Classification of the family Leucosiidae

Fm. Leucosiidae Samouelle 1819
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Subfm. Cryptocneminae Stimpson 1907
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G. Cryptocnemus Stimpson 1858
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Subfm. Ebaliinae Stimpson 1871
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G. Arcania Leach 1817
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Arcania cornuta MacGilchrist 1905
Arcania elongata Yokoya 1933
Arcania globata Stimpson 1858
Arcania gracilis Henderson 1893
Arcania septemspinosa Fabricius 1787
†Arcania shuishuni Hu and Tao 1985
Arcania siamensis Rathbun 1909
Invalid names: Ixoides [synonym]
G. †Cabrillia Nyborg et al. 2018
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†Cabrillia hendyi Nyborg et al. 2018
G. Ebalia Leach 1817
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Ebalia cranchii Leach 1817
Ebalia crassipes Bell 1855
Ebalia cryptocnemoides Takeda and Miyake 1972
Ebalia dentifrons Miers 1886
†Ebalia deshayesi Lucas 1846
†Ebalia ferganensis Ilyin and Alekseev 2005
†Ebalia fucinii Ristori 1893
†Ebalia globulosa Müller 1976
†Ebalia hungarica Müller 1974
Ebalia intermedia Miers 1886
†Ebalia jacqueshermani van Bakel et al. 2003
†Ebalia kakegawaensis Karasawa 1993
†Ebalia lamarmorai Lorenthey 1909
Ebalia longimana Ortmann 1892
†Ebalia meulenkampi Muller and Georgiades 1984
†Ebalia multiangulata Muller 1993
†Ebalia noetlingi Karasawa 2008
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Invalid names: Ebalia tuberculata Noetling 1901 [replaced]
Ebalia nux Milne-Edwards 1883
†Ebalia oersi Müller 1978
Ebalia spinifera Miers 1886
†Ebalia spinosa Tiwari et al. 1997
Ebalia tuberculosa Milne-Edwards 1873
Ebalia tuberosa Pennant 1777 [Pennant's nut crab]
Ebalia tuberosus Pennat 1777
Ebalia tumefacta Montagu 1808 [Bryer's nut crab]
†Ebalia vahldieki Forster and Mundlos 1992
†Ebalia vanstraeleni Bachmayer 1953
G. Heteronucia Alcock 1896
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G. Hiplyra Galil 2009
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Hiplyra platycheir De Haan 1841
G. Ilia Leach 1817
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Ilia nucleus Linnaeus 1758
†Ilia ornata Randall 1839
G. Iliacantha Stimpson 1871
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Iliacantha globosa Stimpson 1981
†Iliacantha liodactylus Rathbun 1938
†Iliacantha panamanica Todd and Collins 2005
G. Ixa Leach 1816
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Ixa cylindrus Fabricius 1777
G. Leucosilia Bell 1855
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†Leucosilia bananensis Rathbun 1918
Leucosilia jurinei Saussure 1853
Leucosilia jurinii De Saussure 1853
G. †Leucosiraja De Angeli et al. 2017
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†Leucosiraja manta De Angeli et al. 2017
G. Lyphira Galil 2009
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Lyphira heterograna Ortmann 1892
G. Merocryptus Milne-Edwards 1873
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†Merocryptus altavillensis De Angeli et al. 2010
Merocryptus lambriformis Poore and Ahyong 2004
G. Myra Leach 1817
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Myra affinis Bell 1855
†Myra amoena Bohm 1922
†Myra arachnoides Bohm 1922
†Myra brevisulcata Morris and Collins 1991
Myra celeris Galil 2001
†Myra emarginata Glaessner 1928
†Myra fijiensis Rathbun 1934
Myra fugax Fabricius 1798 [fleeting purse crab]
Myra mammillaris Bell 1855
†Myra paenitensis Via 1941
†Myra subcarinata Morris and Collins 1991
†Myra trispinosa Morris and Collins 1991
†Myra wenjii Hu and Tao 1985
G. Myropsis Stimpson 1871
G. Nucia Dana 1852
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†Nucia baripadensis Bachmayer and Mohanti 1973
†Nucia borneoensis Morris and Collins 1991
†Nucia calculoides Morris and Collins 1991
†Nucia coxi Morris and Collins 1991
†Nucia fennemai Bohm 1922
