PBDB Collection

Klein Lehmhagen pit, Grimmen (LGA collection): Early/Lower Toarcian, Germany
collected by Ansorge

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Insecta - Eoblattida - Blattogryllidae

	Griphopteron molle Handlirsch 1939	Ansorge 1996	1 specimen
	LGA 850 (1 measurement)
Insecta - Diptera - Psychodidae

	Nannotanyderus krzeminskii n. gen., n. sp. Ansorge 1994	Ansorge 1994	2 specimens
	LGA 1145, 544 (2 measurements)

	Nannotanyderus grimmenensis n. sp. Ansorge and Krzemiński 2002	Ansorge and Krzemiński 2002	1 specimen
	LGA 2222 (1 measurement)

	Liassopsychodina pommerana n. gen., n. sp. Ansorge 1994	Ansorge 1994	2 specimens
	LGA 457, 752 (1 measurement)

	Tanypsycha connexa n. gen., n. sp. Ansorge 1994	Ansorge 1994	1 specimen
	LGA 1233 (1 measurement)
Insecta - Diptera - Ptychopteridae

	Eoptychoptera liasina (Handlirsch 1906)	Ansorge 1996	1 specimen
	synonym of Eoptychoptera simplex
	LGA 1368 (1 measurement)
Insecta - Diptera - Chironomidae

	Aenne liasina n. gen., n. sp. Ansorge 1999	Ansorge 1999	4 specimens
	LGA 1474, 1486, 1894, 1956 (3 measurements)

	Podonominae indet. Thienemann and Edwards 1837	Ansorge 1996	1 specimen
	LGA 1003 (2 measurements)

	? Podonomius tumidus n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 1034 (2 measurements)

	? Oryctochlus toarciensis n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 835 (1 measurement)
Insecta - Diptera - Trichoceridae

	Trichoceridae indet. Kertész 1902	Krzemińska et al. 2009	1 specimen
	LGA 1900

	Mailotrichocera mikereichi Krzemińska et al. 2009	Krzemińska et al. 2009	2 specimens
	LGA 79, 2005 (1 measurement)

	Mailotrichocera zessini n. sp. Krzemińska et al. 2009	Krzemińska et al. 2009	1 specimen
	LGA 611 (1 measurement)

	Mailotrichocera variabilis Krzemińska et al. 2009	Krzemińska et al. 2009	5 specimens
	LGA 288, 681, 1100, 1163, 2309 (3 measurements)
Insecta - Diptera - Heterorhyphidae

	Heterorhyphus triangularis n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 380 (1 measurement)
Insecta - Diptera - Anisopodidae

	Mesorhyphus zirzipanus n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 579 (1 measurement)
Insecta - Diptera - Antefungivoridae

	Archibio mycetophilinus Handlirsch 1939	Ansorge 1996	3 specimens
	LGA 214, 840, 988 (2 measurements)

	Antefungivora germanica n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 394 (1 measurement)
Insecta - Diptera - Pleciofungivoridae

	Pleciofungivora liassica n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 1045 (1 measurement)
Insecta - Diptera - Protorhyphidae

	Protorhyphus simplex (Geinitz 1888)	Ansorge 1996	7 specimens
	LGA 533, 571, 744, 998, 1002, 1007, 1294 (2 measurements)

	Archirhyphus geinitzi Handlirsch 1939	Ansorge 1996	1 specimen
	LGA 999 (1 measurement)
Insecta - Diptera

	Asilomorpha informal sp 1	Ansorge 1996	1 specimen
	LGA 690 (1 measurement)

	Asilomorpha informal sp 2	Ansorge 1996	1 specimen
	LGA 429 (1 measurement)
Insecta - Diptera - Rhagionidae

	Grimmyia baltica n. gen., n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 950 (1 measurement)

	Palaeobrachyceron willmanni n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 269 (1 measurement)
Insecta - Diptera - Eoditomyidae

	Eoditomyia primitiva n. gen., n. sp. Ansorge 1996	Ansorge 1996	2 specimens
	LGA 1305, 836 (2 measurements)
Insecta - Diptera - Protopleciidae

	Protoplecia liasina (Geinitz 1884)	Ansorge 1996	1 specimen
	LGA 1180 (1 measurement)

