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Haptophyta (haptophyte alga)

Taxonomy

Haptophyta was named by Hibberd ex Cavalier-Smith (1986). It is extant.

It was assigned to Chromista by Young and Bown (1997); and to Chromalveolata by Adl et al. (2005).

Synonymy list

Year	Name and author
1986	Haptophyta Hibberd ex Cavalier-Smith
1997	Haptophyta Young and Bown
2005	Haptophyta Adl et al. p. 401 figs. Table 1

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Rank	Name	Author
—	Life
—	Eukaryota	()
—	Chromalveolata

Rank	Name	Author
phylum	Haptophyta	Hibberd ex Cavalier-Smith 1986
phylum	Haptophyta	Hibberd ex Cavalier-Smith 1986

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Ph. Haptophyta Hibberd ex Cavalier-Smith 1986 [haptophyte alga]

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G. Aspidorhabdus Hay and Towe 1962

Cl. Coccolithophyceae Rothmaler 1951

G. †Biantholithus Bramlette and Martini 1964

Or. Braarudosphaerales Aubry 2013

Fm. †Polycyclolithaceae Forchheimer 1972

G. †Assipetra Roth 1973

†Assipetra infracretacea Thierstein 1973

†Assipetra terebrodentarius Applegate and Bralower 1987

G. †Eprolithus Stover 1966

†Eprolithus antiquus Perch-Nielsen 1979

†Eprolithus apertior Black 1973

†Eprolithus floralis Stradner 1962

†Eprolithus moratus Stover 1966

†Eprolithus octopetalus Varol 1992

†Eprolithus rarus Varol 1992

Invalid names: Polycyclolilthus Forchheimer 1968 [synonym]

G. †Farhania Varol 1992

†Farhania varolii Jakubowski 1986

G. †Hayesites Manivit 1971

†Hayesites albiensis Manivit 1971

†Hayesites irregularis Thierstein et al. 1972

G. †Hexalithus Gardet 1955

†Hexalithus gardetiae Bukry 1969

†Hexalithus noeliae Loeblich and Tappan 1966

G. †Lithastrinus Stradner 1962

†Lithastrinus grillii Stradner 1962

†Lithastrinus pentabrachius Varol 1992

†Lithastrinus quadricuspis Farhan 1997

†Lithastrinus septenarius Forchheimer 1972

G. †Micula Vekshina 1959 [coccolithophore]

†Micula adumbrata Burnett 1998

†Micula clypeata Lees and Bown 2005

†Micula concava Stradner 1960

†Micula cubiformis Forchheimer 1972

†Micula decussata Vekshina 1959 [coccolithophore]

†Micula murus Martini 1961

†Micula praemurus Bukry 1973

†Micula premolisilvae Lees and Bown 2005

†Micula prinsii Perch-Nielsen 1979

†Micula staurophora Gardet 1955

†Micula swastica Stradner and Steinmetz 1984

Invalid names: Nannotetraster Martini and Stradner 1960 [synonym]

