PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Anisoceratidae

Cephalopoda - Ammonitida - Anisoceratidae

Taxonomy

Anisoceratidae was named by Hyatt (1900). Its type is Anisoceras.

It was assigned to Hamitaceae by Wright (1952); to Ancylocerataceae by Förster (1975); to Turrilitaceae by Cantu-Chapa (1962), Wright (1979), Kennedy (1984), Kennedy et al. (1988), Kennedy and Cobban (1990), Kennedy (1994), Kennedy (1995), Kennedy et al. (1998), Arkadiev et al. (2000), Szives (2007) and Kennedy and Latil (2007); and to Turrilitoidea by Henderson (1990), Kennedy (2005), Kennedy et al. (2008), Kennedy et al. (2008), Kennedy and Klinger (2009), Klinger et al. (2010), Kennedy and Gale (2015), Sharifi et al. (2016) and Kennedy and Morris (2018).

Subtaxa

Algerites, Allocrioceras, Anisoceras (type genus), Idiohamites, Metahamites, Ndumuiceras, Phlycticrioceras, Prohelicoceras, Protanisoceras, Rossalites, Tarrantites

Synonyms

Algeritidae was named by Spath (1925).

It was synonymized subjectively with Anisoceratidae by Kennedy and Cobban (1990) and Kennedy (1995).

It was assigned to Hamitaceae by Wright (1952).
Phlycticrioceratidae was named by Spath (1926). It is not extant.

It was synonymized subjectively with Anisoceratidae by Kennedy (1984) and Kennedy (1995).

It was assigned to Hamitaceae by Wright (1952); to Ancylocerataceae by Summesberger (1979); and to Ammonoidea by Thomel (1988).

Synonymy list

Year	Name and author
1900	Anisoceratidae Hyatt p. 587
1925	Algeritidae Spath p. 190
1926	Phlycticrioceratidae Spath
1952	Algeritidae Wright
1952	Anisoceratidae Wright
1952	Phlycticrioceratidae Wright
1962	Anisoceratidae Cantu-Chapa p. 31
1975	Anisoceratidae Förster p. 169
1979	Phlycticrioceratidae Summesberger p. 117
1979	Anisoceratidae Wright p. 288
1984	Anisoceratidae Kennedy p. 134
1988	Anisoceratidae Kennedy et al. p. 41
1988	Phlycticrioceratidae Thomel p. 19
1990	Anisoceratidae Henderson p. 137
1990	Anisoceratidae Kennedy and Cobban p. 416
1994	Anisoceratidae Kennedy p. 231
1995	Anisoceratidae Kennedy p. 427
1998	Anisoceratidae Kennedy et al. p. 35
2000	Anisoceratidae Arkadiev et al. p. 115
2005	Anisoceratidae Kennedy p. 406
2007	Anisoceratidae Kennedy and Latil p. 467
2007	Anisoceratidae Szives p. 59
2008	Anisoceratidae Kennedy et al. p. 45
2008	Anisoceratidae Kennedy et al. p. 131
2009	Anisoceratidae Kennedy and Klinger p. 43
2010	Anisoceratidae Klinger et al. p. 91
2015	Anisoceratidae Kennedy and Gale p. 516
2016	Anisoceratidae Sharifi et al. p. 604
2018	Anisoceratidae Kennedy and Morris p. 95

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
phylum	Mollusca

Rank	Name	Author
class	Cephalopoda	Cuvier 1797
subclass	Ammonoidea	()
order	Ammonitida
suborder	Ancyloceratina	Wiedmann 1966
superfamily	Turrilitoidea	(Gill 1871)
family	Anisoceratidae
family	Anisoceratidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Anisoceratidae Hyatt 1900

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G. †Algerites

†Algerites ellipticus Mantell 1822

†Algerites sayni Pervinquière 1910

G. †Allocrioceras Spath 1926

†Allocrioceras angustum Sowerby 1850

Invalid names: Allocrioceras woodsi Spath 1926 [synonym], Hamites geinitzii d'Orbigny 1850 [synonym]

†Allocrioceras billinghursti Klinger 1976

†Allocrioceras burckhardti Kennedy and Cobban 1991

†Allocrioceras cuvieri Schluter 1872

†Allocrioceras dentonense Moreman 1942

†Allocrioceras larvatum Conrad 1855

†Allocrioceras nodiger Roemer 1870

Invalid names: Allocrioceras strangulatum Wright 1979 [synonym]