	Protoplecia klafacki n. sp. Ansorge 1996	Ansorge 1996	2 specimens
	LGA 892, 924 (2 measurements)
Insecta - Diptera

	Archipleciomima sp. Rohdendorf 1962	Ansorge 1996	1 specimen
	LGA 1203 (1 measurement)

	Archipleciomima germanica n. sp. Krzemiński and Ansorge 2018	Kopeć et al. 2018	3 specimens
	LGA 1550, 1803, 2282
Insecta - Diptera - Chaoboridae

	Rhaetomyia herrigi n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 1189 (1 measurement)
Insecta - Blattodea - Blattulidae

	Blattula langfeldti (Geinitz 1880)	Vršanský and Ansorge 2007	29 specimens
	LGA 86, 96, 27, 331/1(2), 380, 425, 579, 594, 888, 893, 1077, 1201, 1275, 1372, 1529, 1578, 1739, 1820, 2169, 13, 118, 154, 880, 1328, 1435, 1519, 1842, 1887, 1926, 2006, 2015, 2196, 2356, 888, 1820

	Blattula dubia (Handlirsch 1939)	Vršanský and Ansorge 2007	1 specimen
	LGA 1725
Insecta - Blattodea - Mesoblattinidae

	Mesoblattina protypa (Geinitz 1880)	Vršanský and Ansorge 2007	2 specimens
	LGA 1320, 1942
Insecta - Plecoptera - Perlariopseidae

	Perlariopseidae indet. Sinitshenkova 1985	Ansorge 1996	1 specimen
	LGA 777

	Dicronemoura furcata n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 1278 (1 measurement)
Insecta - Aerophasmidae

	Durnovaria cf. parallela Whalley 1985	Ansorge 1996	2 specimens
	LGA 1086, 1323 (1 measurement)
Insecta - Orthoptera - Locustopsidae

	Locustopsis informal sp 1	Ansorge 1996	1 specimen
	LGA 1272 (1 measurement)

	Locustopsis informal sp 2	Ansorge 1996	1 specimen
	LGA 1144 (1 measurement)

	Locustopsis informal sp 3	Ansorge 1996	1 specimen
	LGA 1309 (1 measurement)

	Locustopsis gyra Zessin 1983	Ansorge 1996	1 specimen
	LGA 390 (1 measurement)

	Locustopsis kruegeri Zessin 1983	Zessin 1988	1 specimen
	recombined as Plesioschwinzia kruegeri
	LGA 100

	Locustopsis bernstorffi (Geinitz 1880)	Ansorge 1996	2 specimens
	recombined as Plesioschwinzia bernstorffi
	LGA 100, 296 (2 measurements)
Insecta - Orthoptera - Regiatidae

	Protochaeta lanceolata (Handlirsch 1939)	Ansorge 1996	1 specimen
	LGA 235 (1 measurement)

	Protochaeta masculina Handlirsch 1939	Ansorge 1996	11 specimens
	LGA 113, 304, 324, 497, 597, 686, 780, 937, 940, 1114, 1280 (1 measurement)
Insecta - Orthoptera - Protogryllidae

	Protogryllus sp. Handlirsch 1906	Ansorge 1996	13 specimens
	LGA 32, 51, 358, 406, 511, 604, 766, 887, 889, 1040, 1172, 1196, 1238 (1 measurement)

	Protogryllus dobbertinensis (Geinitz 1880)	Ansorge 1996	11 specimens
	recombined as Protogryllus (Protogryllus) dobbertinensis
	LGA 94, 244, 324, 395, 813, 818, 865, 932, 1113, 1136, 1267 (1 measurement)
Insecta - Orthoptera - Elcanidae

	Elcana minima Handlirsch 1906		1 specimen
	recombined as Panorpidium minima
	LGA 61/1/2

	Elcana geinitzi (Heer 1880)	Ansorge 1996	1 specimen
	recombined as Panorpidium geinitzi
	LGA 445 (4 measurements)

	Elcana media Handlirsch 1906		4 specimens
	recombined as Panorpidium media
	LGA 4/1/2, 58/1/2, 63, 402/1/2