G. †Polycostella Thierstein 1971

†Polycostella beckmanii Thierstein 1971

†Polycostella senaria Thierstein 1971

G. †Quadrum Prins and Perch-Nielsen 1977

†Quadrum bengalensis Burnett 1998

†Quadrum eneabrachium Varol 1992

†Quadrum eptabrachium Varol 1992

†Quadrum gartneri Prins and Perch-Nielsen 1977

†Quadrum giganteum Varol 1992

†Quadrum intermedium Varol 1992

†Quadrum octobrachium Varol 1992

†Quadrum svabenickae Burnett 1998

G. †Radiolithus Stover 1966

†Radiolithus hollandicus Varol 1992

†Radiolithus orbiculatus Forchheimer 1972

†Radiolithus planus Stover 1966

G. †Rucinolithus Stover 1966

†Rucinolithus magnus Bukry 1975

†Rucinolithus windleyae Rutledge and Bown 1996

†Rucinolithus wisei Thierstein 1971

G. †Tegulalithus Crux 1986

†Tegulalithus septentrionalis Stradner 1963

†Tegulalithus tessellatus Stradner 1968

G. †Uniplanarius Hattner and Wise 1980

†Uniplanarius clarkei Lees and Bown 2005

†Uniplanarius gothicus Deflandre 1959

†Uniplanarius sissinghii Perch-Nielsen 1986

†Uniplanarius trifidus Stradner 1961

Fm. Ceratolithaceae Norris 1965

G. Amaurolithus Gartner and Bukry 1975

Amaurolithus primus Bukry and Percival 1971

G. Angulolithina Bukry 1973

G. Ceratolithus Kamptner 1950

G. Florisphaera Okada and Honjo 1973

G. †Nicklithus Raffi et al. 1998

Nicklithus amplificus Bukry and Percival 1971

G. †Orthorhabdus Bramlette and Wilcoxon 1967

Or. †Coccolithales Schwarz 1932

Fm. Coccolithaceae Poche 1913

G. †Bicolumnus Wei and Wise 1990

G. †Bramletteius Gartner 1969

G. †Campylosphaera Kamptner 1963

G. †Chiasmolithus Hay et al. 1966

G. †Clausicoccus Prins 1979

G. Coronocyclus Hay et al. 1966

G. †Ericsonia Black 1964

G. †Holodiscolithus Roth 1970

G. †Hughesius Varol 1989

G. †Solidopons Theodoridis 1984

Invalid names: Hymenomonadaceae Senn 1900 [empty], Pleurochrysidaceae Fresnel and Billard 1991 [empty]

G. †Daktylethra Gartner and Bukry 1969

Or. †Discoasterales Hay 1977

Fm. †Fasciculithaceae Hay and Mohler 1967

G. †Fasciculithus Bramlette and Sullivan 1961

G. †Lithoptychius Aubry 2011

Fm. †Heliolithaceae Hay and Mohler 1967

G. †Bomolithus Roth 1973

G. †Heliolithus Bramlette and Sullivan 1961

Invalid names: Discoasteraceae Tan Sin Hok 1927 [empty]

G. †Ellipsolithus Sullivan 1964

G. Gladiolithus Jordan and Chamberlain 1992

G. †Ilselithina Stradner and Adamiker 1966

Or. Isochrysidales Pascher 1910

Fm. Gephyrocapsaceae Heimdal 1997

G. Crenalithus Roth 1973

Fm. Noelaerhabdaceae Jerkovic 1970

G. Apertapetra Hay et al. 1966

G. Cepekiella Roth 1970

G. †Cribrocentrum Perch-Nielsen 1971

G. Dictyococcites Black 1967

G. Emiliania Hay and Mohler 1967

G. Gephyrocapsa Kamptner 1943

G. †Pseudoemiliania Gartner 1969

Fm. †Prinsiaceae Hay and Mohler 1967

G. †Girgisia Varol 1989

G. Hornibrookina Edwards 1973

G. †Neobiscutum Varol 1989

G. †Prinsius Hay and Mohler 1967

Invalid names: Isochrysidaceae Pascher 1910 [empty]

G. †Lanternithus Stradner 1962

G. †Minylitha Bukry 1973

G. Neosphaera Lecal-Schlauder 1949

G. †Pedinocyclus Bukry and Bramlette 1971

Fm. †Rhomboasteraceae Bown 2005

G. †Rhomboaster Bramlette and Sullivan 1961

G. †Tribrachiatus Shamrai 1963

G. Tetralithoides Theodoridis 1984

G. †Transversopontis Hay et al. 1966

Or. Zygodiscales Young and Bown 1997

Fm. Pontosphaeraceae Lemmermann 1908

G. Calciopilleus Müller 1974

G. Scyphosphaera Lohmann 1902

Fm. Zygodiscaceae Hay and Mohler 1967

G. †Isthmolithus Deflandre and Fert 1954

†Isthmolithus recurvus Deflandre 1954

G. †Lophodolithus Deflandre and Fert 1954

†Lophodolithus mochlophorus Deflandre 1954

G. †Nannotetrina Achuthan and Stradner 1969

G. †Neococcolithes Sujkowski 1931

G. †Zygodiscus Bramlette and Sullivan 1961

Invalid names: Helicosphaeraceae Black 1971 [empty]

Invalid names: Phaeocystales Medlin 2000 [empty], Prymnesiales Papenfuss 1955 [empty], Prymnesiophyceae Hibberd 1976 [synonym]

