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Haptophyta (haptophyte alga)

Synonymy list
YearName and author
1986Haptophyta Hibberd ex Cavalier-Smith
1997Haptophyta Young and Bown
2005Haptophyta Adl et al. p. 401 figs. Table 1

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RankNameAuthor
Life
Eukaryota()
Chromalveolata
RankNameAuthor
phylumHaptophytaHibberd ex Cavalier-Smith 1986
phylumHaptophytaHibberd ex Cavalier-Smith 1986

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Ph. Haptophyta Hibberd ex Cavalier-Smith 1986 [haptophyte alga]
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G. Aspidorhabdus Hay and Towe 1962
G. †Camuralithus de Kaenel and Villa 1996
Cl. Coccolithophyceae Rothmaler 1951
+
G. †Athenagalea Hattner and Wise 1980
G. †Biantholithus Bramlette and Martini 1964
Or. Braarudosphaerales Aubry 2013
+
Fm. †Polycyclolithaceae Forchheimer 1972
+
G. †Assipetra Roth 1973
+
Assipetra infracretacea Thierstein 1973
Assipetra terebrodentarius Applegate and Bralower 1987
G. †Eprolithus Stover 1966
+
Eprolithus antiquus Perch-Nielsen 1979
Eprolithus apertior Black 1973
Eprolithus floralis Stradner 1962
Eprolithus moratus Stover 1966
Eprolithus octopetalus Varol 1992
Eprolithus rarus Varol 1992
Invalid names: Polycyclolilthus Forchheimer 1968 [synonym]
G. †Farhania Varol 1992
+
Farhania varolii Jakubowski 1986
G. †Hayesites Manivit 1971
+
Hayesites albiensis Manivit 1971
Hayesites irregularis Thierstein et al. 1972
G. †Hexalithus Gardet 1955
+
Hexalithus gardetiae Bukry 1969
Hexalithus noeliae Loeblich and Tappan 1966
G. †Lithastrinus Stradner 1962
+
Lithastrinus grillii Stradner 1962
Lithastrinus septenarius Forchheimer 1972
G. †Micula Vekshina 1959 [coccolithophore]
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Micula adumbrata Burnett 1998
Micula clypeata Lees and Bown 2005
Micula concava Stradner 1960
Micula cubiformis Forchheimer 1972
Micula decussata Vekshina 1959 [coccolithophore]
Micula murus Martini 1961
Micula praemurus Bukry 1973
Micula premolisilvae Lees and Bown 2005
Micula prinsii Perch-Nielsen 1979
Micula staurophora Gardet 1955
Micula swastica Stradner and Steinmetz 1984
Invalid names: Nannotetraster Martini and Stradner 1960 [synonym]
G. †Polycostella Thierstein 1971
+
Polycostella beckmanii Thierstein 1971
Polycostella senaria Thierstein 1971
G. †Quadrum Prins and Perch-Nielsen 1977
+
Quadrum bengalensis Burnett 1998
Quadrum eneabrachium Varol 1992
Quadrum eptabrachium Varol 1992
Quadrum gartneri Prins and Perch-Nielsen 1977
Quadrum giganteum Varol 1992
Quadrum intermedium Varol 1992
Quadrum octobrachium Varol 1992
Quadrum svabenickae Burnett 1998
G. †Radiolithus Stover 1966
+
Radiolithus orbiculatus Forchheimer 1972
Radiolithus planus Stover 1966
G. †Rucinolithus Stover 1966
+
Rucinolithus magnus Bukry 1975
Rucinolithus windleyae Rutledge and Bown 1996
Rucinolithus wisei Thierstein 1971
G. †Tegulalithus Crux 1986
+
Tegulalithus tessellatus Stradner 1968
G. †Uniplanarius Hattner and Wise 1980
+
Uniplanarius clarkei Lees and Bown 2005
Uniplanarius gothicus Deflandre 1959
Uniplanarius sissinghii Perch-Nielsen 1986
Uniplanarius trifidus Stradner 1961
Fm. Ceratolithaceae Norris 1965
+
G. Amaurolithus Gartner and Bukry 1975
+
Amaurolithus primus Bukry and Percival 1971
G. Angulolithina Bukry 1973
G. Ceratolithus Kamptner 1950
G. Florisphaera Okada and Honjo 1973
G. †Nicklithus Raffi et al. 1998
+
Nicklithus amplificus Bukry and Percival 1971
G. †Orthorhabdus Bramlette and Wilcoxon 1967
Or. †Coccolithales Schwarz 1932
+
Fm. Coccolithaceae Poche 1913
+
G. †Bicolumnus Wei and Wise 1990
G. †Birkelundia Perch-Nielsen 1971
G. †Bramletteius Gartner 1969
G. †Campylosphaera Kamptner 1963
G. †Chiasmolithus Hay et al. 1966
+
Chiasmolithus bidens Bramlette and Sullivan 1961
Chiasmolithus consuetus Bramlette and Sullivan 1961
Chiasmolithus danicus Hay and Mohler 1967
Chiasmolithus eograndis Perch-Nielsen 1971
Chiasmolithus expansus Bramlette and Sullivan 1961
Chiasmolithus grandis Bramlette and Riedel 1954
Chiasmolithus modestus Perch-Nielsen 1971
Chiasmolithus oamaruensis Deflandre 1954
Chiasmolithus solitus Bramlette and Sullivan 1961
G. †Clausicoccus Prins 1979
G. Coronocyclus Hay et al. 1966
