Elmo, MCZ 1927 collection: Artinskian, Kansas
collected by F.M. Carpenter, J.W. Wilson, W.S. Creighton 1927

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Xiphosura - Xiphosurida - Paleolimulidae
Paleolimulus avitus Dunbar 1923
Raymond 1944
synonym of Paleolimulus signatus
Insecta - Meganisoptera - Meganeuridae
Megatypus schucherti Tillyard 1925
Carpenter 1931 1 specimen
MCZ 3048
Typus permianus (Sellards 1906)
Carpenter 1931 6 specimens
original and current combination Tupus permianus
MCZ 3050-5
Oligotypus sp. Carpenter 1931
Carpenter 1931 1 specimen
MCZ 3041
Oligotypus tillyardi n. gen., n. sp. Carpenter 1931
Carpenter 1931 1 specimen
MCZ 3049 (1 measurement)
Insecta - Ditaxineuridae
Ditaxineura anomalostigma Tillyard 1926
Carpenter 1931 4 specimens
MCZ 3043-6
Insecta - Progoneuridae
Progoneura minuta n. gen., n. sp. Carpenter 1931
Carpenter 1931 1 specimen
MCZ 3042 (1 measurement)
Insecta - Permoneuridae
Permoneura lameerei n. gen., n. sp. Carpenter 1931
Carpenter 1931 1 specimen
MCZ 3135 (1 measurement)
Insecta - Trachopterygidae
Trachopteryx martynovi n. gen., n. sp. Carpenter 1976
Carpenter 1976 1 specimen
MCZ 7497 (1 measurement)
Insecta - Apheloneuridae
Apheloneura amplia n. sp. Carpenter 1976
Carpenter 1976 3 specimens
MCZ 8604, 7525, 7522 (1 measurement)
Apheloneura minutissima n. gen., n. sp. Carpenter 1976
Carpenter 1976 2 specimens
MCZ 7527, 7534 (4 measurements)
Insecta - Permopsocida - Permopsocidae
Lithopsocidium permianum n. gen., n. sp. Carpenter 1932
Carpenter 1932 1 specimen
MCZ 3168 (1 measurement)
Orthopsocus singularis n. gen., n. sp. Carpenter 1932
Carpenter 1932 1 specimen
MCZ 3200 (1 measurement)
Progonopsocus permianus Tillyard 1926
Carpenter 1932 3 specimens
MCZ 3175-7 (1 measurement)
Permopsocus latipennis Tillyard 1926
Carpenter 1932 9 specimens
MCZ 3155,3181,2,4,5,8
Insecta - Permopsocida - Psocidiidae
Dichentomum parvulum n. sp. Carpenter 1932
Carpenter 1932 1 specimen
MCZ 3173
Dichentomum tinctum Tillyard 1926
Carpenter 1932 100 specimens
Over 100 specimens including MCZ 3138,3141-7,3162-5 (1 measurement)
Dichentomum minimum (Tillyard 1926)
Carpenter 1932 6 specimens
MCZ 3167 and 3171 are best
Dichentomum latum n. sp. Carpenter 1932
Carpenter 1932 1 specimen
MCZ 3160 (1 measurement)
Insecta - Lophioneurida - Lophioneuridae
Cyphoneura reducta n. sp. Carpenter 1932
Carpenter 1932 1 specimen
recombined as Cyphoneurodes reducta
MCZ 3198 (1 measurement)
Cyphoneura permiana n. gen., n. sp. Carpenter 1932
Carpenter 1932 1 specimen
MCZ 3197 (1 measurement)
Insecta - Hemiptera - Archescytinidae
Permopsylla americana Tillyard 1926
Carpenter 1931 6 specimens
MCZ 3123-3125,?3126,?3128,?3134
Permopsylla permiana n. sp. Carpenter 1931
Carpenter 1931 3 specimens
MCZ 3129-31 (1 measurement)
Permopsylla minuta n. sp. Carpenter 1931
Carpenter 1931 1 specimen
MCZ 3132 (1 measurement)