G. Conococcolithus Hay and Mohler 1967

G. Corannulus Stradner 1962

G. Craspedolithus Kamptner 1963

G. Crassapontosphaera Boudreaux and Hay 1969

G. Cyclolithella Loeblich and Tappan 1963

Fm. †Discoasteridae

G. †Agalmatoaster Klumpp 1953

†Agalmatoaster septemradiatus Klumpp 1953

G. †Trochoaster Klumpp 1953

†Trochoaster simplex Klumpp 1953

G. Ellipsodiscoaster Boudreaux and Hay 1969

G. †Helicosphaera Kamptner 1954

†Helicosphaera carteri Wallich 1877

†Helicosphaera neolophota Barron et al. 1984

†Helicosphaera orientalis Black 1971

†Helicosphaera stalis Theodoridis 1984

Invalid names: Helicopontosphaera Hay and Mohler 1967 [synonym]

G. Heliorthus Brönnimann and Stradner 1960

G. Koczyia Boudreaux and Hay 1969

G. Leptodiscus Bukry and Bramlette 1969

Fm. Lithostromationaceae Deflandre 1959

G. Imperiaster Martini 1970

G. Lithostromation Deflandre 1942

G. Nannocorbis Müller 1974

G. Orthozygus Bramlette and Wilcoxon 1967

G. Pyrocyclus Hay and Towe 1962

G. Reticulofenestra Hay et al. 1966

†Reticulofenestra minuta Roth 1970

†Reticulofenestra pseudoumbilicus Gartner 1967

†Reticulofenestra rotaria Theodoridis 1984

G. Rhabdolithina Reinhardt 1967

G. Sphenaster Wilcoxon 1970

G. Striatococcolithus Bukry 1971

G. †Syracosphaera Lohmann 1902

G. Umbilicosphaera Lohmann 1902

†Umbilicosphaera jafari Muller 1974

Invalid names: Geminilithella Backman 1980 [synonym]

G. Vermiculithina Bukry and Percival 1971

G. Wiseorhabdus Bukry 1981

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

low Mg calcitep

Environment:

oceanic, oligotrophic, marine, brackishp

Locomotion:

passively mobilep

Life habit:

planktonicp

Depth habitat:

surfacep

Diet:

"photoautotroph"p

Created:

2006-10-31 06:24:05

Modified:

2006-10-31 08:24:05

Source: p = phylum

Reference: Kiessling 2003

Age range

Maximum range based only on fossils: base of the Late/Upper Famennian to the top of the Ionian or 364.70000 to 0.12600 Ma
Minimum age of oldest fossil (stem group age): 360.7 Ma

