Classification of the family Trionychidae

Fm. Trionychidae Gray 1825 [softshell turtle]
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G. Apalone Rafinesque 1832 [softshell turtle]
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Subg. Apalone (Platypeltis) Fitzinger 1835
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Platypeltis ferox Schneider 1783 [Florida softshell turtle]
†Platypeltis sternbergi Wiman 1933
†Apalone amorense Valdes et al. 2017
Apalone ferox Schneider 1783 [Florida softshell turtle]
†Apalone lima Cope 1869
Apalone mutica LeSueur 1827 [smooth softshell turtle]
Apalone spinifera LeSueur 1827 [spiny softshell turtle]
†Apalone trionychoides Cope 1872
G. Aspideretes Hay 1904
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Aspideretes gangeticus Cuvier 1825 [Indian softshell]
Aspideretes hurum Gray 1831 [Indian peacock softshell]
Aspideretes leithii Gray 1872 [Leith's softshell]
Aspideretes nigricans Anderson 1875 [Black softshell]
†Aspideretes sinuosus Chow and Yeh 1958
G. Aspidonectes Wagler 1830
Subfm. †Chitrainae Jasinski et al. 2022
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G. Chitra Gray 1844
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Chitra chitra Nutaphand 1986
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Invalid names: Chitra selenkae Jaekel 1911 [replaced]
Chitra indica Gray 1831
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Invalid names: Trionyx indicus Gray 1831 [synonym]
Chitra vandijki McCord and Pritchard 2003 [Burmese narrow-headed softshell turtle]
G. Pelochelys Gray 1864
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Pelochelys bibroni Owen 1853 [Asian giant softshell]
G. †Conchochelys Hay 1905
G. †Crassithecachelys Chkhikvadze 2000
Subfm. Cyclanorbinae Hummel 1929
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G. Cyclanorbis Gray 1854
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Cyclanorbis senegalensis Dumeril and Bibron 1835
†Cyclanorbis turkanensis Meylan et al. 1990
G. Cycloderma Peters 1854
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Cycloderma aubryi Joyce and Lyson 2010
†Cycloderma debroinae Meylan et al. 1990
Cycloderma frenatum Peters 1854
†Cycloderma victoriae Andrews 1914
G. Lissemys Smith 1931
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Lissemys ceylonensis Gray 1856 [Sri Lankan flapshell turtle]
Lissemys punctata Bonnaterre 1789
Lissemys scutata Peters 1868 [Burmese flapshell turtle]
Invalid names: Emyda lineata Lydekker 1885 [nomen dubium], Emyda palaeindica Lydekker 1885 [nomen dubium], Emyda sivalensis Lydekker 1885 [nomen dubium], Lissemys piramensis Prasad 1974 [nomen dubium], Lissemys punctata sinhaleyus Deraniyagala 1953 [nomen dubium]
G. †Nemegtemys Danilov et al. 2014
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†Nemegtemys conflata Danilov et al. 2014
G. †Eugenechelys Chkhikvadze 2000
G. †Hummelichelys Chkhikvadze 2000
Subfm. †Kuhnemydinae Jasinski et al. 2022
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G. †Kuhnemys Chkhikvadze 1999
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†Kuhnemys breviplastra Danilov et al. 2014
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Invalid names: Aspideretes alashanensis Yeh 1965 [synonym], Trionyx alashanensis Yeh 1965 [synonym]
†Kuhnemys orlovi Khosatzky 1976
†Kuhnemys palaeocenica Danilov et al. 2015
G. †Perochelys Li et al. 2015