Permoscytina muiri Tillyard 1926
Carpenter 1931 1 specimen
recombined as Archescytina muiri
MCZ 3122
Permoscytina kansasensis Tillyard 1926
Carpenter 1931 1 specimen
recombined as Archescytina kansasensis
MCZ 3121
Archescytina permiana Tillyard 1926
Carpenter 1931 3 specimens
MCZ 3120,3136-7
Paleoscytina brevistigma n. gen., n. sp. Carpenter 1931
Carpenter 1931 1 specimen
MCZ 3133 (1 measurement)
Insecta - Permembiidae
Permembia delicatula Tillyard 1928
Carpenter 1976 5 specimens
MCZ 7526,7539,7547,7596,7474
Insecta - Gelasopteridae
Gelasopteron gracile n. gen., n. sp. Carpenter 1976
Carpenter 1976 1 specimen
MCZ 7511 (1 measurement)
Insecta - Grylloblattodea - Liomopteridae
Liomopterum ornatum Sellards 1909
Carpenter 1950
36 from lower limestone, at least 1 from 1927
Insecta - Grylloblattodea - Lemmatophoridae
Lisca minuta Sellards 1909
Carpenter 1935 62 specimens
104 total, minus 42 collected in 1932
Artinska ovata (Sellards 1909)
Carpenter 1935 55 specimens
76 total, minus 21 collected in 1932
Artinska clara Sellards 1909
Carpenter 1935 96 specimens
160 total, minus 64 collected in 1932
Lemmatophora typa Sellards 1909
Carpenter 1935
More than 200 total, but years not given (2 measurements)
Paraprisca fragilis (Sellards 1909)
Carpenter 1935
36 total from 1927 and 1932
Paraprisca grandis n. sp. Carpenter 1935
Carpenter 1935 2 specimens
MCZ 3584,5 (1 measurement)
Lecorium elongatum Sellards 1909
Carpenter 1935
46 total, at least 1 from 1927
Insecta - Cnemidolestodea - Parmapteridae
Opisthocladus arcuatus n. gen., n. sp. Carpenter 1976
Carpenter 1976 1 specimen
MCZ 5882 (1 measurement)
Insecta - Cnemidolestodea - Tococladidae
Opisthocladus strictus n. sp. Carpenter 1976
Carpenter 1976 1 specimen
recombined as Elmocladus strictus
MCZ 5881 (2 measurements)
Insecta - Cnemidolestodea - Tillyardembiidae
Heteroptilon costale n. gen., n. sp. Carpenter 1976
Carpenter 1976 1 specimen
MCZ 5878 (1 measurement)
Insecta - Cnemidolestodea - Psoropteridae
Psoroptera cubitalia n. gen., n. sp. Carpenter 1976
Carpenter 1976 1 specimen
MCZ 5840 (1 measurement)
Insecta - Caloneurodea - Caloneuridae
Plesiogramma reducta n. sp. Carpenter 1943
Carpenter 1943 1 specimen
synonym of Pleisiogramma medialis
(1 measurement)
Insecta - Nugonioneuridae
Nugonioneura problematica Tillyard 1937
Carpenter 1976 1 specimen
(1 measurement)
Insecta - Paoliida - Anthracoptilidae
Homocladus grandis n. gen., n. sp. Carpenter 1966
Carpenter 1966 2 specimens
(1 measurement)
Insecta - Paoliida - Blattinopsidae
Glaphyrophlebia ovata (Sellards 1909)
Carpenter 1966 1 specimen
MCZ 4965 (1 measurement)
Glaphyrophlebia speciosa (Sellards 1909)
Carpenter 1966 1 specimen
MCZ 6303 (1 measurement)
Insecta - Orthoptera - Pruvostitidae
Paroedischia recta n. gen., n. sp. Carpenter 1966
Carpenter 1966 4 specimens