Collections (388 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Late/Upper Famennian	364.7 - 360.7	USA (Missouri)	Leptodiscus corrugatus (6027 6029)
Middle Hettangian - Late/Upper Hettangian	201.3 - 196.5	Canada (Alberta)	Mitrolithus jansae, Mitrolithus elegans, Crucirhabdus primulus, Parhabdolithus liasicus, Crepidolithus crassus, Tubirhabdus patulus (197630)
Hettangian - Sinemurian	201.3 - 190.8	Austria	Schizosphaerella punctulata (38056 38057 38058 38059)
Late/Upper Sinemurian - Late/Upper Pliensbachian	196.5 - 183.0	Morocco	Calcivascularis jansae, Crucirhabdus primulus, Parhabdolithus robustus, Parhabdolithus liasicus, Similiscutum cruciulus, Calyculus sp., Schizosphaerella spp., Schizosphaerella sp., Crepidolithus crassus (205771) Calcivascularis jansae, Schizosphaerella sp. (205765) Calcivascularis jansae, Schizosphaerella sp., Schizosphaerella spp. (205777) Lotharingius frodoi, Calcivascularis jansae, Mitrolithus elegans, Parhabdolithus robustus, Parhabdolithus liasicus, Similiscutum cruciulus, Schizosphaerella sp., Schizosphaerella spp. (205772) Lotharingius frodoi, Lotharingius haufii, Calcivascularis jansae, Mitrolithus elegans, Parhabdolithus liasicus, Similiscutum finchii, Similiscutum cruciulus (205774) Lotharingius frodoi, Lotharingius haufii, Lotharingius sigillatus, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus liasicus, Similiscutum finchii, Calyculus sp., Schizosphaerella sp., Schizosphaerella spp. (205779) Lotharingius haufii, Calcivascularis jansae, Similiscutum novum, Similiscutum finchii, Schizosphaerella sp., Schizosphaerella spp., Crepidolithus crassus (205778) Lotharingius haufii, Mitrolithus elegans, Schizosphaerella sp. (205776) Mitrolithus elegans, Mitrolithus lenticularis, Crucirhabdus primulus, Calyculus sp., Schizosphaerella sp., Crepidolithus crassus, Tubirhabdus patulus (205769) Mitrolithus lenticularis, Schizosphaerella sp. (205764) Schizosphaerella sp. (205759 205760 205762 205763 205767 205768 205770 205775) Schizosphaerella sp., Schizosphaerella spp. (205761 205773)
Pliensbachian	190.8 - 182.7	Spain (Jaén)	Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201752) Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Biscutum depravatum, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Calyculus recondita, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201756) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus elegans, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201755) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201753) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201754) Lotharingius sigillatus, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus sp, Schizosphaerella punctulata, Crepidolithus cavus, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201758) Lotharingius sigillatus, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus cavus, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201759)
Spinatum	189.6 - 183.0	Morocco	Lotharingius sigillatus (203735 203736)
Early/Lower Toarcian	183.0 - 182.0	Spain (Granada)	Lotharingius velatus, Lotharingius crucicentralis, Watznaueria colacicchii, Biscutum intermedium, Discorhabdus striatus, Similiscutum sp. 1, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199494)
Toarcian	182.7 - 174.1	Spain (Jaén)	Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Tubirhabdus patulus (201675) Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Diductius constans, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201750) Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Diductius constans, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus depressa, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201678) Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201751) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus elegans, Mitrolithus lenticularis, Diductius constans, Parhabdolithus distinctus, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus depressa, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201667) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Axopodorhabdus atavus, Carinolithus poulnabronei, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus cavus, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201668) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum depravatum, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus recondita, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Tubirhabdus patulus (201673) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201670) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Calyculus depressa, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201671) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Diductius constans, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201674) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Diductius constans, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Carinolithus poulnabronei, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201669) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201677) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Carinolithus poulnabronei, Calyculus depressa, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201666) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201685) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Carinolithus poulnabronei, Calyculus depressa, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201665) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum grande, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus recondita, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201672) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Bussonius prinsii, Calcivascularis jansae, Mitrolithus lenticularis, Diductius constans, Crucirhabdus primulus, Parhabdolithus distinctus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Axopodorhabdus atavus, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus (201676)
