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Bemella
Taxonomy
Bemella was named by Rozanov et al. (1969) [Missarzhevsky in Rozanov et al. 1969; Sepkoski's age data: V N-Da-u Cm mMid-l Sepkoski's reference number: 268,699,885,1030]. Its type is Helcionella jacutica.
It was replaced with Securiconus by Vasil’yeva (1998).
It was assigned to Securiconidae by Missarzhevsky (1989); to Cyrtonellida by Bengtson (1992); to Stenothecidae by Elicki (1994); to Helcionelloida by Sepkoski (2002); to Helcionellinae by Parkhaev and Demidenko (2010); and to Helcionellidae by Rozanov et al. (1969), Runnegar and Jell (1976), Parkhaev (2002), Kouchinsky et al. (2017) and Wagner (2023).
It was replaced with Securiconus by Vasil’yeva (1998).
It was assigned to Securiconidae by Missarzhevsky (1989); to Cyrtonellida by Bengtson (1992); to Stenothecidae by Elicki (1994); to Helcionelloida by Sepkoski (2002); to Helcionellinae by Parkhaev and Demidenko (2010); and to Helcionellidae by Rozanov et al. (1969), Runnegar and Jell (1976), Parkhaev (2002), Kouchinsky et al. (2017) and Wagner (2023).
Species
B. ambita, B. amplaperata (syn. B. emeiensis), B. atava, B. barskovae (syn. B. minuta), B. bella, B. communis, B. flexa, B. incomparabilis, B. jacutica, B. malycanica, B. minuta, B. multicarinata, B. parula, B. rotundata, B. septata, B. villemsonae, B. vonbitteri, B. xinjiensis, Kalbyella poulseni, Sacciconus yanjiaheensis
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1969 | Bemella Rozanov et al. p. 163 |
| 1976 | Bemella Runnegar and Jell p. 116 |
| 1978 | Kalbyella Berg-Madsen and Peel |
| 1979 | Latirostratus Yü p. 244 |
| 1980 | Sacciconus Jiang p. 118 |
| 1984 | Sichuanospira He |
| 1989 | Bemella Missarzhevsky p. 175 |
| 1990 | Charaulachella Vassiljeva |
| 1992 | Bemella Bengtson p. 1026 |
| 1994 | Bemella Elicki p. 83 |
| 1994 | Repenoconus Feng et al. p. 4 |
| 2000 | Striatoconus Feng et al. p. 370 |
| 2002 | Bemella Parkhaev p. 34 |
| 2002 | Bemella Sepkoski |
| 2002 | Kalbyella Sepkoski |
| 2002 | Latirostratus Sepkoski |
| 2002 | Rostroconus Sepkoski |
| 2002 | Sacciconus Sepkoski |
| 2002 | Sichuanospira Sepkoski |
| 2010 | Bemella Parkhaev and Demidenko pp. 1024, 1025 |
| 2017 | Bemella Kouchinsky et al. p. 334 |
| 2023 | Bemella Wagner p. G1331 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Bemella Rozanov et al. 1969
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†Bemella ambita Barskova 1987
†Bemella atava He 1984
†Bemella bella Chen and Zhang 1980
†Bemella communis Parkhaev 2001
†Bemella flexa Barskova 1988
†Bemella incomparabilis Parkhaev 2001
†Bemella jacutica Rozanov and Missarzhevsky 1966
†Bemella malycanica Rozanov and Missarzhevsky 1966
†Bemella minuta Luo et al. 1982
†Bemella multicarinata Chen and Zhang 1980
†Bemella parula Rozanov et al. 1969
†Bemella rotundata Chen et al. 1981
†Bemella septata Rozanov and Missarzhevsky 1966
†Bemella villemsonae Barskova 1988
†Bemella vonbitteri Landing et al. 1989
†Bemella xinjiensis Feng et al. 1994
Invalid names: Charaulachella Vassiljeva 1990 [synonym], Kalbyella Berg-Madsen and Peel 1978 [synonym], Latirostratus Yü 1979 [synonym], Repenoconus Feng et al. 1994 [synonym], Rostroconus [synonym], Sacciconus Jiang 1980 [synonym], Sichuanospira He 1984 [synonym], Striatoconus Feng et al. 2000 [synonym]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| P. Y. Parkhaev and Y. Demidenko 2010 | "Shell cap-shaped, low or even depressed, moderately compressed laterally. Apex inclined posteriorly and shifted up to posterior apertural margin and usually projects over it. Anterior field convex, posterior concave, rather short. Aperture simple, without sinuses and flaring, oval or irregularly-oval. Ornamentation usually represented by concentric folds, rarely radial ribs or striation can be present. |