G. †Anisoceras Pictet 1854

Subg. †Anisoceras (Anisoceras) Pictet 1854

†Anisoceras armatum Sowerby 1817

Invalid names: Anisoceras jacobi Breistroffer 1946 [synonym]

†Anisoceras arrogans Giebel 1852

Invalid names: Anisoceras campichei Spath 1926 [synonym]

†Anisoceras charlottense Anderson 1958

†Anisoceras draconis Anderson 1958

†Anisoceras exoticum Spath 1939

†Anisoceras haasi Cooper and Kennedy 1979

†Anisoceras oldhamianum Stoliczka 1861

†Anisoceras perarmatum Pictet and Campiche 1861

Invalid names: Anisoceras pseudopunctatum Pictet and Campiche 1861 [synonym], Anisoceras vraconense Renz 1968 [synonym]

†Anisoceras phillipsi Cooper and Kennedy 1979

†Anisoceras plicatilis Sowerby 1819

†Anisoceras pseudoelegans Pictet and Campiche 1861

†Anisoceras raynaudi Boule et al. 1907

†Anisoceras reidi Wright 1979

†Anisoceras saussureanum spinosa Haas 1942

†Anisoceras saussureanus Pictet 1847

†Anisoceras spathi Wiedmann 1962

†Anisoceras subarcuatum Spath 1939

†Anisoceras subundulatum Yokoyama 1890

G. †Idiohamites Spath 1925

†Idiohamites bispinosus Kennedy and Cobban 1990

†Idiohamites elegantulus Spath 1939

†Idiohamites fremonti Marcou 1858

†Idiohamites pulchellus Kennedy and Cobban 1990

†Idiohamites pygmaeus Cooper and Kennedy 1979

†Idiohamites spiniger Sowerby 1818

†Idiohamites subspiniger Spath 1939

†Idiohamites tuberculatus Sowerby 1818

G. †Metahamites

G. †Ndumuiceras Kennedy and Klinger 2009

†Ndumuiceras variabile Kennedy and Klinger 2009

G. †Phlycticrioceras

†Phlycticrioceras rude Kennedy 1995

†Phlycticrioceras trinodosus Geinitz 1850

Invalid names: Ancyloceras douvillei de Grossouvre 1894 [synonym], Phlycticrioceras oregonense Reeside 1927 [synonym]

G. †Prohelicoceras

G. †Protanisoceras

Subg. †Protanisoceras (Heteroclinus) Casey 1961

Subg. †Protanisoceras (Protanisoceras) Spath 1923

†Protanisoceras (Protanisoceras) creutzbergi Kakabadze and Hoedemaeker 1997

Subg. †Protanisoceras (Torquistylus) Casey 1961

†Protanisoceras acteon d'Orbigny 1850

†Protanisoceras blancheti Pictet and Campiche 1861

G. †Rossalites

G. †Tarrantites Klinger et al. 2010

†Tarrantites adkinsi Scott 1928

Invalid names: Algeritidae Spath 1925 [synonym], Phlycticrioceratidae Spath 1926 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

aragoniteo

Environment:

marinec

Locomotion:

fast-movingo

Life habit:

nektonico

Diet:

carnivoreo

Vision:

well-developedc

Created:

2005-01-28 02:27:03

Modified:

2009-10-08 14:14:55

Source: o = order, c = class

References: Kiessling 2004, Kiessling 2003

Age range: base of the Late/Upper Aptian to the top of the Santonian or 122.46000 to 83.60000 Ma

Collections (220 total)