G. †Ericsonia Black 1964
G. †Gephyrorhabdus Hill 1976
G. †Holodiscolithus Roth 1970
+
Holodiscolithus solidus Deflandre and Fert 1954
G. †Hughesius Varol 1989
G. †Praeprinsius Varol and Jakubowski 1989
G. †Solidopons Theodoridis 1984
Invalid names: Hymenomonadaceae Senn 1900 [empty], Pleurochrysidaceae Fresnel and Billard 1991 [empty]
G. †Daktylethra Gartner and Bukry 1969
Or. †Discoasterales Hay 1977
+
Fm. †Fasciculithaceae Hay and Mohler 1967
+
G. †Fasciculithus Bramlette and Sullivan 1961
G. †Lithoptychius Aubry 2011
Fm. †Heliolithaceae Hay and Mohler 1967
+
G. †Bomolithus Roth 1973
G. †Heliolithus Bramlette and Sullivan 1961
Invalid names: Discoasteraceae Tan Sin Hok 1927 [empty]
Fm. †Ellipsagelosphaeraceae Noël 1965
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Subfm. †Actinosphaeroideae Noël 1965
+
G. †Calolithus Noël 1965
G. †Ellipsolithus Sullivan 1964
G. Gladiolithus Jordan and Chamberlain 1992
G. †Ilselithina Stradner and Adamiker 1966
Or. Isochrysidales Pascher 1910
+
Fm. Gephyrocapsaceae Heimdal 1997
+
G. Crenalithus Roth 1973
Fm. Noelaerhabdaceae Jerkovic 1970
+
G. Apertapetra Hay et al. 1966
G. Cepekiella Roth 1970
G. †Cribrocentrum Perch-Nielsen 1971
G. Dictyococcites Black 1967
+
Dictyococcites scrippsae Bukry and Percival 1973
G. Emiliania Hay and Mohler 1967
G. Gephyrocapsa Kamptner 1943
G. †Pseudoemiliania Gartner 1969
Fm. †Prinsiaceae Hay and Mohler 1967
+
G. †Futyania Varol 1989
G. †Girgisia Varol 1989
G. Hornibrookina Edwards 1973
+
Hornibrookina teuriensis Edwards 1973
Hornibrookina weimerae Self-Trail 2011
G. †Neobiscutum Varol 1989
G. †Prinsius Hay and Mohler 1967
+
Prinsius dimorphosus Perch-Nielsen 1977
Prinsius martinii Haq 1971
Prinsius tenuiculus Perch-Nielsen 1984
Invalid names: Isochrysidaceae Pascher 1910 [empty]
G. †Lanternithus Stradner 1962
+
Lanternithus minutus Stradner 1962
G. †Minylitha Bukry 1973
G. Neosphaera Lecal-Schlauder 1949
G. †Pedinocyclus Bukry and Bramlette 1971
Fm. †Rhomboasteraceae Bown 2005
+
G. †Rhomboaster Bramlette and Sullivan 1961
G. †Tribrachiatus Shamrai 1963
G. Tetralithoides Theodoridis 1984
G. †Transversopontis Hay et al. 1966
Or. Zygodiscales Young and Bown 1997
+
Fm. Pontosphaeraceae Lemmermann 1908
+
G. Calciopilleus Müller 1974
G. Scyphosphaera Lohmann 1902
Fm. Zygodiscaceae Hay and Mohler 1967
+
G. †Isthmolithus Deflandre and Fert 1954
+
Isthmolithus recurvus Deflandre 1954
G. †Lophodolithus Deflandre and Fert 1954
+
Lophodolithus mochlophorus Deflandre 1954
G. †Nannotetrina Achuthan and Stradner 1969
G. †Neococcolithes Sujkowski 1931
+
Neococcolithes dubius Deflandre and Fert 1954
G. †Zygodiscus Bramlette and Sullivan 1961
Invalid names: Helicosphaeraceae Black 1971 [empty]
Invalid names: Phaeocystales Medlin 2000 [empty], Prymnesiales Papenfuss 1955 [empty], Prymnesiophyceae Hibberd 1976 [synonym]
G. Conococcolithus Hay and Mohler 1967
G. Corannulus Stradner 1962
G. Craspedolithus Kamptner 1963
G. Crassapontosphaera Boudreaux and Hay 1969
G. †Crassidiscus Okada 1990
G. †Cricosphaera Braarud 1960
G. Cyclolithella Loeblich and Tappan 1963
G. †Cycloperfolithus Lehotayova and Priewalder 1978
+
G. †Agalmatoaster Klumpp 1953
+
G. †Trochoaster Klumpp 1953
+
Trochoaster simplex Klumpp 1953
G. †Duplexipodorhabdus Varol and Girgis 1994
G. Ellipsodiscoaster Boudreaux and Hay 1969
G. †Helicosphaera Kamptner 1954
+
Helicosphaera carteri Wallich 1877
Helicosphaera lophota Bramlette and Sullivan 1961
Helicosphaera neolophota Barron et al. 1984
Helicosphaera stalis Theodoridis 1984
Invalid names: Helicopontosphaera Hay and Mohler 1967 [synonym]
G. Heliorthus Brönnimann and Stradner 1960
+
Heliorthus tenuis Stradner 1961
G. Koczyia Boudreaux and Hay 1969
G. Leptodiscus Bukry and Bramlette 1969
Fm. Lithostromationaceae Deflandre 1959
+
G. Imperiaster Martini 1970
G. Lithostromation Deflandre 1942
G. Nannocorbis Müller 1974
G. Orthozygus Bramlette and Wilcoxon 1967
G. †Pervilithus Crux 1981
G. †Phanulithus Wise and Wind 1977
G. Pyrocyclus Hay and Towe 1962
G. Reticulofenestra Hay et al. 1966
+
Reticulofenestra dictyoda Stradner 1968
Reticulofenestra reticulata Roth and Thierstein 1972
Reticulofenestra rotaria Theodoridis 1984
Reticulofenestra scrippsae Bukry and Percival 1973
