Classification of the subfamily Harpalii

Subfm. Harpalinae Bonelli 1810
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Tr. Abacetini Chaudoir 1873
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G. Loxandrus Leconte 1852
G. †Aethocarabus Ren 1995
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Tr. Chlaeniini Brullé 1834
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Subtr. Chlaeniina Brullé 1834
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G. Chlaenius Bonelli 1810
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Subg. Chlaenius (Agostenus) Fischer von Waldheim 1829
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Subg. Chlaenius (Chlaeniellus) Reitter 1808
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Chlaeniellus nigricornis Fabricius 1787
†Chlaenius electrinus Giebel 1862
†Chlaenius furvus Hong 1984
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Invalid names: Shanwangicarabus paucilus Hong and Wang 1985 [synonym]
Chlaenius gebleri Ganglbauer 1891
†Chlaenius plicatipennis Wickham 1917
†Chlaenius primaevus Piton and Théobald 1939
†Chlaenius punctulatus Horn 1876
Invalid names: Shanwangicarabus Hong 1984 [synonym]
Tr. Galeritini Kirby 1825
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Subtr. Galeritina Kirby 1825
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G. Galerita Fabricius 1801
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Galerita bicolor Drury 1770
†Galerita marshii Scudder 1900
Tr. Harpalini Bonelli 1810
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Subtr. Anisodactylina Lacordaire 1854
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G. Anisodactylus Dejean 1829
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Invalid names: Eurytrichi Le Conte 1847 [synonym]
Subtr. Harpalina Bonelli 1810
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G. Cratacanthus Dejean 1829
G. Euryderus Le Conte 1846
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Invalid names: Nothopus Leconte 1852 [objective synonym]
G. Harpalus Latreille 1802
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†Harpalus abolitus von Heyden and von Heyden 1866
†Harpalus bruckmanni Heer 1862
†Harpalus conditus Scudder 1900
†Harpalus constrictus Heer 1862
†Harpalus deletus Oustalet 1874
†Harpalus dilatatus Theobald 1937
†Harpalus excavatus Förster 1891
†Harpalus lheritieri Piton 1940
†Harpalus maceratus Wickham 1911
†Harpalus nero Oustalet 1874
†Harpalus nitens Heer 1870
†Harpalus nuperus Scudder 1900
†Harpalus obtusus Förster 1891
†Harpalus oustaleti Meunier 1915
†Harpalus pleistocenicus Lomnicki 1894
Harpalus pusillus Motschulsky 1850
†Harpalus redivivus Wickham 1917
†Harpalus sinis Heer 1847
†Harpalus stierlini Heer 1862
†Harpalus stygius Heer 1862
†Harpalus tabidus Heer 1847
†Harpalus tardigradus Heer 1862
†Harpalus ulomaeformis Wickham 1917
†Harpalus veterum Cockerell 1921
Harpalus vittatus Gelber 1833
†Harpalus whitfieldii Scudder 1900
G. Ophonus Dejean 1821
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Subg. Ophonus (Metophonus) Bedel 1897
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G. Selenophorus Dejean 1829
Invalid names: Dapti Le Conte 1847 [synonym]
Subtr. Stenolophina Kirby 1837
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G. Bradycellus Erichson 1837
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Bradycellus ruficollis Stephens 1828
G. Dicheirotrichus Jacquelin du Val 1857
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G. Stenolophus Dejean 1821
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†Stenolophus religatus Scudder 1900
Tr. Helluonini Hope 1838
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Subtr. Omphrina Jedlicka 1941
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G. Creagris Nietner 1857
G. Helluomorpha De Laporte 1834
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†Helluomorpha protogaea Giebel 1862
Tr. Lebiini Bonelli 1810
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G. †Agatoides Motschulsky 1856
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†Agatoides carinulatus Motschulsky 1856
Subtr. Calleidina Chaudoir 1873
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G. Plochionus Latreille and Dejean 1821
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†Plochionus lesquereuxi Scudder 1900
Subtr. Cymindidina Laporte 1834
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G. Cymindis Latreille 1806
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Subg. Cymindis (Tarulus) Bedel 1906
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Cymindis angularis Gyllenhal 1810