Time interval	Ma	Country or state	Original ID and collection number
Aptian - Albian	125.0 - 100.5	Azerbaijan	Ancyloceras blancheti (209384)
Late/Upper Aptian	122.46 - 113.0	Madagascar	Metahamites parcetuberculatus (83997)
Late/Upper Aptian	122.46 - 113.0	France	Protanisoceras sp. (56673)
Late/Upper Aptian	122.46 - 113.0	South Africa	Protanisoceras sp. (65971)
Albian	113.0 - 100.5	France	Ancyloceras blancheti (2166)
Albian	113.0 - 100.5	Canada (British Columbia)	Anisoceras sp. (87506)
Albian	113.0 - 100.5	Colombia	Protanisoceras creutzbergi (71083)
Albian	113.0 - 100.5	Madagascar	Metahamites dubius, Protanisoceras flexuosum (98268) Metahamites gignouxi, Protanisoceras hourcqi, Protanisoceras komihevitraense (98244) Metahamites octosignatus, Metahamites problematicus (98264) Metahamites spathi, Metahamites gignouxi (98227) Protanisoceras mareranense (98245) Protanisoceras tuberculatum, Protanisoceras gradatum, Protanisoceras crassecostatum, Protanisoceras hourcqi (98234)
Albian	113.0 - 100.5	Japan	Anisoceras sp. (122346)
Albian	113.0 - 100.5	Switzerland	Ancyloceras blancheti (2167) Anisoceras perarmatus, Anisoceras alternatus, Anisoceras pseudopunctatus, Anisoceras saussureanus, Anisoceras pseudoelegans (2237)
Albian	113.0 - 100.5	Madagascar (Mahajanga)	Anisoceras trituberculatum, Metahamites gignouxi, Metahamites spathi, Metahamites sablieri, Metahamites besairiei, Protanisoceras breistrofferi, Protanisoceras hourcqi, Protanisoceras superbum, Protanisoceras besairiei (86497)
Albian	113.0 - 100.5	USA (Oregon)	Anisoceras merriami (219049 229463)
Early/Lower Albian	112.03 - 109.0	South Africa	Ndumuiceras variabile (177185) Protanisoceras sp. (66073)
Early/Lower Albian	112.03 - 109.0	Peru (Cajamarca)	Protanisoceras blancheti (88170)
Early/Lower Albian	112.03 - 109.0	Madagascar	Protanisoceras convexum (98233)
Early/Lower Albian	112.03 - 109.0	United Kingdom (England)	Metahamites sp., Prohelicoceras anglicum, Protanisoceras raulinianum, Protanisoceras cantianum, Protanisoceras lardyi, Protanisoceras vaucherianum, Protanisoceras halleri, Protanisoceras sp., Protanisoceras blancheti, Protanisoceras acteon (1783) Protanisoceras acteon (1863) Protanisoceras cantianum, Protanisoceras acteon (1834) Protanisoceras cantianum, Protanisoceras vaucherianum, Protanisoceras sp., Protanisoceras blancheti, Protanisoceras acteon (1827) Protanisoceras major (77949) Protanisoceras raulinianum, Protanisoceras acteon (1846) Protanisoceras raulinianum, Protanisoceras cantianum (1818) Protanisoceras sp. (1833)
Early/Lower Albian	112.03 - 109.0	Switzerland	Protanisoceras cantianum (197617)
Early/Lower Albian	112.03 - 109.0	Peru	Protanisoceras blancheti (88173)
Early/Lower Albian	112.03 - 109.0	France	Protanisoceras raulinianum (56521)
Early/Lower Albian	112.03 - 109.0	Pakistan	Tarrantites adkinsi, Anisoceras arrogans (177362)
Early/Lower Albian - Middle Albian	112.03 - 105.3	France	Anisoceras pararmatum, Anisoceras elegantulus, Anisoceras armatum, Anisoceras pseudoelegans (13499) Protanisoceras blancheti (58124) Protanisoceras buvigneri, Protanisoceras alternotuberculatum, Protanisoceras raulinianum, Protanisoceras sp. (56845)