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†Perochelys hengshanensis Brinkman et al. 2017
†Perochelys lamadongensis Li et al. 2015
G. †Petrochelys Vitek et al. 2017
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†Petrochelys kyrgyzensis Nessov 1995
G. †Paleotrionyx Schmidt 1945
G. †Striatochelys Massonne et al. 2023
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†Striatochelys baba Massonne et al. 2023
G. †Temnotrionyx Hay 1908
Subfm. Trionychinae Gray 1825 [softshell turtle]
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G. †Aulacochelys Lydekker 1889
G. †Axestemys Hay 1899
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†Axestemys byssina Cope 1872
†Axestemys byssinus Cope 1872
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Invalid names: Eugenechelys robertemryi Chkhikvadze 2008 [synonym], Temnotrionyx manducans Hay 1908 [synonym]
†Axestemys cerevisia Vitek 2012
†Axestemys erquelinnensis Perez-Garcia and Smith 2021
†Axestemys infernalis Joyce et al. 2019
†Axestemys montinsana Vitek 2012
†Axestemys quinni Schmidt 1945
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Invalid names: Aspideretes granifer Hay 1909 [synonym], Aspideretes planus Parks 1933 [synonym]
†Axestemys vittata Pomel 1847
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Invalid names: Eurycephalochelys fowleri Moody and Walker 1970 [synonym], Trionyx levalensis Dollo 1909 [synonym]
Invalid names: Axestus Cope 1872 [synonym], Eurycephalochelys Moody and Walker 1970 [synonym]
Tr. Chitrini Gray 1870
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G. Amyda Geoffroy Saint-Hilaire 1809
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Amyda cartilaginea Boddaert 1770
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Invalid names: Trionyx trinilensis Jaekel 1911 [synonym]
†Amyda exquisita Hay 1908
G. †Palaeoamyda Cadena 2016
G. Dogania Gray 1844
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†Dogania maortuensis Yeh 1965
Dogania subplana Geoffrey Saint-Hilaire 1809 [Malayan softshell]
G. Nilssonia Gray 1869 [Burmese peacock softshells]
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Nilssonia formosa Gray 1869 [Burmese peacock softshell]
Nilssonia gangetica Cuvier 1825 [Indian softshell turtle]
Nilssonia hurum Gray 1830
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Invalid names: Trionyx hurum Gray 1831 [synonym]
Nilssonia leithii Gray 1872 [Leiths softshell turtle]
Nilssonia nigricans Anderson 1875 [Black softshell turtle]
Invalid names: Trionyx hurum sivalensis Lydekker 1889 [synonym]
G. Palea Siebenrock 1906 [Wattleneck softshells]
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Palea steindachneri Siebenrock 1906 [Wattleneck softshell]
G. Pelodiscus Fitzinger 1835
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Pelodiscus axenaria Zhou et al. 1991 [Hunan softshell turtle]
†Pelodiscus gracilia Yeh 1963
Pelodiscus maackii Brandt 1857 [Northern Chinese softshell turtle]
Pelodiscus parviformis Tang 1997 [Lesser Chinese softshell turtle]
Pelodiscus sinensis Wiegmann 1835 [Chinese softshell]
Invalid names: Trionyx sinuosus Chow and Yeh 1958 [nomen dubium]
G. Rafetus Gray 1864
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†Rafetus bohemicus Liebus 1930
Rafetus euphraticus Daudin 1802 [Euphrates softshell]
Rafetus swinhoei Gray 1873
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Invalid names: Pelochelys taihuensis Zhang 1984 [synonym], Trionyx liupani Tao 1986 [synonym]