†Nucia platyspinosa Collins et al. 2003
Nucia speciosa Dana 1852
G. Nursia Leach 1817
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†Nursia bilobata Karasawa et al. 2008
Nursia japonica Sakai 1935
G. Paranursia Serene and Soh 1976
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†Paranursia acharyai Bachmayer and Mohanti 1973
G. Persephona Leach 1817
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†Persephona bigranulata Rathbun 1926
†Persephona enigmatica Todd and Collins 2005
†Persephona invalida Rathbun 1929
†Persephona manningi Todd and Collins 2005
†Persephona niemeyeri Blow 2003
†Persephona prepunctata Rathbun 1919
Persephona punctata Linnaeus 1758
Persephona punctatus Linnaeus 1758
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Invalid names: Persephona latreillii Leach 1817 [synonym]
†Persephona rodesae Blow 2003
Invalid names: Guaia Leach 1817 [synonym]
G. Philyra Leach 1817
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†Philyra alveola Hu and Tao 2000
†Philyra collinsi Karasawa 2010
†Philyra cranium Desmarest 1822
†Philyra ferrica Hu and Tao 1996
†Philyra granulosa Morris and Collins 1991
†Philyra hayasakai Karasawa and Inoue 1992
†Philyra hormozganensis Vega et al. 2010
†Philyra miyamotoi Karasawa and Kishimoto 1996
†Philyra nishimotoi Karasawa 1989
Philyra olivacea Rathbun 1909
†Philyra plana Karasawa 1989
Philyra scabriuscula Fabricius 1793
†Philyra shihcheni Hu and Tao 1979
Philyra syndactyla Ortmann 1892
†Philyra tanakai Karasawa 1993
†Philyra tridentata Karasawa 1993
†Philyra trusanensis Collins et al. 2003
G. Pseudophilyra Miers
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†Pseudophilyra elongatella Hu and Tao 1996
G. Pyrhila Galil 2009
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Pyrhila pisum De Haan 1841
G. Randallia Stimpson 1857
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†Randallia jingomao Hu and Tao 2004
†Randallia ornata Randall 1839
†Randallia pleistocenica Rathbun 1926
†Randallia prolanata Hu and Tao 1996
†Randallia saitoensis Karasawa 1993
†Randallia strouhali Bachmayer 1953
G. Ryphila Galil 2009
G. Speloeophorus Milne Edwards 1865
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†Speloeophorus subcircularis Todd and Collins 2005
G. Tokoyo Galil 2003
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Tokoyo eburnea Alcock 1896
†Tokoyo trinucloidea Hu and Tao 1996
G. Uhlias Stimpson 1871
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Uhlias limbatus Stimpson 1871
Subfm. Leucosiinae Samouelle 1819
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G. Euclosiana Galil and Ng 2010
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†Euclosiana obtusifrons de Haan 1841
Invalid names: Euclosia Galil 2003 [replaced]
G. Leucosia Weber 1795
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Leucosia anatum Herbst 1783
†Leucosia calcarata Collins et al. 2003
Leucosia craniolaris Linnaeus 1758
†Leucosia longiangulata Morris and Collins 1991
†Leucosia martini Karasawa 2008
†Leucosia ovalata Hu and Tao 1996
Leucosia perlata De Haan 1841
†Leucosia persica Vega et al. 2010
Leucosia pubescens Miers 1877
†Leucosia serenei Morris and Collins 1991
†Leucosia subrhomboidalis Desmarest 1822
†Leucosia subrhomboidea Hu and Tao 1996
†Leucosia taiwanica Hu and Tao 1996
†Leucosia takamii Karasawa 1993
†Leucosia tricarinata Martin 1879
†Leucosia tutongensis Morris and Collins 1991
G. †Ristoria Garassino 2012
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†Ristoria pliocaenica Ristori 1891
G. Seulocia Galil 2005
G. Urnalana Galil 2005
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Urnalana haematosticta Adams and White 1848
Subfm. Philyrinae Leach 1817
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G. Bellidilia Kinahan 1856
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G. †Zannatoius Beschin et al. 2018
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†Zannatoius vicentinus Beschin et al. 2018