G. Rhabdolithina Reinhardt 1967
G. Sphenaster Wilcoxon 1970
G. Striatococcolithus Bukry 1971
G. †Syracosphaera Lohmann 1902
+
Syracosphaera heimii Lohmann 1919
Syracosphaera mediterranea Gaarder and Heirndal 1977
G. Umbilicosphaera Lohmann 1902
+
Umbilicosphaera jafari Muller 1974
Invalid names: Geminilithella Backman 1980 [synonym]
G. Vermiculithina Bukry and Percival 1971
G. Wiseorhabdus Bukry 1981
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: low Mg calcitep
Environment: oceanic, oligotrophic, marine, brackishp
Locomotion: passively mobilep
Life habit: planktonicp
Depth habitat: surfacep
Diet: "photoautotroph"p
Created: 2006-10-31 06:24:05
Modified: 2006-10-31 08:24:05
Source: p = phylum
Reference: Kiessling 2003

Age range

Maximum range based only on fossils: base of the Late/Upper Famennian to the top of the Ionian or 364.70000 to 0.12900 Ma
Minimum age of oldest fossil (stem group age): 360.7 Ma

Collections (444 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Late/Upper Famennian364.7 - 360.7USA (Missouri) Leptodiscus corrugatus (6027 6029)
Middle Hettangian - Late/Upper Hettangian201.3 - 196.5Canada (Alberta) Crepidolithus crassus, Tubirhabdus patulus, Mitrolithus elegans, Mitrolithus jansae, Parhabdolithus liasicus, Crucirhabdus primulus (197630)
Hettangian - Sinemurian201.3 - 190.8Austria Schizosphaerella punctulata (38056 38057 38058 38059)
Late/Upper Sinemurian - Late/Upper Pliensbachian196.5 - 183.0Morocco Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus liasicus, Crucirhabdus primulus, Schizosphaerella sp., Schizosphaerella spp., Calyculus sp., Similiscutum finchii, Lotharingius frodoi, Lotharingius haufii, Lotharingius sigillatus, Bussonius prinsii (205779) Calcivascularis jansae, Schizosphaerella sp. (205765) Calcivascularis jansae, Schizosphaerella sp., Schizosphaerella spp. (205777) Crepidolithus crassus, Calcivascularis jansae, Parhabdolithus liasicus, Parhabdolithus robustus, Crucirhabdus primulus, Schizosphaerella spp., Schizosphaerella sp., Calyculus sp., Similiscutum cruciulus (205771) Crepidolithus crassus, Calcivascularis jansae, Schizosphaerella sp., Schizosphaerella spp., Similiscutum novum, Similiscutum finchii, Lotharingius haufii (205778) Crepidolithus crassus, Tubirhabdus patulus, Mitrolithus elegans, Mitrolithus lenticularis, Crucirhabdus primulus, Schizosphaerella sp., Calyculus sp. (205769) Mitrolithus elegans, Calcivascularis jansae, Parhabdolithus liasicus, Parhabdolithus robustus, Schizosphaerella sp., Schizosphaerella spp., Similiscutum cruciulus, Lotharingius frodoi (205772) Mitrolithus elegans, Calcivascularis jansae, Parhabdolithus liasicus, Similiscutum finchii, Similiscutum cruciulus, Lotharingius frodoi, Lotharingius haufii (205774) Mitrolithus elegans, Schizosphaerella sp., Lotharingius haufii (205776) Mitrolithus lenticularis, Schizosphaerella sp. (205764) Schizosphaerella sp. (205759 205760 205762 205763 205767 205768 205770 205775) Schizosphaerella sp., Schizosphaerella spp. (205761 205773)
Pliensbachian190.8 - 182.7Spain (Jaén) Crepidolithus cavus, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201758) Crepidolithus cavus, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201759) Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201752) Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201753) Crepidolithus crucifer, Crepidolithus cavus, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Mitrolithus elegans, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus, Schizosphaerella punctulata, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201755) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Schizosphaerella punctulata, Calyculus recondita, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201756) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Schizosphaerella punctulata, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201754)
Spinatum189.6 - 183.0Morocco Lotharingius sigillatus (203735 203736)
Early/Lower Toarcian183.0 - 182.0Spain (Granada) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum sp. 1, Biscutum intermedium, Discorhabdus striatus, Lotharingius crucicentralis, Lotharingius velatus, Watznaueria colacicchii (199494)