Toarcian	182.7 - 174.1	Spain (Murcia)	Discorhabdus striatus, Schizosphaerella punctulata, Crepidolithus crassus (201778) Lotharingius crucicentralis, Lotharingius hauffii, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201781) Lotharingius hauffii, Discorhabdus striatus, Carinolithus superbus, Schizosphaerella punctulata (201774) Lotharingius sigillatus, Biscutum intermedium, Discorhabdus striatus, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201787) Lotharingius sigillatus, Discorhabdus striatus, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201792) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Bussonius prinsii, Watznaueria colacicchii, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201779) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Bussonius prinsii, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Triscutum sullivanii, Biscutum dubium, Biscutum intermedium, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201789) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201776) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Watznaueria colacicchii, Watznaueria sp. B, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201769) Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus, Zeugrhabdotus erectus (201790) Lotharingius sigillatus, Lotharingius hauffii, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201772) Lotharingius sigillatus, Lotharingius hauffii, Bussonius prinsii, Watznaueria colacicchii, Watznaueria sp. A, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201777) Lotharingius sigillatus, Lotharingius hauffii, Watznaueria colacicchii, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus (201771) Lotharingius sigillatus, Lotharingius hauffii, Watznaueria colacicchii, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201780) Lotharingius sigillatus, Watznaueria colacicchii, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201788) Lotharingius umbriensis, Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Watznaueria colacicchii, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus (201768) Lotharingius umbriensis, Lotharingius velatus, Lotharingius sigillatus, Lotharingius hauffii, Bussonius prinsii, Watznaueria colacicchii, Watznaueria sp. A, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201783) Lotharingius velatus, Lotharingius sigillatus, Bussonius prinsii, Watznaueria colacicchii, Biscutum dubium, Biscutum intermedium, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201793) Lotharingius velatus, Lotharingius sigillatus, Bussonius prinsii, Watznaueria colacicchii, Watznaueria sp. A, Triscutum sullivanii, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201773) Lotharingius velatus, Lotharingius sigillatus, Lotharingius barozii, Lotharingius hauffii, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Biscutum dubium, Biscutum intermedium, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201791) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Watznaueria colacicchii, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201794) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Watznaueria colacicchii, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201764) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Bussonius prinsii, Watznaueria colacicchii, Biscutum dubium, Biscutum intermedium, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201775) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Watznaueria colacicchii, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201765 201766) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Triscutum sullivanii, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201770) Lotharingius velatus, Lotharingius sigillatus, Lotharingius hauffii, Watznaueria colacicchii, Watznaueria sp. A, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201782)
Toarcian	182.7 - 174.1	Saudi Arabia	Ellipsagelosphaera britannica, Tetralithus goticus (45191) Tetralithus gothicus, Ellipsagelosphaera britannica (45089)
Levesquei	182.0 - 175.6	Germany	Bussonius leufuensis (59428) Lotharingius crucicentralis, Crucirhabdus primulus, Triscutum sullivanii, Retecapsa incompta, Carinolithus superbus, Crepidolithus crassus (59434) Lotharingius velatus, Lotharingius crucicentralis, Bussonius leufuensis, Watznaueria contracta, Crucirhabdus primulus, Triscutum tiziense, Triscutum sullivanii, Retecapsa incompta, Discorhabdus criotus, Carinolithus crassus, Carinolithus superbus, Schizosphaerella sp., Crepidolithus crassus (59430) Lotharingius velatus, Watznaueria contracta, Retecapsa incompta, Discorhabdus criotus, Schizosphaerella sp. (59431)
Middle Toarcian	182.0 - 175.6	Spain (Murcia)	Bussonius prinsii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Crepidolithus crassus (201809) Lotharingius hauffii, Biscutum dubium, Discorhabdus cf. D. criotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201810) Lotharingius hauffii, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201811) Lotharingius hauffii, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201805) Lotharingius hauffii, Mitrolithus lenticularis, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201808) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201799) Lotharingius sigillatus, Lotharingius hauffii, Mitrolithus lenticularis, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201803) Lotharingius sigillatus, Lotharingius hauffii, Watznaueria colacicchii, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201795) Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Watznaueria sp. A, Mitrolithus lenticularis, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201796) Lotharingius umbriensis, Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Watznaueria colacicchii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201801) Lotharingius umbriensis, Lotharingius velatus, Lotharingius sigillatus, Lotharingius hauffii, Mitrolithus lenticularis, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201802) Lotharingius velatus, Lotharingius hauffii, Biscutum dubium, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201807) Lotharingius velatus, Lotharingius hauffii, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201806) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Watznaueria colacicchii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Schizosphaerella punctulata, Crepidolithus crassus (201804) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Watznaueria colacicchii, Mitrolithus lenticularis, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201798) Lotharingius velatus, Lotharingius sigillatus, Lotharingius hauffii, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201812) Lotharingius velatus, Lotharingius sigillatus, Lotharingius hauffii, Bussonius prinsii, Watznaueria colacicchii, Watznaueria sp. A, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201797) Lotharingius velatus, Lotharingius sigillatus, Lotharingius hauffii, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Schizosphaerella punctulata, Crepidolithus crassus (201813) Lotharingius velatus, Lotharingius sigillatus, Lotharingius hauffii, Watznaueria colacicchii, Mitrolithus lenticularis, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201800)