MCZ 5897 (type from here; 5 specimens in total) (1 measurement)
Insecta - Protelytroptera - Elytroneuridae
Elytroneura permiana Carpenter 1933
Carpenter 1933
(1 measurement)
Insecta - Protomeroptera - Protomeropidae
Platychorista venosa Tillyard 1926
8 specimens
MCZ 3001-8 (2 measurements)
Insecta - Mecoptera - Permopanorpidae
Permopanorpa inaequalis Tillyard 1926
25 specimens
MCZ 3009-3033 (3 measurements)
Insecta - Mecoptera - Permochoristidae
Protopanorpa permiana Tillyard 1926
2 specimens
MCZ 3034-5 (1 measurement)
Protochorista tetraclada Tillyard 1926
1 specimen
MCZ 3036
Agetopanorpa maculata n. gen., n. sp. Carpenter 1930
1 specimen
MCZ 3037 (1 measurement)
Insecta - Permithonidae
Elmothone martynovae n. gen., n. sp. Carpenter 1976
Carpenter 1976 1 specimen
MCZ 5585 (1 measurement)
Insecta - Palaeodictyoptera - Elmoboriidae
Elmoboria piperi n. gen., n. sp. Carpenter 1976
Carpenter 1976 1 specimen
MCZ 5839 (1 measurement)
Insecta - Palaeodictyoptera - Spilapteridae
Dunbaria fasciipennis Tillyard 1924
Carpenter 1931 3 specimens
MCZ 3056-8
Insecta - Megasecoptera - Protohymenidae
Protohymen permianus Tillyard 1924
Carpenter 1930 11 specimens
MCZ 3060-8,3070
Protohymen elongatus n. sp. Carpenter 1930
Carpenter 1930 1 specimen
MCZ 3069 (1 measurement)
Permohymen schucherti Tillyard 1924
Carpenter 1930 4 specimens
MCZ 3071-4
Insecta - Diaphanopterodea - Asthenohymenidae
Doter minor Sellards 1907
Carpenter 1930 30 specimens
MCZ 3059,3074-94,3111-3119 (1 measurement)
    = Asthenohymen dunbari Tillyard 1924
Carpenter 1933
Doter affinis (Tillyard 1926)
Carpenter 1930 1 specimen
original and current combination Asthenohymen affinis
MCZ 3095
Doter pusillus (Tillyard 1926)
Carpenter 1930 2 specimens
original and current combination Asthenohymen pusillus
MCZ 3097 (2 measurements)
Insecta - Eoblattida
Elmonympha carpenteri n. gen., n. sp. Aristov 2019
Vršanský et al. 2019 1 specimen
MCZ 3622 (year not stated)
see common names

Geography
Country:United States State/province:Kansas
Coordinates: 38.6° North, 97.2° West (view map)
Paleocoordinates:3.6° North, 25.8° West
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Permian Epoch:Cisuralian
Stage:Artinskian 10 m.y. bin:Permian 2
Key time interval:Artinskian
Age range of interval:290.10000 - 283.50000 m.y. ago
Stratigraphy
Formation:Wellington
Stratigraphic resolution:group of beds
Stratigraphy comments: The Elmo limestone itself is a chalky, soft, almost white deposit, about five feet thick; only the very bottom, more massive layer of this stratum contains insects.
Lithology and environment
Primary lithology: lithified "limestone"
Environment:pond Tectonic setting:cratonic basin
Geology comments: Presence of Myalinella may suggest brackish-marine conditions, but stratigraphic position and presence of conchostracans and freshwater ostracods suggest a lacustrine/pond setting.