Toarcian182.7 - 174.1Saudi Arabia Tetralithus gothicus, Ellipsagelosphaera britannica (45089) Tetralithus goticus, Ellipsagelosphaera britannica (45191)
Toarcian182.7 - 174.1Spain (Murcia) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Lotharingius hauffii, Lotharingius sigillatus (201772) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Bussonius prinsii (201775) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Bussonius prinsii (201779) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus (201776) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus superbus, Discorhabdus striatus, Lotharingius sigillatus (201792) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii (201765 201766) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Triscutum sullivanii, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B (201770) Crepidolithus crassus, Schizosphaerella punctulata, Discorhabdus striatus (201778) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Bussonius prinsii (201777) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Triscutum sullivanii, Lotharingius velatus, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Bussonius prinsii (201773) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii (201780) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B (201791) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius velatus, Lotharingius sigillatus, Watznaueria colacicchii, Bussonius prinsii (201793) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Triscutum sullivanii, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Bussonius prinsii (201789) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Lotharingius umbriensis, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Bussonius prinsii (201783) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Bussonius prinsii (201794) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius hauffii (201781) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Discorhabdus striatus, Discorhabdus criotus, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A (201782) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius sigillatus, Watznaueria colacicchii (201788) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Biscutum intermedium, Discorhabdus striatus, Lotharingius sigillatus (201787) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Lotharingius sigillatus, Watznaueria colacicchii (201764) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. B (201769) Schizosphaerella punctulata, Carinolithus superbus, Discorhabdus striatus, Lotharingius hauffii (201774) Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus superbus, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii (201771) Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus striatus, Discorhabdus criotus, Lotharingius umbriensis, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii (201768) Zeugrhabdotus erectus, Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B (201790)
Toarcian182.7 - 174.1Spain (Jaén) Crepidolithus cavus, Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Carinolithus poulnabronei, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201668) Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201677) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus depressa, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201671) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Axopodorhabdus atavus, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201672) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Calyculus depressa, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201665) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201670) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Diductius constans, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus depressa, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201678) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Diductius constans, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201750) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Diductius constans, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201674) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Diductius