Late/Upper Toarcian	180.1 - 175.6	Spain (Granada)	Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus (202117) Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus (202110) Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (202114) Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202103) Discorhabdus striatus, Schizosphaerella punctulata, Crepidolithus crassus (202106) Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Schizosphaerella punctulata, Crepidolithus crassus (202108) Lotharingius crucicentralis, Watznaueria colacicchii, Watznaueria sp. B, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199492) Lotharingius crucicentralis, Watznaueria sp. A, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202109) Lotharingius sigillatus, Bussonius prinsii, Watznaueria sp. A, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202099) Lotharingius sigillatus, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus sp. cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (202097) Lotharingius sigillatus, Lotharingius crucicentralis, Bussonius prinsii, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199187) Lotharingius sigillatus, Lotharingius crucicentralis, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199496) Lotharingius sigillatus, Lotharingius crucicentralis, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202100) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus, Zeugrhabdotus erectus (199177) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Biscutum depravatum, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (201880) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Watznaueria sp. B, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Calyculus sp, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus, Zeugrhabdotus erectus (199488) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Bussonius prinsii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Biscutum depravatum, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus, Zeugrhabdotus erectus (199166) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (201948) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Watznaueria colacicchii, Watznaueria sp. A, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus, Zeugrhabdotus erectus (199229) Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Watznaueria colacicchii, Watznaueria sp. A, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199195) Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria colacicchii, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199487) Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria colacicchii, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199248) Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria colacicchii, Watznaueria sp. B, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199173) Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Biscutum depravatum, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus (199167) Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria sp. A, Watznaueria sp. B, Biscutum depravatum, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (199181) Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria sp. A, Watznaueria sp. B, Biscutum dubium, Biscutum intermedium, Discorhabdus sp. cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus, Zeugrhabdotus erectus (199490) Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria sp. B, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199491) Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria sp. B, Biscutum dubium, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (199231) Lotharingius sigillatus, Watznaueria colacicchii, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199198) Lotharingius sigillatus, Watznaueria colacicchii, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus (202105) Lotharingius sigillatus, Watznaueria colacicchii, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (202115) Lotharingius sigillatus, Watznaueria sp. A, Discorhabdus striatus, Carinolithus cantaluppii, Carinolithus poulnabronei, Schizosphaerella punctulata, Crepidolithus crassus (202112) Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria colacicchii, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Zeugrhabdotus erectus (199253) Lotharingius umbriensis, Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria colacicchii, Triscutum sullivanii, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199495) Lotharingius umbriensis, Lotharingius velatus, Lotharingius sigillatus, Biscutum dubium, Discorhabdus sp. cf. D. criotus, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (202101) Lotharingius umbriensis, Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Watznaueria sp. A, Watznaueria sp. B, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus, Zeugrhabdotus erectus (199247) Lotharingius velatus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202111) Lotharingius velatus, Discorhabdus striatus, Similiscutum novum, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202116) Lotharingius velatus, Lotharingius crucicentralis, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (199235) Lotharingius velatus, Lotharingius sigillatus, Discorhabdus striatus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus (202098) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Bussonius prinsii, Discorhabdus striatus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (202107) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Bussonius