Tr. Trionychini Gray 1825
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Subtr. Apalonina Meylan 1987 [softshell turtle]
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G. †Ulutrionyx Kordikova 1994
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†Ulutrionyx ninae Chkhikvadze 1971
G. †Gobiapalone Danilov et al. 2014
G. †Oliveremys Vitek 2011
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†Oliveremys uintaensis Cope 1873
G. †Rafetoides Karl 1998
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†Rafetoides messelianus Reinach 1900
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Invalid names: Rafetoides austriacus Peters 1858 [synonym], Trionyx messelianus kochi Hummel 1927 [synonym]
Invalid names: Trionyx messelianus lepsiusi Hummel 1927 [synonym]
Invalid names: Rafetoidina Karl 1998 [empty], Trionychina Gray 1825 [empty]
G. Trionyx Geoffroy Saint-Hilaire 1809
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†Trionyx admirabilis Hay 1905
†Trionyx aequa Hay 1908
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Invalid names: Amyda crassa Hay 1908 [synonym]
†Trionyx allani Gilmore 1923
†Trionyx austerus Hay 1908
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Invalid names: Aspideretes fontanus Hay 1908 [synonym], Aspideretes vorax Hay 1908 [synonym]
†Trionyx baynshirensis Danilov et al. 2014
†Trionyx beecheri Hay 1904
†Trionyx boulengeri Reinach 1900
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Invalid names: Trionyx clavatomarginatus Lörenthey 1903 [synonym]
†Trionyx capellinii Negri 1892
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Invalid names: Trionyx affinis Negri 1892 [synonym], Trionyx capellinii gracilina Sacco 1895 [synonym], Trionyx capellinii montevialensis Negri 1892 [synonym], Trionyx capellinii perexpansa Sacco 1895 [synonym], Trionyx gemmellaroi Negri 1892 [synonym], Trionyx insolitus Bergounioux 1954 [synonym], Trionyx intermedius Bergounioux 1954 [synonym], Trionyx schaurothianus Negri 1893 [synonym]
†Trionyx dissolutus Vitek and Danilov 2014
†Trionyx ellipticus Hay 1908
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Invalid names: Aspideretes annae Zangerl 1944 [synonym], Aspideretes grangeri Hay 1908 [synonym]
†Trionyx eloisae Gilmore 1919
†Trionyx foveatus Leidy 1856
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Invalid names: Aspideretes maturus Lambe 1914 [synonym], Aspideretes rugosus Parks 1933 [synonym], Aspideretes subquadratus Lambe 1914 [synonym]
Trionyx gangeticus Cuvier 1824
†Trionyx gilbentuensis Danilov et al. 2014
†Trionyx gobiensis Danilov et al. 2014
†Trionyx gregarius Gilmore 1934
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Invalid names: Rafetus gilmorei Chkhikvadze 1999 [synonym]
†Trionyx henrici Owen and Bell 1849
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Invalid names: Trionyx barbarae Owen and Bell 1849 [synonym], Trionyx circumsulcatus Owen and Bell 1849 [synonym], Trionyx incrassatus Owen and Bell 1849 [synonym], Trionyx marginatus Owen and Bell 1849 [synonym], Trionyx planus Owen and Bell 1849 [synonym], Trionyx rivosus Owen and Bell 1849 [synonym]
†Trionyx ikoviensis Danilov et al. 2011
†Trionyx jixiensis Li et al. 2015
†Trionyx johnsoni Gilmore 1931
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Invalid names: Amyda neimenguensis Yeh 1965 [synonym]
†Trionyx kansaiensis Vitek and Danilov 2010
†Trionyx latus Gilmore 1919
†Trionyx linchuensis Yeh 1962
