Classification of the subfamily Scarabaeinae

Subfm. Scarabaeinae Latreille 1802
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G. Anachalcos Hope 1837
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†Anachalcos mfwangani Paulian 1976
Tr. Ateuchini Perty 1830
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Subtr. Ateuchina Perty 1830
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G. Ateuchus Weber 1801
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Invalid names: Choeridium Lepeletier de Saint-Fargeau and Audinet-Serville 1828 [synonym], Choeridium ebeninum Horn 1876 [nomen dubium]
G. †Lobateuchus Montreuil et al. 2010
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†Lobateuchus parisii Montreuil et al. 2010
G. Canthochilum Chapin 1934
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†Canthochilum alleni Tarasov and Vaz-de-Mello 2016
†Canthochilum philipsivieorum Tarasov and Vaz-de-Mello 2016
Tr. Coprini Leach 1815
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G. Copris Geoffroy 1762
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Subg. Copris (Copris) Geoffroy 1762
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†Copris (Copris) kartlinus Kabakov 1988
†Copris druidum Heer 1862
Copris incertus Say 1835
†Copris leakeyorum Paulian 1976
Copris lunaris Linnaeus 1758 [horned dung beetle]
†Copris pristinus Pierce 1946
Tr. Deltochilini Lacordaire 1856
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G. Canthon Hoffmannsegg 1817
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G. Deltochilum Eschscholtz 1822
G. Scybalophagus Martínez 1953
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†Scybalophagus brellenthinae Tello et al. 2023
Tr. Gymnopleurini Lacordaire 1856
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G. Gymnopleurus Illiger 1803
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†Gymnopleurus sisyphus Heer 1847
G. Heliocopris Hope 1837
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†Heliocopris antiquus Fujiyama 1968
G. Metacatharsius Paulian 1939
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†Metacatharsius rusingae Paulian 1976
Tr. Oniticellini Kolbe 1905
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Subtr. Drepanocerina van Lansberge 1875
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G. Eodrepanus Barbero et al. 2009
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†Eodrepanus coopei Barbero et al. 2009
Subtr. Oniticellina Kolbe 1905
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G. Oniticellus Dejean 1821
Tr. Onitini Laporte 1840
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G. Onitis Fabricius 1798
Tr. Onthophagini Burmeister 1846
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G. Onthophagus Latreille 1802
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†Onthophagus arvernus Piton and Théobald 1939
†Onthophagus crassus Heer 1862
†Onthophagus everestae Pierce 1946
Onthophagus fracticornis Preyssler 1790
Onthophagus janus Panzer 1794
Onthophagus lenzii Harold 1874
†Onthophagus ovatulus Heer 1847
†Onthophagus pilauco Tello et al. 2021
†Onthophagus prodromus Heer 1862
†Onthophagus statzi Krell 1990
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Invalid names: Onthophagus muelleri Statz 1952 [replaced]
Onthophagus taurus Schreber 1759
G. Phalops Erichson 1848
Tr. Phanaeini Hope 1838
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G. Phanaeus Macleay 1819
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†Phanaeus labreae Pierce 1946
†Phanaeus violetae Zunino 2013
Invalid names: Palaeocopris Pierce 1946 [synonym], Phanaeus antiquus Horn 1876 [nomen dubium]
Tr. Scarabaeini Latreille 1802
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G. Scarabaeus Linnaeus 1758
Invalid names: Onthophagus spitsbergeniensis Krell 2010 [nomen dubium]