford 2005
†Eomellivora ursogulo Orlov 1948
†Eomellivora wimani Zdansky 1924
+
Invalid names: Eomellivora orlovi Kretzoi 1965 [synonym], Eomellivora rumana Simionescu 1938 [synonym]
Invalid names: Perunium Orlov 1947 [synonym]
G. †Moralesictis Valenciano and Baskin 2022
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†Moralesictis intrepidus Valenciano and Baskin 2022
G. †Hoplictis Ginsburg 1961
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†Hoplictis grangerensis Bryant 1968
†Hoplictis noueli Mayet 1908
Invalid names: Beckia Bryant 1968 [synonym]
Tr. †Mellivorini Valenciano and Govender 2020
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G. †Howellictis de Bonis et al. 2009
+
†Howellictis valentini de Bonis et al. 2009
G. Mellivora Storr 1780 [ratel]
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†Mellivora benfieldi Hendy 1978
Mellivora capensis Schreber 1776 [honey badger]
Mellivora indica Kerr 1792
†Mellivora punjabiensis Lydekker 1884
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: scansorialf
Diet: carnivoref
Diet 2: omnivoref
Reproduction: viviparoussubc
Created: 2005-06-08 10:11:09
Modified: 2005-08-22 19:38:29
Source: f = family, subc = subclass, c = class, subp = subphylum
References: Hendy et al. 2009, Ji et al. 2002, Nowak 1991, Carroll 1988, Lillegraven 1979

Age range

Maximum range based only on fossils: base of the Tortonian to the top of the Holocene or 11.62000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 11.608 Ma

Collections (63 total)


Time interval Ma Country or state Original ID and collection number
Miocene23.03 - 5.333USA (Washington) Beckia grangerensis (18903)
Miocene - Pliocene23.03 - 2.588South Africa Mellivora capensis (22229)
Middle Miocene15.97 - 11.608India (Jammu and Kashmir) Eomellivora sp. (11797)
Clarendonian12.5 - 9.4USA (California) Beckia grangerensis (19391) Eomellivora wimani (19393)
Late/Upper Clarendonian12.5 - 9.4USA (Florida) Ischyrictis sp. (18554)
Tortonian11.62 - 7.246Germany (Bavaria) Eomellivora piveteaui (206345)
Tortonian11.62 - 7.246Ukraine (Odessa) Eomellivora wimani (75797)
Tortonian11.62 - 7.246China (Yunnan) Eomellivora wimani (11798)
Tortonian11.62 - 7.246Ukraine Perunium ursogulo (75800)
Tortonian - Messinian11.62 - 5.333Kenya Mellivorinae indet., Ekorus ekakeran (22241)
Tortonian - Zanclean11.62 - 3.6Ethiopia Mellivora benfieldi (22139)
Vallesian11.608 - 8.7Spain (Segovia) Eomellivora liguritor (40768)
Vallesian11.608 - 8.7Greece Eomellivora wimani (191070)
MN 911.1 - 9.7Slovakia (Trnava) Eomellivora wimani (69719)
Hemphillian10.3 - 4.9USA (Texas) Eomellivora sp. (18072 18198)
Turolian8.7 - 5.333China (Yunnan) Eomellivora wimani (44221)
Messinian7.246 - 5.333Italy (Ravenna) Mellivora benfieldi (144566)
Messinian - Zanclean7.246 - 3.6Kenya Mellivorinae indet. (22242)
Messinian - Piacenzian7.246 - 2.588South Africa (Cape) Mellivora punjabiensis, Mellivora capensis (59167)
Early/Lower Pliocene5.333 - 3.6Senegal Mellivorinae indet. (221108)
Zanclean5.333 - 3.6South Africa Mellivora benfeldi (22221 22222)
Pliocene5.333 - 2.588Tanzania (Eastern Drift Valley) Mellivora capensis (32050)
Late/Upper Pliocene3.6 - 2.588South Africa (Langebaanweg) Mellivora punjabiensis (21713)
Late/Upper Pliocene3.6 - 2.588Morocco Mellivora capensis (32052)
Piacenzian3.6 - 2.588Tanzania Mellivora capensis (21818)
Pleistocene2.588 - 0.0117Kenya Mellivora capensis (21963)
Pleistocene2.588 - 0.0117South Africa Mellivora capensis (21695 21916 21929 21931 21956 115191 115205) Mellivora sivalensis (21825) Mellivora sp. (21920)
Pleistocene - Holocene2.588 - 0.0Morocco Mellivora sp. (21856 21857)
Pleistocene - Holocene2.588 - 0.0Swaziland Mellivora capensis (21925)
Pleistocene - Holocene2.588 - 0.0South Africa Mellivora capensis (21711 21948 21949 21951 21952)
Pleistocene - Holocene2.588 - 0.0Zambia Mellivora capensis (21924)
Middle Pleistocene0.774 - 0.129Morocco Mellivora sp. (34287)
Middle Pleistocene - Late/Upper Pleistocene0.774 - 0.0117Kenya Mellivora capensis (21276 21277)
Late/Upper Pleistocene0.129 - 0.0117South Africa Mellivora capensis (90070 115184)
Late/Upper Pleistocene (interglacial)0.129 - 0.0117South Africa Mellivora capensis (115167)
Late/Upper Pleistocene0.129 - 0.0117Kenya (Athi-Kapiti plains) Mellivora capensis (32115 32124)
Late/Upper Pleistocene - Holocene0.129 - 0.0South Africa Mellivora capensis (21922)
Holocene0.0117 - 0.0Sudan Mellivora capensis (21903 208787)
Holocene0.0117 - 0.0South Africa (Orange Free) Mellivora capensis (21911)
Holocene0.0117 - 0.0South Africa (N. Transraal) Mellivora capensis (21693)
Holocene0.0117 - 0.0South Africa Mellivora capensis (21921 21930)
Holocene0.0117 - 0.0Tanzania Mellivora capensis (21690)