†Trionyx miensis Okazaki and Yoshida 1977
†Trionyx minusculus Chkhikvadze 1973
†Trionyx miocaenus Matthew 1924
†Trionyx mira Hay 1908
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Invalid names: Platypeltis extensa Hay 1908 [synonym], Platypeltis postera Hay 1908 [synonym]
†Trionyx ninae Chkhikvadze 1971
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Invalid names: Trionyx turgaicus Kuznetsov and Chkhikvadze 1977 [synonym], Trionyx zaisanensis Chkhikvadze 1973 [synonym]
†Trionyx onomatoplokos Kuznetsov and Chkhikvadze 1987
Trionyx phayrei Theobald 1868
†Trionyx pliocenicus Fucini 1912
†Trionyx reesidei Gilmore 1919
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Invalid names: Aspideretes quadratus Gilmore 1919 [synonym]
†Trionyx riabinini Kuznetsov and Chkhikvadze 1987
†Trionyx robustus Gilmore 1919
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Invalid names: Aspideretes ovatus Gilmore 1935 [synonym]
†Trionyx shiluutulensis Danilov et al. 2014
†Trionyx silvestris Walker and Moody 1974
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Invalid names: Trionyx michauxi Broin 1977 [synonym]
†Trionyx singularis Hay 1907
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Invalid names: Aspideretes puercensis Hay 1908 [synonym], Aspideretes sagatus Hay 1908 [synonym]
Trionyx triunguis Forksal 1775 [African softshell turtle]
†Trionyx uintaensis Cope 1873
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Invalid names: Amyda franciscae Hay 1908 [synonym], Amyda nelsoni Case 1927 [synonym], Aspidonectes tritor Hay 1904 [synonym]
†Trionyx vindobonensis Peters 1855
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Invalid names: Trionyx brunhuberi Ammon 1911 [synonym], Trionyx gergensi Reinach 1900 [synonym], Trionyx hoernesi Heritsch 1909 [synonym], Trionyx peneckei Heritsch 1909 [synonym], Trionyx petersi Hoernes 1881 [synonym], Trionyx rostratus Arthaber 1898 [synonym], Trionyx septemcostatus Hoernes 1881 [synonym], Trionyx teiritzbergensis Gemel 2002 [synonym], Trionyx teyleri Winkler 1869 [synonym]
Invalid names: Francedebroinella Chkhikvadze 1999 [synonym], Sinamyda Chkhikvadze 2000 [synonym], Trionyx capellinii conjugens Sacco 1894 [synonym], Trionyx nopcsai Szalai 1934 [nomen dubium]
Invalid names: Trionyx (Amyda) subangularis Bergounioux 1954 [nomen dubium], Trionyx bambolii Ristori 1891 [nomen dubium], Trionyx blayaci Bergounioux 1933 [nomen dubium], Trionyx desmostyli Matsumoto 1918 [nomen dubium], Trionyx dieupentalensis Bergounioux 1935 [nomen dubium], Trionyx harmati Szalai 1934 [nomen dubium], Trionyx irregularis Bergounioux 1954 [nomen dubium], Trionyx pliopedemontana Sacco 1889 [nomen dubium], Trionyx stadleri Teppner 1913 [nomen dubium], Trionyx stiriacus Peters 1855 [nomen dubium]
G. †Zangerlichelys Chkhikvadze 2000
Invalid names: Altaytrionyx Chkhikvadze 2008 [nomen dubium], Altaytrionyx burtschaki Chkhikvadze 2008 [nomen dubium], Altaytrionyx devjatkini Chkhikvadze 2008 [nomen dubium], Altaytrionyx phiruzae Chkhikvadze 2008 [nomen dubium], Amyda menneri Chkhikvadze 1988 [nomen dubium], Aspideretes jaxarticus Riabinin 1938 [nomen nudum], Aspideretes muyuensis Lei and Ye 1985 [nomen dubium], Aspilus Portis 1885 [nomen dubium], Aspilus cortesii Portis 1885 [nomen dubium], Chitra minor Jaekel 1911 [nomen dubium], Lissemydinae Williams 1950 [empty], Paleotrionyx jimenezfuentesi Chkhikvadze 1995 [nomen dubium], Plastomenus gabunii Chkhikvadze 1984 [nomen dubium], Plastomenus