Measurements
No measurements are available
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| Source: g = genus, f = family, c = class | |||||
| References: Hendy 2009, Feng et al. 2000, Hendy et al. 2009 | |||||
Age range: base of the Meishucunian to the top of the Undillan or 530.70000 to 500.50000 Ma
Collections (78 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Nemakit-Daldynian | Russian Federation (Khabarovsk) | B. jacutica (99465) | |
| Nemakit-Daldynian | Russian Federation (Khabarovsk Krai) | Rostroconus sp., B. jacutica (50367) | |
| Nemakit-Daldynian | Russian Federation (Krasnoyarsk) | B. sp. (99473) | |
| Nemakit-Daldynian - Tommotian | Russian Federation (Sakha) | B. jacutica (99229) | |
| Placentian | USA (Massachusetts) | B. bella (56829) | |
| Cambrian | China (Yunnan) | B. jacutica (201549 201563 201564 201565 201566 201567 201576 201585 201586 201587 201590 201591 201594) | |
| Meishucunian - Nemakit-Daldynian | China (Yunnan) | Striatoconus sp. (87093) | |
| Meishucunian | China (Yunnan) | Rostroconus sp. (108257 108258) Rostroconus sp., Latirostratus amplaperatus (108259) | |
| Meishucunian | China (Anhui) | Sacciconus sacciformis (107772) | |
| Meishucunian | China (Hubei) | B. rotundata (108415) Sacciconus yanjiaheensis (106525) | |
| Meishucunian | China (Shaanxi) | Rostroconus multisegmentata (107790) | |
| Meishucunian | China (Sichuan) | B. sp. (107389) Sichuanospira atava (107390) | |
| Meishucunian - Tommotian | China (Yunnan) | B. emeiensis (50504) | |
| Tommotian | Russian Federation (Yakutia) | B. jacutica (45850 45852 96245) B. jacutica, B. parula (45848) B. jacutica, B. parula, B. septata (45847 45849) B. jacutica, B. septata (45851) | |
| Tommotian | Canada (Newfoundland and Labrador) | B. vonbitteri (87332 87334 87353) | |
| Tommotian | Russian Federation (Uchur-Maya) | B. ambita (9998) | |
| Tommotian | Russian Federation (Khabarovsk Krai) | B. jacutica, B. septata (50314) | |
| Tommotian | Russian Federation (Krasnoyarsk) | B. jacutica (99484) | |
| Tommotian | Russian Federation (Sakha) | B. jacutica (233169) B. malycanica (10046) | |
| Tommotian | Russian Federation (Krasnoyarsk Krai) | B. jacutica (96392 96396 233166) B. jakutica (232906) Rostroconus sp., B. jacutica (50429) | |
| Tommotian | Russian Federation | B. jacutica (90089) | |
| Tommotian | Russian Federation (Yakutiya) | B. jacutica (56858 90088 96391 96394) B. jacutica, B. malycanica (80796) B. jacutica, B. parula (96226) Helcionella jacutica, B. malycanica (96251) | |
| Tommotian | USA (Massachusetts) | B. bella (56825) | |
| Tommotian | China (Hubei) | Latirostratus amplaperatus (56807) Sacciconus yanjiaheensis (155441) | |
| Tommotian | Russian Federation (Siberia) | B. jacutica (50319 50321) B. jacutioa (50428) | |
| Tommotian - Botomian | Iran | B. sp. (10530) | |
| Atdabanian | Russian Federation (Siberia) | B. flexa, B. minuta (10011) B. sp. (174257) | |
| Nangaoian | China (Henan) | Repenoconus xinjiensis (96525) | |
| Atdabanian - Lenian | Germany (Saxony) | B. jacutica (6899) | |
| Botomian | Australia (South Australia) | B. communis (67980 67981 96958 96959 97730) | |
| Toyonian | Australia (South Australia) | B. communis (97991) | |
| Amgan | Denmark | Kalbyella poulseni (9876) | |
| Solvan | Morocco (High Atlas) | B. bella (50548) | |
| Solvan | Australia (New South Wales) | Helcionopsis sp. (56811) | |
| Undillan | Germany | B. sp. A (186151) |