constans, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Biscutum dubium, Axopodorhabdus atavus, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201676) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Diductius constans, Crucirhabdus primulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201669) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Schizosphaerella punctulata, Carinolithus poulnabronei, Calyculus depressa, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201666) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Axopodorhabdus atavus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201685) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Schizosphaerella punctulata, Orthogonoides hamiltoniae, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201751) Crepidolithus crucifer, Crepidolithus crassus, Crepidolithus granulatus, Tubirhabdus patulus, Mitrolithus elegans, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Diductius constans, Schizosphaerella punctulata, Calyculus depressa, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201667) Crepidolithus crucifer, Crepidolithus crassus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus recondita, Calyculus sp, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum grande, Biscutum dubium, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201675) Crepidolithus crucifer, Crepidolithus crassus, Tubirhabdus patulus, Calcivascularis jansae, Mitrolithus lenticularis, Parhabdolithus distinctus, Crucirhabdus primulus, Schizosphaerella punctulata, Calyculus recondita, Similiscutum finchii, Similiscutum novum, Similiscutum precarium, Similiscutum cruciulus, Biscutum depravatum, Biscutum grande, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Bussonius prinsii (201673)
Levesquei182.0 - 175.6Germany Bussonius leufuensis (59428) Crepidolithus crassus, Crucirhabdus primulus, Carinolithus superbus, Retecapsa incompta, Triscutum sullivanii, Lotharingius crucicentralis (59434) Crepidolithus crassus, Crucirhabdus primulus, Schizosphaerella sp., Carinolithus crassus, Carinolithus superbus, Retecapsa incompta, Discorhabdus criotus, Triscutum tiziense, Triscutum sullivanii, Lotharingius crucicentralis, Lotharingius velatus, Watznaueria contracta, Bussonius leufuensis (59430) Schizosphaerella sp., Retecapsa incompta, Discorhabdus criotus, Lotharingius velatus, Watznaueria contracta (59431)
Middle Toarcian182.0 - 175.6Spain (Murcia) Crepidolithus crassus, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Bussonius prinsii (201809) Crepidolithus crassus, Mitrolithus lenticularis, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus criotus, Lotharingius umbriensis, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria sp. A, Bussonius prinsii (201796) Crepidolithus crassus, Mitrolithus lenticularis, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Lotharingius umbriensis, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus (201802) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Biscutum dubium, Discorhabdus striatus, Lotharingius velatus, Lotharingius hauffii (201807) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Bussonius prinsii (201804) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Similiscutum novum, Discorhabdus striatus, Discorhabdus ignotus, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus (201813) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Lotharingius hauffii (201805) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus (201799) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus superbus, Discorhabdus striatus, Lotharingius velatus, Lotharingius hauffii (201806) Crepidolithus crassus, Tubirhabdus patulus, Mitrolithus lenticularis, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Calyculus noeliae, Similiscutum novum, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Lotharingius hauffii (201808) Crepidolithus crassus, Tubirhabdus patulus, Mitrolithus lenticularis, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii (201798) Crepidolithus crassus, Tubirhabdus patulus, Mitrolithus lenticularis, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii (201800) Crepidolithus