jaxarticus Riabinin 1938 [nomen nudum], Plastomenus mlynarskii Chkhikvadze 1970 [nomen dubium], Platypeltis subcircularis Chow and Yeh 1957 [nomen dubium], Procyclanorbis Portis 1901 [nomen dubium], Rafetini Chkhikvadze 2000 [empty], Rafetus karkhualexandri Chkhikvadze 2000 [nomen nudum], Rafetus yexiangkuii Chkhikvadze 1999 [nomen dubium], Trionyx acutiformis Bergounioux 1935 [nomen dubium], Trionyx amansii Gray 1831 [nomen dubium], Trionyx anthracotheriorum Portis 1883 [nomen dubium], Trionyx aquitanicus Delfortrie 1869 [nomen dubium], Trionyx aspidiformis Laube 1900 [nomen dubium], Trionyx australiensis De Vis 1894 [nomen dubium], Trionyx austriacus Peters 1859 [nomen dubium], Trionyx borkenensis Gramann 1956 [nomen dubium], Trionyx bowerbanki Lydekker 1889 [nomen dubium], Trionyx bruxelliensis Winkler 1869 [nomen dubium], Trionyx burdigalensis Bergounioux 1935 [nomen dubium], Trionyx capellinii bulgaricus Khosatzky et al. 1983 [nomen dubium], Trionyx chaubeti Bergounioux 1935 [nomen dubium], Trionyx ciryi Bergounioux 1935 [nomen dubium], Trionyx cliftii Fitzinger 1835 [nomen nudum], Trionyx croaticus Koch 1915 [nomen dubium], Trionyx danovi Chkhikvadze 1989 [nomen dubium], Trionyx dodunii Gray 1831 [nomen nudum], Trionyx elongatus Liebus 1930 [nomen dubium], Trionyx erquelinnensis Dollo 1909 [nomen nudum], Trionyx hilberi Hoernes 1892 [nomen dubium], Trionyx ishiharaensis Miura and Uyama 1987 [nomen dubium], Trionyx italicus Schauroth 1865 [nomen dubium], Trionyx jakhimovitchae Chkhikvadze 1989 [nomen dubium], Trionyx kazusensis Otsuka 1969 [nomen dubium], Trionyx khosatzkyi Chkhikvadze 1983 [nomen dubium], Trionyx laurillardii Gray 1831 [nomen dubium], Trionyx lockardi Gray 1831 [nomen nudum], Trionyx lorioli Portis 1882 [nomen dubium], Trionyx manouri Gray 1831 [nomen dubium], Trionyx marini Hernández Sampelayo and Bataller 1944 [nomen dubium], Trionyx moldaviensis Khosatzky 1986 [nomen dubium], Trionyx mourieri Bergounioux 1935 [nomen dubium], Trionyx munzenbergensis Hummel 1927 [nomen nudum], Trionyx oligocenica Negri 1892 [nomen nudum], Trionyx oweni Reinach 1900 [nomen dubium], Trionyx parisiensis Gray 1831 [nomen dubium], Trionyx partschii Peters 1855 [nomen dubium], Trionyx pedemontana Portis 1879 [nomen dubium], Trionyx petersi trifailensis Teppner 1914 [nomen dubium], Trionyx pliocaenicus Reinach 1903 [nomen dubium], Trionyx pompignanensis Bergounioux 1933 [nomen dubium], Trionyx pontanus Laube 1895 [nomen dubium], Trionyx portisi Ristori 1891 [nomen dubium], Trionyx preschenensis Laube 1900 [nomen dubium], Trionyx propinquus Ristori 1891 [nomen dubium], Trionyx pseudovindobonensis Szalai 1934 [nomen dubium], Trionyx pustulatus Owen and Bell 1849 [nomen dubium], Trionyx ragusensis De Gregorio 1883 [nomen dubium], Trionyx reticulatus Rieppel 1979 [nomen dubium], Trionyx rocchettiana Portis 1882 [nomen dubium], Trionyx roncensis Harrassowitz 1919 [nomen dubium], Trionyx rotundiformis Bergounioux 1933 [nomen dubium], Trionyx sculptus Gilmore 1931 [nomen dubium], Trionyx senckenbergianus Reinach 1903 [nomen dubium], Trionyx senensis Ristori 1891 [nomen dubium], Trionyx siegeri Heritsch 1909 [nomen dubium], Trionyx sophiae Heritsch 1909 [nomen dubium], Trionyx stormsi Delheid 1899 [nomen dubium], Trionyx ubeensis Chitani 1925 [nomen dubium], Trionyx valdensis Portis 1882 [nomen dubium], Trionyx zakhidovi Khosatzky 1966 [nomen dubium], Zaisanonyx Chkhikvadze 2008 [nomen dubium]