Cymindis arctica Kryzhanovskij and Emets 1979
†Cymindis aurora Horn 1876
†Cymindis extorpescens Scudder 1898
†Cymindis pulchella Heer 1847
G. †Cymindoides Motschulsky 1856
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†Cymindoides sculptipennis Motschulsky 1856
Subtr. Dromiusina Bonelli 1810
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G. †Antephilorhizus Kirichenko-Babko et al. 2021
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†Antephilorhizus pripiatiensis Kirichenko-Babko et al. 2021
†Antephilorhizus zerovae Kirichenko-Babko and Perkovsky 2022
G. Dromius Bonelli 1810
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Dromius angusticollis Sahlberg 1880
†Dromius bakeri Abdullah 1969
†Dromius resinatus Germar 1813
Invalid names: Lebina Germar 1813 [synonym]
G. Microlestes Schmidt-Göbel 1846
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G. Paradromius Fowler 1887
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Paradromius linearis Olivier 1795
G. Syntomus Hope 1838
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Syntomus americanus Dejean 1831
Syntomus truncatellus Linnaeus 1761
Subtr. Lebiina Bonelli 1810
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G. Lebia Latreille 1802 [colorful foliage ground beetle]
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†Lebia amissa von Heyden and von Heyden 1865
†Lebia harrelli Cockerell 1936
†Lebia minuscula Piton 1936
†Lebia newberryi Scudder 1900
†Lebia protospiloptera Cockerell 1921
G. †Miolebidia Zhang 1989
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†Miolebidia gaura Zhang 1989
Subtr. Pericalina Hope 1838
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G. Coptodera Dejean 1825
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†Coptodera elektra Gamboa and Ortuño 2015
G. †Protoscalidion Schaufuss 1889
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†Protoscalidion rugiae Schaufuss 1889
Invalid names: Brachinus repressus Scudder 1900 [nomen dubium]
Tr. Licinini Bonelli 1810
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Subtr. Dicaelina Laporte 1834
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G. Dicaelus Bonelli 1813
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†Dicaelus alutaceus Horn 1876
G. Diplocheila Brullé 1834
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†Diplocheila henshawi Scudder 1890
Invalid names: Rembus Latreille 1817 [replaced]
Subtr. Licinina Bonelli 1810
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G. Badister Clairville 1806
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†Badister antecursor Scudder 1900
†Badister debilis Heer 1847
†Badister fragilis Heer 1862
†Badister grandis Heer 1862
†Badister prodromus Heer 1847
Tr. Odacanthini Laporte 1834
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Invalid names: Casnoniae LeConte 1861 [synonym]
Tr. Panagaeini Bonelli 1810
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Subtr. Panagaeina Bonelli 1810
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G. Panagaeus Latreille 1802
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†Panagaeus dryadum Oustalet 1874
Tr. Pseudomorphini Hope 1838
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G. †Burmapseudomorphus Beutel et al. 2020
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†Burmapseudomorphus planus Beutel et al. 2020
Tr. Pterostichini Bonelli 1810
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G. Cyclotrachelus Chaudoir 1838
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Subg. Cyclotrachelus (Evarthrus) Leconte 1853
†Cyclotrachelus tenebricus Scudder 1900
Subtr. Pterostichina Bonelli 1810
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G. Abax Bonelli 1810
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†Abax durhamianus Lesne 1926
Abax parallelus Duftschmid 1812
G. Myas Dejean 1828
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†Myas rigefactus Scudder 1900
†Myas umbrarum Scudder 1900
G. Platysma Bonelli 1810
G. Plocamostethus Britton 1940
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G. Poecilus Bonelli 1810
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Poecilus kugelanni Panzer 1797
Poecilus nearcticus Lindroth 1966
†Poecilus virgatus Pongrácz 1935
Invalid names: Feronia Latreille 1816 [synonym]
G. Pterostichus Bonelli 1810
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Subg. Pterostichus (Argutor) Dejean 1821
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†Argutor antiquus Heer 1847
†Argutor pliocenicus Lesne 1920
Subg. Pterostichus (Eosteropus) Tschitschérine 1902
Subg. Pterostichus (Melanius) Bonelli 1810