crassus, Tubirhabdus patulus, Mitrolithus lenticularis, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Lotharingius hauffii, Lotharingius sigillatus (201803) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus (201812) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Lotharingius umbriensis, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii (201801) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus cf. D. criotus, Lotharingius hauffii (201810) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Discorhabdus cf. D. criotus, Discorhabdus striatus, Lotharingius hauffii (201811) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus criotus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii (201795) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Bussonius prinsii (201797)
Late/Upper Toarcian180.1 - 175.6Spain (Granada) Crepidolithus crassus, Schizosphaerella punctulata (202113) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Discorhabdus striatus, Lotharingius velatus (202116) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Discorhabdus striatus (202110) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Discorhabdus striatus, Lotharingius sigillatus, Watznaueria sp. A (202112) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Similiscutum novum, Discorhabdus striatus, Discorhabdus criotus, Lotharingius sigillatus, Watznaueria colacicchii (202105) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus cantaluppii, Similiscutum novum, Discorhabdus striatus (202108) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Biscutum dubium, Discorhabdus striatus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria sp. B (199231) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius velatus (199235) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus (202103) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Discorhabdus criotus, Lotharingius sigillatus, Watznaueria sp. A, Bussonius prinsii (202099) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Lotharingius velatus (202111) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Lotharingius velatus, Lotharingius sigillatus (202098) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Biscutum depravatum, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria sp. A, Watznaueria sp. B (199181) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Discorhabdus striatus, Lotharingius crucicentralis, Lotharingius sigillatus (202100) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Discorhabdus striatus, Lotharingius crucicentralis, Watznaueria sp. A (202109) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Similiscutum novum, Similiscutum sp. 1, Discorhabdus striatus, Discorhabdus criotus (202117) Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus superbus, Biscutum depravatum, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B (201880) Crepidolithus crassus, Schizosphaerella punctulata, Discorhabdus striatus (202106) Crepidolithus crassus, Schizosphaerella punctulata, Similiscutum novum, Discorhabdus striatus, Watznaueria sp. A (202104) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Lotharingius sigillatus, Watznaueria colacicchii (202115) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus cantaluppii, Carinolithus poulnabronei, Discorhabdus striatus (202114) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A (199493) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B, Bussonius prinsii (199187) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Calyculus sp, Similiscutum novum, Similiscutum sp. 1, Biscutum intermedium, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius sigillatus (202097) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Lotharingius crucicentralis, Lotharingius sigillatus (199496) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Discorhabdus striatus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii (199487) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Lotharingius sigillatus, Watznaueria colacicchii (199198) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Discorhabdus striatus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. B (199173) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Biscutum