Early/Lower Albian - Late/Upper Albian	112.03 - 99.6	France	Anisoceras sp., Idiohamites favrinus, Protanisoceras lardyi, Protanisoceras sp., Protanisoceras blancheti (58861) Anisoceras sp., Protanisoceras sp. (57922)
Middle Albian	109.0 - 105.3	South Africa	Ndumuiceras variabile (177187)
Middle Albian	109.0 - 105.3	France	Anisoceras arrogans (56550) Metahamites arrogans, Protanisoceras buvignieri (233805) Protanisoceras blancheti, Protanisoceras raulinianum, Protanisoceras lardyi (233804) Protanisoceras raulinianum (56540) Protanisoceras sp. (56529 58430 58933)
Middle Albian	109.0 - 105.3	Angola	Protanisoceras sp. (211028)
Middle Albian	109.0 - 105.3	Madagascar (Mahajanga)	Idiohamites androiavensis, Anisoceras armatum, Anisoceras exoticum (86503)
Middle Albian	109.0 - 105.3	USA (Texas)	Tarrantites adkinsi (177360 177361)
Middle Albian - Late/Upper Albian	109.0 - 99.6	Madagascar (Mahajanga)	Metahamites gignouxi (86506)
Middle Albian - Late/Upper Albian	109.0 - 99.6	Switzerland	Anisoceras pseudelegans, Anisoceras armatum, Anisoceras perarmatum (197618)
Late/Upper Albian	105.3 - 99.6	Nigeria	Anisoceras perarmatum (82735) Anisoceras spp., Anisoceras perarmatum (210106)
Late/Upper Albian	105.3 - 99.6	USA (California)	Anisoceras sp. (1784) Idiohamites sp. (225652 225674)
Late/Upper Albian	105.3 - 99.6	Ukraine	Anisoceras perarmatum (183305)
Late/Upper Albian	105.3 - 99.6	South Africa	Anisoceras sp., Idiohamites sp. (82478)
Late/Upper Albian	105.3 - 99.6	India	Anisoceras perarmatum (201887 201889)
Late/Upper Albian	105.3 - 99.6	Australia (Northern Territory)	Anisoceras sp. (70283) Anisoceras sp., Idiohamites subspiniger, Idiohamites tuberculatus (70245)
Late/Upper Albian	105.3 - 99.6	United Kingdom (England)	Anisoceras exoticum (82856) Anisoceras sp., Anisoceras perarmatum (82808)
Late/Upper Albian	105.3 - 99.6	Italy	Anisoceras perarmatum (12838)
Late/Upper Albian	105.3 - 99.6	Angola	Anisoceras armatum, Anisoceras perarmatum, Anisoceras haasi, Anisoceras phillipsi, Anisoceras arrogans, Anisoceras exoticum, Anisoceras subarcuatum, Anisoceras spathi (232170) Anisoceras armatum, Anisoceras subundulatum (210888) Anisoceras oldhamianum (210847) Anisoceras saussureanum, Idiohamites subspinigero (211248) Anisoceras sp., Idiohamites sp., Anisoceras armatum, Anisoceras perarmatum (83840) Idiohamites elegantulus, Anisoceras perarmatum, Anisoceras armatum, Anisoceras haasi, Idiohamites pygmaeus (232171) Idiohamites spiniger, Idiohamites sp. (211249)
Late/Upper Albian	105.3 - 99.6	Switzerland	Idiohamites sp. (58113)
Late/Upper Albian	105.3 - 99.6	Madagascar (Antsiranana)	Anisoceras armatum, Idiohamites spiniger (98215) Anisoceras saussurei, Anisoceras diegoensis, Anisoceras perarmatum (86507)
Late/Upper Albian	105.3 - 99.6	Spain	Anisoceras armatum (3824) Anisoceras perarmatum (3829)
Late/Upper Albian	105.3 - 99.6	Armenia	Anisoceras perarmatum (183304)
Late/Upper Albian	105.3 - 99.6	Madagascar (Mahajanga)	Anisoceras saussurei (98220) Protanisoceras actaeon, Protanisoceras raulianum, Protanisoceras madagasyense, Protanisoceras blancheti (98221)
Late/Upper Albian	105.3 - 99.6	Belgium	Anisoceras armatum (174924)
Late/Upper Albian	105.3 - 99.6	Japan	Anisoceras sp. (122277)
Late/Upper Albian	105.3 - 99.6	Mozambique	Anisoceras raynaudi (174811) Anisoceras sp. (174827)