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Invalid names: Pterostichus (Omaseus) Dejean 1821 [synonym]
Subg. Pterostichus (Metallophilus) Chaudoir 1838
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Invalid names: Pterostichus (Lyperopherus) Motschulsky 1844 [synonym]
Subg. Pterostichus (Phonias) des Gozis 1886
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Subg. Pterostichus (Steropus) Dejean 1821
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Invalid names: Pterostichus (Steroderus) Motschulsky 1850 [synonym]
Pterostichus abnormis Sahlberg 1880
†Pterostichus abrogatus Scudder 1890
Pterostichus cancellatus Motschulsky 1860
Pterostichus costatus Ménétriès 1851
†Pterostichus depletus Scudder 1900
†Pterostichus destitutus Scudder 1890
†Pterostichus destructus Scudder 1890
†Pterostichus distinctus Piton and Théobald 1939
†Pterostichus dormitans Scudder 1890
†Pterostichus fernquisti Wickham 1931
†Pterostichus fractus Scudder 1890
†Pterostichus gelidus Scudder 1877
†Pterostichus gurnetensis Cockerell 1921
†Pterostichus laevigatus Scudder 1890
Pterostichus minor Gyllenhal 1827
†Pterostichus molestus Theobald 1937
Pterostichus niger Schaller 1783
Pterostichus nigrita Paykull 1790
Pterostichus parens Tschitscherine 1897
†Pterostichus pumpellyi Scudder 1900
Pterostichus sublaevis Sahlberg 1880
Pterostichus tundrae Tschitscherine 1894
†Pterostichus vetustus Heer 1862
†Pterostichus walcotti Scudder 1900
G. Stomis Clairville 1806
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†Stomis elegans Oustalet 1874
Invalid names: Feroniens Dejean 1825 [synonym]
G. †Sinis Heer 1862
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†Sinis brevicollis Heer 1862
Tr. Sphodrini Laporte 1834
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Subtr. Calathina Laporte 1834
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G. Calathus Bonelli 1810
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Calathus fuscipes Goeze 1777
Calathus piceus Marsham 1802
G. †Quasicalathus Schmidt and Will 2022
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†Quasicalathus agonicollis Schmidt and Will 2022
†Quasicalathus conservans Schmidt and Will 2022
†Quasicalathus elpis Ortuño and Arillo 2009
Subtr. Sphodrina Laporte 1834
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G. Pristonychus Dejean 1828
G. Sphodrus Clairville 1806
Subtr. Synuchina Lindroth 1956
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G. Synuchus Gyllenhal 1810
Tr. Zabrini Bonelli 1810
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Subtr. Amarina Zimmerman 1832
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G. Amara Bonelli 1810
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Subg. Amara (Amara) Bonelli 1810
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†Amara (Amara) subaenea Lomnicki 1894
Subg. Amara (Bradytus) Stephens 1827
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Subg. Amara (Celia) Zimmerman 1832
Subg. Amara (Curtonotus) Stephens 1827
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Curtonotus alpinus Paykull 1790
Curtonotus fodinae Mannerheim 1825
Subg. Amara (Paracelia) Bedel 1899
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Amara aenea De Geer 1774
Amara arcticola Poppius 1906
Amara aulica Panzer 1796
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Invalid names: Carabus ruficornis De Geer 1774 [replaced]
Amara bifrons Gyllenhal 1810
Amara brunnea Gyllenhal 1810
†Amara cockerelli Wickham 1912
Amara conflata Le Conte 1855
†Amara danae Scudder 1900
Amara erratica Duftschmid 1812
†Amara foersteri Handlirsch 1907
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Invalid names: Amara sinuata Förster 1891 [replaced]
Amara insignis Dejean 1831
Amara municipalis Duftschmid 1812
†Amara occlusa Scudder 1900
†Amara paleomelas Scudder 1879
†Amara pinguicula Heer 1862
†Amara powellii Scudder 1900
†Amara primigenia Heer 1862
†Amara princeps Heer 1862
†Amara procera Förster 1891
Amara pseudobrunnea Lindroth 1968
†Amara revocata Scudder 1900
†Amara sterilis Scudder 1900
†Amara veterata Scudder 1900
Subtr. Zabrina Bonelli 1810
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G. Zabrus Clairville 1806
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Tr. Zuphiini Bonelli 1810
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Subtr. Zuphiina Bonelli 1810
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G. Polistichus Bonelli 1810
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†Polystichus hopei Oustalet 1874
Invalid names: Dryptini Bonelli 1810 [empty], Licininae Bonelli 1810 [synonym]