depravatum, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria sp. B, Bussonius prinsii (199172) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A (199195) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Biscutum dubium, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Discorhabdus criotus, Lotharingius umbriensis, Lotharingius velatus, Lotharingius sigillatus (202101) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Triscutum sullivanii, Lotharingius umbriensis, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii (199495) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Discorhabdus striatus, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius sigillatus, Bussonius prinsii (202107) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum sp. 1, Discorhabdus striatus, Discorhabdus ignotus, Lotharingius crucicentralis, Watznaueria colacicchii, Watznaueria sp. B (199492) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Similiscutum novum, Discorhabdus striatus, Discorhabdus ignotus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii (199248) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Bussonius prinsii (199169) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B (201948) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A (199168) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria sp. B (199491) Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B (199164) Schizosphaerella punctulata, Carinolithus poulnabronei, Similiscutum novum, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Watznaueria sp. B (202102) Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius umbriensis, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria colacicchii (199253) Zeugrhabdotus erectus, Crepidolithus crassus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum depravatum, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B (199167) Zeugrhabdotus erectus, Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Biscutum depravatum, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria contracta, Watznaueria sp. A, Watznaueria sp. B, Bussonius prinsii (199166) Zeugrhabdotus erectus, Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Calyculus sp, Similiscutum novum, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius sigillatus, Watznaueria sp. B (199488) Zeugrhabdotus erectus, Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A (199229) Zeugrhabdotus erectus, Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Lotharingius umbriensis, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria sp. A, Watznaueria sp. B, Bussonius prinsii (199247) Zeugrhabdotus erectus, Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum novum, Similiscutum sp. 1, Biscutum intermedium, Discorhabdus striatus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius sigillatus (199177) Zeugrhabdotus erectus, Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus poulnabronei, Carinolithus superbus, Similiscutum sp. 1, Biscutum dubium, Biscutum intermedium, Discorhabdus sp. cf. D. criotus, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Lotharingius crucicentralis, Lotharingius sigillatus, Watznaueria sp. A, Watznaueria sp. B (199490) Zeugrhabdotus erectus, Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum depravatum, Biscutum dubium, Biscutum intermedium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Triscutum sullivanii, Lotharingius crucicentralis, Lotharingius barozii, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria contracta, Watznaueria sp. A (199165) Zeugrhabdotus erectus, Crepidolithus crassus, Tubirhabdus patulus, Schizosphaerella punctulata, Carinolithus superbus, Similiscutum novum, Biscutum dubium, Discorhabdus striatus, Discorhabdus ignotus, Discorhabdus criotus, Triscutum sullivanii, Lotharingius crucicentralis, Lotharingius velatus, Lotharingius hauffii, Lotharingius sigillatus, Watznaueria colacicchii, Watznaueria sp. A, Watznaueria sp. B (199171)