Late/Upper Albian	105.3 - 99.6	USA (Texas)	Anisoceras armatum (82371) Anisoceras perarmatum (82364 82366) Anisoceras sp. A (175118)
Late/Upper Albian	105.3 - 99.6	Iran	Idiohamites fremonti (183699)
Late/Upper Albian	105.3 - 99.6	France	Anisoceras armatum (58426 58533 174469) Anisoceras armatum, Anisoceras perarmatum (174468 174482) Anisoceras armatum, Anisoceras perarmatum, Idiohamites elegantulus (174484) Anisoceras armatum, Idiohamites elegantulus (174480) Anisoceras perarmatum (174466 174467 174481 174487) Anisoceras perarmatum, Anisoceras pseudoelegans, Idiohamites elegantulus (174485) Anisoceras perarmatum, Idiohamites elegantulus (174477 174486) Anisoceras sp. (56678 58122 58359 58364 58560) Anisoceras sp., Anisoceras armatum, Anisoceras perarmatum (58527) Anisoceras sp., Anisoceras auberti, Idiohamites sp., Anisoceras armatum, Anisoceras perarmatum, Anisoceras saussureanum (56680) Anisoceras sp., Anisoceras perarmatum (58363) Anisoceras sp., Idiohamites sp. (56674 58126) Anisoceras sp., Idiohamites sp., Anisoceras perarmatum (56669 58431) Anisoceras sp., Idiohamites sp., Anisoceras perarmatum, Anisoceras saussureanum (56523) Anisoceras sp., Idiohamites tuberculatus (58125) Idiohamites elegantulus (174471 174472 174474 174475 174479) Idiohamites sp. (56552 56554 57941)
Late/Upper Albian - Cenomanian	105.3 - 93.9	Madagascar	Anisoceras subcompressum, Anisoceras armatum (98219)
Cenomanian	100.5 - 93.9	USA (Arizona)	Allocrioceras sp. (3082 3083)
Cenomanian	100.5 - 93.9	USA (Utah)	Allocrioceras annulatum (5608)
Early/Lower Cenomanian - Late/Upper Albian	99.6 - 99.6	Spain	Anisoceras perarmatum (3832)
Early/Lower Cenomanian	99.6 - 93.5	USA (Texas)	Anisoceras sp. B (175263)
Early/Lower Cenomanian	99.6 - 93.5	Tunisia	Anisoceras leckenbyi, Anisoceras plicatile (178213) Idiohamites collignoni, Algerites ellipticus (217133)
Early/Lower Cenomanian	99.6 - 93.5	France	Algerites ellipticus (13500)
Early/Lower Cenomanian	99.6 - 93.5	France (Normandie)	Anisoceras sp. nov., Anisoceras plicatile (230776)
Early/Lower Cenomanian	99.6 - 93.5	Mexico (Zacatecas)	Anisoceras neohispanicum, Anisoceras armatum (87607)
Early/Lower Cenomanian	99.6 - 93.5	United Kingdom (England)	Anisoceras auberti, Anisoceras sp., Idiohamites alternatus, Idiohamites collignoni, Idiohamites ellipticus, Idiohamites sp. (82798) Anisoceras sp. (82814)
Early/Lower Cenomanian	99.6 - 93.5	Japan	Anisoceras sp. (123126)
Early/Lower Cenomanian	99.6 - 93.5	USA (California)	Anisoceras sp. (1786 1788)
Early/Lower Cenomanian	99.6 - 93.5	Tanzania	Algerites sayni (193269)
Early/Lower Cenomanian	99.6 - 93.5	Kazakhstan	Algerites ellipticus (175009)
Early/Lower Cenomanian - Middle Cenomanian	99.6 - 93.5	France	Anisoceras saussurei (230858) Anisoceras sp. (58626)
Late/Upper Cenomanian	99.6 - 93.5	USA (Wyoming)	Idiohamites bispinosus (167808 167809 167810)
Late/Upper Cenomanian	99.6 - 93.5	France	Protanisoceras sp. (58969)
Late/Upper Cenomanian	99.6 - 93.5	USA (Arizona)	Allocrioceras annulatum (53495) Allocrioceras sp. (55564) Anisoceras coloradoense (55562 55563 55817)
Late/Upper Cenomanian	99.6 - 93.5	USA (Montana)	Allocrioceras annulatum (175421) Idiohamites bispinosus (175423)