prinsii, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199169) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Triscutum sullivanii, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus, Zeugrhabdotus erectus (199165) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius barozii, Watznaueria colacicchii, Watznaueria sp. A, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199493) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Bussonius prinsii, Watznaueria sp. B, Biscutum depravatum, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus poulnabronei, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199172) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Similiscutum sp. 1, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199164) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Lotharingius hauffii, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Triscutum sullivanii, Biscutum dubium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus, Zeugrhabdotus erectus (199171) Lotharingius velatus, Lotharingius sigillatus, Lotharingius crucicentralis, Watznaueria colacicchii, Watznaueria sp. A, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus criotus, Discorhabdus striatus, Discorhabdus ignotus, Similiscutum novum, Carinolithus superbus, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus (199168) Schizosphaerella punctulata, Crepidolithus crassus (202113) Watznaueria sp. A, Discorhabdus striatus, Similiscutum novum, Schizosphaerella punctulata, Crepidolithus crassus (202104) Watznaueria sp. B, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Similiscutum novum, Carinolithus poulnabronei, Schizosphaerella punctulata (202102)
Late/Upper Toarcian	180.1 - 175.6	Germany	Lotharingius haufforum, Lotharingius sp. 3, Lotharingius sp. 4, Lotharingius sigillatus, Lotharingius contractus, Lotharingius barozii, Bussonius sp. 1, Bussonius leufuensis, Bussonius prinsii, Parhabdolithus cavus, Stradnerlithus humilis, Biscutum striatum, Palaeopontosphaera dubia, Biscutum intermedium, Discorhabdus novus, Axopodorhabdus cylindratus, Axopodorhabdus atavus, Carinolithus superbus, Calyculus sp. 1, Calyculus sp. 2, Calyculus noelae, Catillus hommerilii, Calyculus cribrum, Proculithus fistulatus, Schizosphaerella punctulata, Crepidolithus crassus, Vagalapilla quadriarculla, Tubirhabdus patulus, Zeugrhabdotus choffatii, Zeugrhabdotus erectus (59451) Lotharingius haufforum, Lotharingius sp. 4, Lotharingius sigillatus, Lotharingius contractus, Lotharingius barozii, Bussonius sp. 1, Bussonius leufuensis, Bussonius prinsii, Crucirhabdus magnus, Parhabdolithus cavus, Stradnerlithus humilis, Triscutum beaminsterensene, Triscutum sullivanii, Biscutum criotum, Biscutum sp. 2, Biscutum sp. 4, Biscutum striatum, Palaeopontosphaera dubia, Axopodorhabdus depravatus, Biscutum intermedium, Discorhabdus novus, Axopodorhabdus cylindratus, Axopodorhabdus atavus, Ethmorhabdus crucifer, Carinolithus sceptrum, Carinolithus superbus, Calyculus noelae, Catillus hommerilii, Calyculus cribrum, Proculithus charlottei, Schizosphaerella punctulata, Crepidolithus crassus, Tubirhabdus patulus, Zeugrhabdotus choffatii, Zeugrhabdotus erectus (59453) Lotharingius sp. 2, Lotharingius haufforum, Lotharingius sp. 3, Lotharingius sp. 4, Lotharingius sigillatus, Lotharingius contractus, Lotharingius barozii, Bussonius sp. 1, Bussonius leufuensis, Bussonius prinsii, Crucirhabdus magnus, Parhabdolithus cavus, Triscutum pricatillus, Biscutum criotum, Biscutum striatum, Palaeopontosphaera sp. 1, Palaeopontosphaera dubia, Axopodorhabdus depravatus, Biscutum intermedium, Discorhabdus novus, Axopodorhabdus cylindratus, Axopodorhabdus atavus, Carinolithus sp. 1, Carinolithus sceptrum, Carinolithus superbus, Calyculus sp. 2, Proculithus fistulatus, Proculithus charlottei, Schizosphaerella punctulata, Crepidolithus crassus, Vagalapilla quadriarculla, Tubirhabdus patulus, Zeugrhabdotus choffatii, Zeugrhabdotus sp. 1, Zeugrhabdotus sp. 2, Zeugrhabdotus erectus (59449) Retecapsa incompta, Discorhabdus striatus, Carinolithus superbus, Schizosphaerella sp., Crepidolithus crassus (59454)
Early/Lower Aalenian	175.6 - 171.6	Germany	Bussonius prinsii, Calyculus sp. (59458) Bussonius prinsii, Retecapsa incompta, Discorhabdus striatus, Carinolithus superbus, Calyculus sp., Schizosphaerella sp., Crepidolithus crassus (59457)
Bajocian	170.3 - 168.3	Spain	Cyclagelosphaera margerelii, Watznaueria communis, Watznaueria contracta, Watznaueria britannica (214379)
Bajocian	170.3 - 168.3	Saudi Arabia	Cyclagelosphaera margereli (45268) Tetralithus gothicus, Ellipsagelosphaera britannica, Schizosphaerella punctulata (45265)
Bathonian	168.3 - 166.1	Saudi Arabia	Tetralithus gothicus, Cyclagelosphaera margereli, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Hexalithus magharensis, Schizosphaerella punctulata (45464) Tetralithus gothicus, Diazomatolithus lehmani, Cyclagelosphaera margereli, Ellipsagelosphaera britannica (45519)
Callovian	166.1 - 163.5	Saudi Arabia	Cyclagelosphaera margereli, Ellipsagelosphaera communis (45555) Cyclagelosphaera margereli, Watznaueria manivitae, Ellipsagelosphaera britannica, Ellipsagelosphaera communis (45540) Cyclagelosphaera margereli, Watznaueria manivitae, Ellipsagelosphaera communis, Ellipsagelosphaera britannica, Stephanolithion bigoti, Polypodorhabdus escaigii, Ethmorhabdus anglicus, Zeugrhabdotus erectus (47890) Tetralithus gothicus, Cyclagelosphaera margereli (45558) Tetralithus gothicus, Cyclagelosphaera margereli, Ellipsagelosphaera britannica, Ellipsagelosphaera communis (45549) Tetralithus gothicus, Diazomatolithus lehmani, Cyclagelosphaera margereli, Watznaueria manivitae, Ellipsagelosphaera communis, Ellipsagelosphaera britannica (45547) Zygodiscus sp., Tetralithus gothicus, Cyclagelosphaera margereli, Watznaueria manivitae, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Stephanolithion bigoti, Discorhabdus sp., Ethmorhabdus gallicus, Zeugrhabdotus erectus (45541)
Oxfordian	163.5 - 157.3	Saudi Arabia	Cyclagelosphaera margereli, Watznaueria manivitae, Ellipsagelosphaera communis (46052) Watznaueria manivitae, Biscutum sp., Ethmorhabdus sp., Vekshinella stradneri (45896)