PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Pararhabdodon isonensis

Reptilia - Hadrosauridae

Taxonomy

Pararhabdodon isonense was named by Casanovas Cladellas (1992) [A nomen vanum in this paper.]. Its type specimen is IPS-SRA 1, a set of vertebrae (mid-cervical vertebra), and it is a 3D body fossil. Its type locality is Sant Romà d'Abella (SRA), which is in a Maastrichtian crevasse splay sandstone in the Talarn Formation of Spain.

It was corrected as Pararhabdodon isonensis by Le Loeuff (1998), Casanovas-Cladellas et al. (1998), Casanovas et al. (1999), Sanz (2000), Prieto-Márquez et al. (2006), Lund and Gates (2006), Dalla Vecchia (2006), Suñer et al. (2008), Riera et al. (2009), Dalla Vecchia (2009), Pereda-Suberbiola et al. (2009), Prieto-Márquez and Wagner (2010), Prieto-Márquez (2010), Cruzado-Caballero et al. (2010), Dalla Vecchia (2010), Prieto-Márquez (2010), Godefroit and Motchurova-Dekova (2010), Prieto-Marquez and Salinas (2010), Prieto-Márquez (2011), Galobart et al. (2011), Prieto-Márquez (2011), Godefroit et al. (2012), Ramírez-Velasco et al. (2012), Cruzado-Caballero et al. (2013), Fondevilla et al. (2013), Sellés et al. (2014), Bolotsky et al. (2014), Dalla Vecchia (2014), Dalla Vecchia et al. (2014), Company et al. (2015), Blanco et al. (2015), Cruzado-Caballero and Powell (2017), Fondevilla et al. (2018), Párraga and Prieto-Márquez (2019), Dalla Vecchia (2020) and Sellés et al. (2021).

Synonyms

Koutalisaurus kohlerorum was named by Prieto-Márquez et al. (2006). Its type specimen is IPS SRA 27, a mandible, and it is not a trace fossil. Its type locality is Les Llaus (LL), which is in a Maastrichtian coarse channel fill sandstone/sandstone in the Conques Formation of Spain. It is the type species of Koutalisaurus.

It was synonymized subjectively with Pararhabdodon isonensis by Riera et al. (2009), Dalla Vecchia (2009), Prieto-Márquez and Wagner (2010), Prieto-Márquez (2010), Dalla Vecchia (2010), Godefroit and Motchurova-Dekova (2010), Godefroit et al. (2012) and Dalla Vecchia et al. (2014); it was considered a nomen dubium by Cruzado-Caballero et al. (2010) and Fondevilla et al. (2018).

