Classification of Krell 2000

Superfm. Scarabaeoidea Latreille 1802 [scarab beetle]
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G. †Aliscarabaeus Hong 1984
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G. †Avitortor Ponomarenko 1977
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†Avitortor leptoscelis Nikritin 1977
Fm. Bolboceratidae Mulsant 1842 [dor beetle]
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G. Bolboceras Kirby 1819
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†Bolboceras inermis Piton 1940
G. †Cretobolbus Nikolajev 1996
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†Cretobolbus rohdendorfi Nikolajev 1996
G. †Cretoserica Nikolajev 1998
Fm. Geotrupidae Latreille 1802 [earth-boring dung beetle]
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Subfm. †Cretogeotrupinae Nikolajev 1996
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G. †Cretogeotrupes Nikolajev 1992
Subfm. Geotrupinae Latreille 1802
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G. Geotrupes Latreille 1796
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†Geotrupes atavus Oustalet 1874
Invalid names: Obitiscarabaeus Hong 1984 [synonym]
G. †Geotrupoides Handlirsch 1906
Fm. Glaphyridae MacLeay 1819 [bumble bee scarab beetle]
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G. Amphicoma Latreille 1807
G. Glaphyrus Latreille 1802
Fm. Glaresidae Prudhomme de Borre 1886 [enigmatic scarab beetle]
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Invalid names: Aphodiites Heer 1864 [nomen dubium]
G. †Hongscarabaeus Krell 2000
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Invalid names: Proscarabaeus Hong 1982 [replaced]
Fm. Hybosoridae Erichson 1847 [scavenger scarab beetle]
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Subfm. Anaidinae Nikolajev 1996
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G. †Cretanaides Nikolajev 1996
G. †Coprologus Heer 1847
Subfm. Hybosorinae Erichson 1847
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G. Phaeochrous Laporte 1840
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†Phaeochrous tertiarium Deichmüller 1881
G. †Hybosorites Nikolajev 1996
Fm. Lucanidae Latreille 1804 [stag beetle]
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Subfm. Aesalinae MacLeay 1819
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G. †Cretaesalus Nikolajev 1993
G. †Ceruchites Statz 1952
G. †Dorcasoides Motschulsky 1856
Subfm. Lucaninae Latreille 1804
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G. †Miocenidorcus Riou 1999
G. Platycerus Geoffroy 1762
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†Platycerus zherichini Nikolayev 1990
G. †Succiniplatycerus Nikolayev 1990
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Invalid names: Lucanus fossilis Wickham 1913 [nomen dubium], Platycerus sepultus Germar 1837 [nomen dubium]
G. †Paleognathus Waga 1883
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†Paleognathus succini Waga 1883
Subfm. Syndesinae MacLeay 1819
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G. Ceruchus MacLeay 1819
G. †Mesoscarabaeus Hong 1982
Fm. Ochodaeidae Streubel 1846 [sand-loving scarab beetle]
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Subfm. †Cretochodaeinae Nikolajev 1995
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G. †Cretochodaeus Nikolajev 1995
G. †Mioochodaeus Nikolajev 1995
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†Mioochodaeus proaevus Germar 1849
G. †Opiselleipon Bode 1953
Fm. Passalidae Leach 1815 [bess beetle]
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G. Passalus Fabricius 1792
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Subg. Passalus (Passalus) Fabricius 1792
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G. †Serrulus Hong 1984
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†Serrulus sinicus Hong 1984
Fm. Pleocomidae Leconte 1861 [rain beetle]
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G. †Proteroscarabaeus Grabau 1923
Fm. Scarabaeidae Latreille 1802 [scarab beetle]
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Subfm. Aegialiinae Laporte 1840
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G. Aegialia Latreille 1807
G. †Cretaegialia Nikolayev 1993
Subfm. Aphodiinae Leach 1815
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Tr. Aphodiini Leach 1815
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G. Aphodius Hellwig 1798
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†Aphodius anteactus Krell 2000
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Invalid names: Aphodius antiquus Heer 1847 [replaced]
†Aphodius theobaldi Krell 2000