Taphonomy
Modes of preservation:mold/impression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:selective quarrying,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:MCZ
Collectors:F.M. Carpenter, J.W. Wilson, W.S. Creighton Collection dates:1927
Metadata
Database number:111217
Authorizer:M. Clapham Enterer:M. Clapham, J. Karr
Modifier:M. Clapham Research group:paleoentomology
Created:2011-06-21 10:00:45 Last modified:2014-04-28 18:25:44
Access level:the public Released:2013-06-21 10:00:45
Creative Commons license:CC BY
Reference information

Primary reference:
5018.ETE 730F. M. Carpenter. 1930. The Lower Permian Insects of Kansas: Part 1. Introduction and the Order Mecoptera. Bulletin of the Museum of Comparative Zoology 70(2):69-101 [C. Labandeira/C. Labandeira/M. Clapham]

Secondary references:

5017ETE 729F. M. Carpenter. 1930. The Lower Permian Insects of Kansas: Part 3. The Protohymenoptera. Psyche 37(4):343-374 [C. Labandeira/C. Labandeira/R. Day]
5019ETE 731F. M. Carpenter. 1931. The Lower Permian Insects of Kansas: Part 4. The Order Hemiptera, and additions to the Paleodictyoptera and Protohymenoptera. American Journal of Science 5(22):113-130 [C. Labandeira/C. Labandeira/R. Day]
4972ETE 684F. M. Carpenter. 1931. The Lower Permian Insects of Kansas. Part 2: The Orders Paleodictyoptera, Protodonata and Odonata. American Journal of Science 5(21):97-139 [C. Labandeira/C. Labandeira/R. Day]
4973ETE 685F. M. Carpenter. 1932. The Lower Permian Insects of Kansas. Part 5: Psocoptera and Additions to the Homoptera. American Journal of Science 5(24):1-22 [C. Labandeira/C. Labandeira/R. Day]
4974ETE 686F. M. Carpenter. 1933. The Lower Permian Insects of Kansas. Part 6: Delopteridae, Protelytroptera, Plectoptera and a New Collection of Protodonata, Odonata, Megasecoptera, Homoptera and Psocoptera. Proceedings of the American Academy of Arts and Sciences 68(11):411-505 [C. Labandeira/C. Labandeira/J. Karr]
37051 F. M. Carpenter. 1935. The Lower Permian insects of Kansas, Part 7: the Order Protoperlaria. Proceedings of the American Academy of Arts and Sciences 70:103-146 [M. Clapham/M. Clapham]
37088 F. M. Carpenter. 1943. The Lower Permian Insects of Kansas. Part 9. The orders Neuroptera, Raphidiodea, Caloneurodea and Protorthoptera (Probnisidae), with additional Protodonata and Megasecoptera. Proceedings of the American Academy of Arts and Sciences 75:55-84 [M. Clapham/M. Clapham]
4980ETE 692F. M. Carpenter. 1950. The Lower Permian insects of Kansas. Part 10. The order Protorthoptera: the family Liomopteridae and its relatives. Proceedings of the Academy of Arts and Sciences 78(4):185-219 [C. Labandeira/C. Labandeira/M. Clapham]
4984ETE 696F. M. Carpenter. 1966. The Lower Permian Insects of Kansas: Part 11. The Orders Protorthoptera and Orthoptera. Psyche 73(1):46-88 [C. Labandeira/C. Labandeira/M. Clapham]
4986ETE 698F. M. Carpenter. 1976. The Lower Permian Insects of Kansas: Part 12. Protorthoptera (continued), Neuroptera, Additional Palaeodictyoptera & Families of Uncertain Position. Psyche 83:336-376 [C. Labandeira/C. Labandeira/R. Day]
45492 J. Kukalová-Peck. 1971. The structure of Dunbaria (Palaeodictyoptera). Psyche 78:306-318 [M. Clapham/M. Clapham]
74929 P. E. Raymond. 1944. Late Paleozoic xiphosurans. Bulletin of the Museum of Comparative Zoology 94:475-508 [M. Clapham/M. Clapham]
67816 P. Vršanský, H. Sendi, D. Aristov, G. Bechly, P. Müller, S. Ellenberger, D. Azar, K. Ueda, P. Barna and T. Garcia. 2019. Ancient roaches further exemplify 'no land return' in aquatic insects. Gondwana Research 68:22-33 [M. Clapham/M. Clapham/M. Clapham]