Synonymy list

Year	Name and author
1992	Pararhabdodon isonense Casanovas Cladellas p. 148 figs. Pl. 1
1993	Pararhabdodon isonense Casanovas-Cladellas et al. p. 122
1995	Pararhabdodon isonense Casanovas-Cladellas et al. p. 278
1998	Pararhabdodon isonensis Casanovas-Cladellas et al. p. 16
1998	Pararhabdodon isonensis Le Loeuff p. 56
1999	Pararhabdodon isonensis Casanovas et al. p. 353
2000	Pararhabdodon isonensis Sanz p. 297
2006	Pararhabdodon isonensis Dalla Vecchia
2006	Pararhabdodon isonensis Lund and Gates p. 273
2006	Pararhabdodon isonensis Prieto-Márquez et al. p. 931
2006	Koutalisaurus kohlerorum Prieto-Márquez et al. p. 936 figs. 7-8
2008	Koutalisaurus kohlerorum Suñer et al. p. 404
2008	Pararhabdodon isonensis Suñer et al. p. 404
2009	Pararhabdodon isonensis Dalla Vecchia p. 13
2009	Koutalisaurus kohlerorum Pereda-Suberbiola et al. p. 560
2009	Pararhabdodon isonensis Pereda-Suberbiola et al. p. 560
2009	Pararhabdodon isonensis Riera et al. p. 168
2010	Pararhabdodon isonensis Cruzado-Caballero et al. p. 154
2010	Pararhabdodon isonensis Dalla Vecchia p. 31
2010	Pararhabdodon isonensis Godefroit and Motchurova-Dekova p. 168
2010	Pararhabdodon isonense Juárez Valieri et al. p. 223
2010	Pararhabdodon isonensis Prieto-Marquez and Salinas p. 835 fig. 20
2010	Pararhabdodon isonensis Prieto-Márquez p. 5 fig. 2
2010	Pararhabdodon isonensis Prieto-Márquez and Wagner p. 1240
2011	Pararhabdodon isonensis Galobart et al. p. 18
2011	Pararhabdodon isonensis Prieto-Márquez p. 64
2012	Pararhabdodon isonensis Godefroit et al. p. 535
2012	Pararhabdodon isonensis Ramírez-Velasco et al. p. 392
2013	Pararhabdodon isonensis Cruzado-Caballero et al.
2013	Pararhabdodon isonensis Fondevilla et al. p. 67
2014	Pararhabdodon isonensis Bolotsky et al. p. 327–328
2014	Pararhabdodon isonensis Dalla Vecchia p. 270
2014	Pararhabdodon isonensis Dalla Vecchia et al. p. 304
2014	Pararhabdodon isonensis Sellés et al. p. 725
2015	Pararhabdodon isonensis Blanco et al. p. 447
2015	Pararhabdodon isonensis Company et al. p. 72
2017	Pararhabdodon isonensis Cruzado-Caballero and Powell p. 3
2018	Pararhabdodon isonensis Fondevilla et al. p. 2
2019	Pararhabdodon isonensis Párraga and Prieto-Márquez p. 253
2020	Pararhabdodon isonensis Dalla Vecchia p. 599
2021	Pararhabdodon isonensis Sellés et al. p. 3

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda

Rank	Name	Author
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Ornithischia	()
—	Genasauria
—	Cerapoda
—	Ornithopoda	()
—	Clypeodonta
—	Iguanodontia	()
—	Dryomorpha
—	Styracosterna
—	Hadrosauriformes
—	Hadrosauroidea	()
family	Hadrosauridae
subfamily	Lambeosaurinae
—	Arenysaurini
genus	Pararhabdodon
species	isonensis	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Pararhabdodon isonensis Casanovas Cladellas 1992

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Invalid names: Koutalisaurus kohlerorum Prieto-Márquez et al. 2006 [synonym]

Diagnosis

Reference	Diagnosis
A. Prieto-Márquez and J. R. Wagner 2010	Differs from all other hadrosaurids, except Tsintaosaurus spinorhinus, in elevation of jugal facet of maxilla such that ventralmost extent is well above level of lateral margin of ectopterygoid shelf, maxilla forms acute embayment extending ventral to jugal process between jugal facet and ectopterygoid shelf; jugal facet of maxilla anteroposteriorly foreshortened, likely with correspondingly anteroposteriorly narrow anterior jugal; anterior dentary with symphysial process projecting medially such that distance between symphysis and lateral surface of dentary is three times labiolingual thickness of alveolar chamber. Differs from Tsintaosaurus spinorhinus in broader, subrectangular anterodorsal region of the maxilla.

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

gregariouso

Environment:

terrestrialo

Locomotion:

actively mobileo

Life habit:

ground dwellingo

Diet:

herbivoresubo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2009-01-03 20:40:05

Modified:

2009-01-03 22:40:05

Source: subo = suborder, o = order

References: Benton 1983, Marsh 1875

Age range: Late/Upper Maastrichtian or 70.60000 to 66.00000 Ma

Collections (6 total)

Time interval	Ma	Country or state	Original ID and collection number
Early/Lower Maastrichtian	70.6 - 66.0	Spain (Cataluña)	Hadrosauridae indet. (60474) Pararhabdodon isonensis (171099)
Late/Upper Maastrichtian	70.6 - 66.0	Spain (Cataluña)	Hadrosauridae indet. (148953) Pararhabdodon isonense (171102) Pararhabdodon isonensis (67138) Rhabdodon sp. (type locality: 38893)