	Elcana magna Handlirsch 1906		2 specimens
	recombined as Panorpidium magna
	LGA 34/1/2, 62 (2 measurements)

	Parelcana tenuis Handlirsch 1906	Ansorge 1996	2 specimens
	LGA 487, 1351 (1 measurement)
Insecta - Grylloblattodea - Bajanzhargalanidae

	Nele jurassica n. gen., n. sp. Ansorge 1996	Ansorge 1996	4 specimens
	LGA 1310, 1111, 211, 977 (1 measurement)
Insecta - Coleoptera

	Coleoptera indet. Linnaeus 1758	Ansorge 2003	1 specimen
	LGA 1230
Insecta - Coleoptera - Trachypachidae

	Liassodites obsti n. gen., n. sp. Kirejtshuk and Ansorge 2022	Kirejtshuk and Ansorge 2022	1 specimen
	LGA 2850 (2 measurements)
Insecta - Neuroptera - Prohemerobiidae

	Actinophlebia sp. Handlirsch 1906	Ansorge 1996	2 specimens
	LGA 1080, 324 (2 measurements)

	Sibithone prodroma (Handlirsch 1906)	Ansorge 1996	3 specimens
	original and current combination Prohemerobius prodromus
	LGA 1001, 607, 452 (1 measurement)

	Prohemerobius latus Handlirsch 1920	Ansorge 1996	1 specimen
	LGA 124 (1 measurement)
Insecta - Neuroptera - Ithonidae

	Polyosmylus excelsus n. gen., n. sp. Ansorge 1996	Ansorge 1996	2 specimens
	LGA 1033, 540 (1 measurement)
Insecta - Neuroptera - Osmylidae

	Mesosmylina falciferum n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	corrected as Mesosmylina falcifera
	LGA 1295 (1 measurement)
Insecta - Necrotrichoptera - Necrotauliidae

	Mesotrichopteridium intermedium (Handlirsch 1906)	Ansorge 2002	8 specimens
	LGA 348, 533, 675, 789, 830, 874, 784, 1425 (4 measurements)

	Necrotaulius parvulus (Geinitz 1884)	Ansorge 2002	65 specimens
	(11 measurements)
Insecta - Trichoptera - Philopotamidae

	Liadotaulius maior (Handlirsch 1906)	Ansorge 2002	13 specimens
	LGA 290, 458, 626, 649, 1995, 2040, 2188, 220, 1198, 1710, 672, 713, 934 (10 measurements)
Insecta - Lepidoptera - Eolepidopterigidae

	Lepidoptera indet. Linnaeus 1758	Ansorge 1996	1 specimen
	LGA 968
	= ? Eolepidopterigidae indet. Rasnitsyn 1983	Zhang et al. 2018

	Micropterygidae indet. (Cotes 1889)	Ansorge 2003	1 specimen
	LGA 1500
	= ? Eolepidopterigidae indet. Rasnitsyn 1983	Zhang et al. 2018

	? Eolepidopterigidae indet. Rasnitsyn 1983	Zhang et al. 2018	1 specimen
	LGA 2150
Insecta - Mecoptera - Pseudopolycentropodidae

	Pseudopolycentropus sp. Handlirsch 1906	Ansorge 1996	2 specimens
	LGA 769, 993

	Pseudopolycentropus triangularis Handlirsch 1920	Ansorge 1996	6 specimens
	LGA 636, 925, 20, 738, 795, 1151 (2 measurements)
Insecta - Mecoptera - Orthophlebiidae

	Orthophlebia cf. elongata Handlirsch 1939	Ansorge 1996	1 specimen
	LGA 737 (1 measurement)

	Orthophlebia germanica Handlirsch 1906	Ansorge 1996	2 specimens
	LGA 811, 1362 (2 measurements)

	Orthophlebia brunsvicensis Bode 1905	Ansorge 1996	1 specimen
	LGA 790 (1 measurement)
Insecta - Mecoptera - Cimbrophlebiidae

	Cimbrophlebiidae indet. Willmann 1977	Ansorge 2003	1 specimen
	LGA 1608
Insecta - Mecoptera - Bittacidae

	Bittacidae indet. Handlirsch 1906	Ansorge 1996	1 specimen
	LGA 500 (1 measurement)