Late/Upper Cenomanian	99.6 - 93.5	USA (New Mexico)	Allocrioceras annulatum (175425) Allocrioceras sp. (44282) Anisoceras coloradoense (229802)
Late/Upper Cenomanian	99.6 - 93.5	United Kingdom (England)	Anisoceras plicatile (82857 82899 82913) Idiohamites alternatus (82918)
Late/Upper Cenomanian	99.6 - 93.5	USA (Colorado)	Anisoceras plicatile (78164) Anisoceras sp. (177116)
Late/Upper Cenomanian	99.6 - 93.5	USA (Texas)	Allocrioceras annulatum (175717) Allocrioceras conlini (230848)
Late/Upper Cenomanian	99.6 - 93.5	USA (Utah)	Allocrioceras sp. (3289 3326)
Middle Cenomanian	99.6 - 93.5	France	Anisoceras plicatile (82843)
Middle Cenomanian	99.6 - 93.5	USA (Montana)	Idiohamites pulchellus, Idiohamites bispinosus (175422)
Middle Cenomanian	99.6 - 93.5	USA (New Mexico)	Anisoceras plicatile (178790)
Middle Cenomanian	99.6 - 93.5	Germany (Westphalia)	Anisoceras sp., Idiohamites alternatus, Anisoceras plicatile (4067)
Turonian - Coniacian	93.9 - 86.3	USA (Wyoming)	Anisoceras sp. (87757)
Early/Lower Turonian	93.5 - 89.3	Canada (British Columbia)	Allocrioceras sp. (210519)
Early/Lower Turonian	93.5 - 89.3	USA (Colorado)	Anisoceras sp., Allocrioceras larvatum (230901)
Early/Lower Turonian	93.5 - 89.3	USA (Texas)	Allocrioceras larvatum (4898) Allocrioceras larvatum, Allocrioceras dentonense (80991)
Late/Upper Turonian	93.5 - 89.3	Germany	Allocrioceras angustum (12681 12682 12684 12685 12693 12704) Allocrioceras billinghursti (154797) Allocrioceras billinghursti, Allocrioceras strangulatum (74196) Anisoceras sp. (12686)
Late/Upper Turonian	93.5 - 89.3	France	Allocrioceras billinghursti (175633) Allocrioceras nodiger (175639 175640)
Late/Upper Turonian	93.5 - 89.3	United Kingdom (England)	Allocrioceras billinghursti (175763 175772 175773) Allocrioceras billinghursti, Allocrioceras strangulatum, Anisoceras reidi, Allocrioceras angustum (112693) Allocrioceras cuvieri (175774) Allocrioceras sp., Allocrioceras ellipticum (175792) Allocrioceras strangulatum (175771) Allocrioceras strangulatum, Allocrioceras angustum (175764) Anisoceras reidi (175770)
Late/Upper Turonian	93.5 - 89.3	Ukraine	Allocrioceras strangulatum (183311)
Middle Turonian - Late/Upper Turonian	93.5 - 89.3	Germany	Allocrioceras angustum (12680)
Coniacian	89.8 - 86.3	USA (New Mexico)	Phlycticrioceras oreonense (82251)
Coniacian	89.8 - 86.3	USA (Wyoming)	Anisoceras sp. (87769) Anisoceras sp., Phlycticrioceras oregonense (87761) Phlycticrioceras oregonense (87763)
Coniacian	89.8 - 86.3	USA (California)	Anisoceras draconis (44270)
Early/Lower Coniacian	89.3 - 85.8	USA (New Mexico)	Phlycticrioceras oregonense (87847)
Late/Upper Coniacian	89.3 - 85.8	France	Phlycticrioceras trinodosus (179102)
Middle Coniacian	89.3 - 85.8	South Africa	Allocrioceras sp. (82443)
Middle Coniacian	89.3 - 85.8	USA (Colorado)	Phlycticrioceras oregonense (87851)
Middle Coniacian	89.3 - 85.8	USA (Wyoming)	Phlycticrioceras oregonense (87760)
Late/Upper Coniacian - Early/Lower Santonian	89.3 - 83.5	France	Phlycticrioceras trinodosus (69871)
Santonian	86.3 - 83.6	USA (California)	Allocrioceras sp. (215307)
Late/Upper Santonian	85.8 - 83.5	France	Phlycticrioceras rude (179119 179761)
Middle Santonian	85.8 - 83.5	France	Allocrioceras burkhardti (179609)
Middle Santonian	85.8 - 83.5	USA (New Mexico)	Phlycticrioceras oregonense (87848)