Late/Upper Toarcian180.1 - 175.6Germany Crepidolithus crassus, Schizosphaerella sp., Carinolithus superbus, Retecapsa incompta, Discorhabdus striatus (59454) Zeugrhabdotus choffatii, Zeugrhabdotus erectus, Crepidolithus crassus, Tubirhabdus patulus, Stradnerlithus humilis, Parhabdolithus cavus, Crucirhabdus magnus, Schizosphaerella punctulata, Carinolithus sceptrum, Carinolithus superbus, Calyculus noelae, Calyculus cribrum, Proculithus charlottei, Catillus hommerilii, Biscutum criotum, Biscutum sp. 2, Biscutum sp. 4, Biscutum striatum, Palaeopontosphaera dubia, Axopodorhabdus depravatus, Biscutum intermedium, Discorhabdus novus, Axopodorhabdus cylindratus, Axopodorhabdus atavus, Ethmorhabdus crucifer, Triscutum beaminsterensene, Triscutum sullivanii, Lotharingius haufforum, Lotharingius sp. 4, Lotharingius barozii, Lotharingius sigillatus, Lotharingius contractus, Bussonius sp. 1, Bussonius prinsii, Bussonius leufuensis (59453) Zeugrhabdotus choffatii, Zeugrhabdotus erectus, Crepidolithus crassus, Vagalapilla quadriarculla, Tubirhabdus patulus, Stradnerlithus humilis, Parhabdolithus cavus, Schizosphaerella punctulata, Carinolithus superbus, Calyculus sp. 1, Calyculus sp. 2, Calyculus noelae, Calyculus cribrum, Proculithus fistulatus, Catillus hommerilii, Biscutum striatum, Palaeopontosphaera dubia, Biscutum intermedium, Discorhabdus novus, Axopodorhabdus cylindratus, Axopodorhabdus atavus, Lotharingius haufforum, Lotharingius sp. 3, Lotharingius sp. 4, Lotharingius barozii, Lotharingius sigillatus, Lotharingius contractus, Bussonius sp. 1, Bussonius prinsii, Bussonius leufuensis (59451) Zeugrhabdotus choffatii, Zeugrhabdotus sp. 1, Zeugrhabdotus sp. 2, Zeugrhabdotus erectus, Crepidolithus crassus, Vagalapilla quadriarculla, Tubirhabdus patulus, Parhabdolithus cavus, Crucirhabdus magnus, Schizosphaerella punctulata, Carinolithus sp. 1, Carinolithus sceptrum, Carinolithus superbus, Calyculus sp. 2, Proculithus fistulatus, Proculithus charlottei, Biscutum criotum, Biscutum striatum, Palaeopontosphaera sp. 1, Palaeopontosphaera dubia, Axopodorhabdus depravatus, Biscutum intermedium, Discorhabdus novus, Axopodorhabdus cylindratus, Axopodorhabdus atavus, Triscutum pricatillus, Lotharingius sp. 2, Lotharingius haufforum, Lotharingius sp. 3, Lotharingius sp. 4, Lotharingius barozii, Lotharingius sigillatus, Lotharingius contractus, Bussonius sp. 1, Bussonius prinsii, Bussonius leufuensis (59449)
Early/Lower Aalenian175.6 - 171.6Germany Calyculus sp., Bussonius prinsii (59458) Crepidolithus crassus, Schizosphaerella sp., Carinolithus superbus, Calyculus sp., Retecapsa incompta, Discorhabdus striatus, Bussonius prinsii (59457)
Bajocian170.3 - 168.3Saudi Arabia Cyclagelosphaera margereli (45268) Tetralithus gothicus, Schizosphaerella punctulata, Ellipsagelosphaera britannica (45265)
Bajocian170.3 - 168.3Spain Watznaueria communis, Watznaueria contracta, Watznaueria britannica, Cyclagelosphaera margerelii (214379)
Bathonian168.3 - 166.1Saudi Arabia Tetralithus gothicus, Ellipsagelosphaera britannica, Cyclagelosphaera margereli, Diazomatolithus lehmani (45519) Tetralithus gothicus, Schizosphaerella punctulata, Hexalithus magharensis, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli (45464)
Callovian166.1 - 163.5Saudi Arabia Ellipsagelosphaera communis, Cyclagelosphaera margereli (45555) Tetralithus gothicus, Cyclagelosphaera margereli (45558) Tetralithus gothicus, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli (45549) Tetralithus gothicus, Watznaueria manivitae, Ellipsagelosphaera communis, Ellipsagelosphaera britannica, Cyclagelosphaera margereli, Diazomatolithus lehmani (45547) Tetralithus gothicus, Zeugrhabdotus erectus, Stephanolithion bigoti, Discorhabdus sp., Ethmorhabdus gallicus, Watznaueria manivitae, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli, Zygodiscus sp. (45541) Watznaueria manivitae, Ellipsagelosphaera britannica, Ellipsagelosphaera communis, Cyclagelosphaera margereli (45540) Zeugrhabdotus erectus, Stephanolithion bigoti, Polypodorhabdus escaigii, Ethmorhabdus anglicus, Watznaueria manivitae, Ellipsagelosphaera communis, Ellipsagelosphaera britannica, Cyclagelosphaera margereli (47890)
Oxfordian163.5 - 157.3Saudi Arabia Vekshinella stradneri, Biscutum sp., Ethmorhabdus sp., Watznaueria manivitae (45896) Watznaueria manivitae, Ellipsagelosphaera communis, Cyclagelosphaera margereli (46052)