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Invalid names: Aphodius incertus Theobald 1937 [replaced]
Tr. Eupariini Schmidt 1910
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G. Ataenius Harold 1867
G. Oxyomus Kirby and Spence 1828
G. Saprosites Redtenbacher 1858
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†Saprosites succini Zang 1905
Subfm. Cetoniinae Leach 1815 [flower chafer]
Subfm. †Cretoscarabaeinae Nikolajev 1995
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G. †Cretoscarabaeus Nikolajev 1995
Subfm. Dynastinae MacLeay 1819 [rhinoceras beetle]
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G. Ligyrus Burmeister 1847
G. Oryctes Hellwig 1798
G. Pentodon Hope 1837
G. Strategus Kirby 1828
Invalid names: Pentodon proserpinae Heer 1862 [nomen dubium]
Subfm. Euchirinae Hope 1840
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G. Cheirotonus Hope 1840
Subfm. Hopliinae Latreille 1829
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G. Lepitrix Lepeletier and Serville 1828
Subfm. †Lithoscarabaeinae Nikolajev 1992
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G. †Lithoscarabaeus Nikolajev 1992
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†Lithoscarabaeus baissensis Nikritin 1977
Subfm. Melolonthinae Leach 1819
Subfm. Rutelinae MacLeay 1819
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G. Adoretus Dejean 1833
G. Anomala Samouelle 1819
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†Anomala furva Hong 1984
†Anomala palaeobrunnea Krell 2000
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Invalid names: Magniscarabaeus brunneus Hong 1985 [replaced]
Invalid names: Magniscarabaeus Hong 1984 [synonym]
G. †Anomalites Frič 1885
G. Holotrichia Hope 1837
G. Mimela Kirby 1823
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†Mimela rhenana von Heyden 1862
G. †Miolachnosterna Wickham 1914
G. †Pelidnotites Cockerell 1920
G. †Petraeianus Zhang 1989
G. †Petulantis Zhang et al. 1994
G. †Zhangsunia Krell 2000
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†Zhangsunia extumida Zhang et al. 1994
Invalid names: Dolichopoda Zhang et al. 1994 [replaced]
Subfm. Scarabaeinae Latreille 1802
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G. Anachalcos Hope 1837
G. †Ateuchites Meunier 1898
G. Choeridium Lepeletier de Saint-Fargeau and Audinet-Serville 1828
G. Copris Geoffroy 1762
G. Gymnopleurus Illiger 1803
G. Heliocopris Hope 1837
G. Metacatharsius Paulian 1939
G. Oniticellus Dejean 1821
G. Onitis Fabricius 1798
G. Onthophagus Latreille 1802
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†Onthophagus urusheeri Krell 2000
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Invalid names: Onthophagus urus Heer 1847 [replaced]
G. Phanaeus Macleay 1819
G. †Prionocephale Lin 1980
G. †Scelocopris Zhang 1989
Subfm. Sericinae Kirby 1837
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G. †Lithanomala Nikolajev 1992
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†Lithanomala crassa Ponomarenko 1990
†Lithanomala oblonga Ponomarenko 1990
†Lithanomala sibirica Ponomarenko 1990
G. Maladera Mulsant 1871
G. †Mioserica Zhang 1989
G. Serica MacLeay 1819
Subfm. Trichiinae Fleming 1821
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G. Gnorimus Lepeletier and Serville 1828
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†Gnorimus aedilis Heer 1862
G. Trichius Fabricius 1775
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†Trichius lugubris Heer 1862
Subfm. Valginae Mulsant 1842
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G. Valgus Scriba 1790
Fm. Trogidae MacLeay 1819 [hide beetle]
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G. Trox Fabricius 1775
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†Trox antiquus Wickham 1909
Invalid names: Trox oustaleti Scudder 1879 [nomen dubium]
Invalid names: Holcorobeini Nikolajev 1992 [empty]
Fm. Melolonthinae Leach 1819
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G. Diplotaxis Kirby 1837
G. †Eophyllocerus Haupt 1950
G. Listrochelus Blanchard 1851
G. Macrodactylus Latreille 1827
G. Melolontha Fabricius 1775
G. †Melolonthites Heer 1847
G. Phyllophaga Harris 1826
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†Phyllophaga avus Cockerell 1921
G. Rhizotrogus Latreille 1825
Fm. Belostomatidae Leach 1815 [giant water bug]
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G. †Scarabaeides Germar 1839
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†Scarabaeides deperditus Germar 1839
Invalid names: Mesobelostomum Haase 1890 [objective synonym]
Fm. Cetonida Leach 1815 [flower chafer]
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G. †Genonota Zhang 1989
G. Macronota Hoffmansegg 1817