	Mesobittacus minutus Handlirsch 1939	Ansorge 1996	4 specimens
	LGA 133, 876, 1269, 1298 (2 measurements)

	Neorthophlebia maculipennis Handlirsch 1906	Ansorge 1996	1 specimen
	LGA 1369 (1 measurement)
Insecta - Hymenoptera - Karatavitidae

	Grimmaratavites mirabilis n. gen., n. sp. Rasnitsyn et al. 2006	Rasnitsyn et al. 2006	1 specimen
	LGA 2510 (2 measurements)
Insecta - Hemiptera - Liadopsyllidae

	Liadopsylla obtusa n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 731 (1 measurement)
Insecta - Hemiptera - Protopsyllidiidae

	Aphidulum ciliatum n. sp. Ansorge 1996	Ansorge 1996	3 specimens
	LGA 264, 641, 931 (1 measurement)

	Toarcopsyllidium ernsti n. gen., n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 1185 (1 measurement)
Insecta - Hemiptera - Oviparosiphidae

	Grimmenaphis magnifica n. gen., n. sp. Ansorge 1996	Ansorge 1996	2 specimens
	LGA 1313, 903 (1 measurement)
Insecta - Hemiptera - Archiconiopterygidae

	Archiconiopteryx liasina (Handlirsch 1906)	Ansorge 1996	3 specimens
	LGA 620, 666, 747 (2 measurements)
Insecta - Hemiptera

	Heteroptera indet. Latreille 1810	Ansorge 2003	1 specimen
	LGA 936
Insecta - Hemiptera - Archijassidae

	Archijassidae indet. Becker-Migdisova 1962	Ansorge 1996	4 specimens
	LGA 240, 380, 384, 701

	Archijassus heeri (Geinitz 1880)	Ansorge 1996	8 specimens
	LGA 505, 640, 757, 1204, 1288, 1340, 1300, 108 (2 measurements)

	Mesoledra pachyneura (Handlirsch 1939)	Ansorge 1996	7 specimens
	LGA 138, 206, 742, 1050, 1086, 1299, 927 (1 measurement)

	Ardela grimmenensis n. gen., n. sp. Ansorge 1996	Ansorge 1996	10 specimens
	LGA 1348, 673, 856, 1104, 1216, 505, 350, 1325, 163, 266 (2 measurements)
Insecta - Hemiptera - Hylicellidae

	Liassocercopis schnicki n. gen., n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 814 (1 measurement)
Insecta - Hemiptera - Procercopidae

	Procercopidae indet. Handlirsch 1906	Ansorge 1996	1 specimen
	LGA 676 (1 measurement)

	Procercopina frenzeli n. sp. Ansorge 1996	Ansorge 1996	2 specimens
	LGA 565, 380 (2 measurements)
Insecta - Hemiptera - Fulgoridiidae

	Fulgoridium informal sp 1	Ansorge 1996	1 specimen
	LGA 820 (1 measurement)

	Fulgoridium informal sp 2	Ansorge 1996	1 specimen
	LGA 123 (1 measurement)

	Fulgoridium sp. Handlirsch 1906	Ansorge 1996	3 specimens
	LGA 569, 749, 1183
Insecta - Hemiptera

	Fulgoridulum egens Handlirsch 1939	Ansorge 1996	18 specimens
	LGA 70, 89, 198, 714, 1313, 876, 1206, 628, 893, 1349, 920, 899, 896, 893, 1303, 920, 896, 1313 (1 measurement)
Insecta - Lophioneurida - Lophioneuridae

	Undacypha europaea n. sp. Ansorge 1996	Ansorge 1996	17 specimens
	LGA 233, 223, 292, 378, 490, 645, 746, 871, 883, 967, 983, 1274, 1333, 1342, 1339, 1364, 1365 (2 measurements)
Insecta - Permopsocida - Psocidiidae

	Liassopsocus lanceolatus n. gen., n. sp. Ansorge 1996	Ansorge 1996	2 specimens
	LGA 1246, 997 (1 measurement)
Insecta - Permopsocida - Archipsyllidae

	Archipsylla primitiva Handlirsch 1906	Ansorge 1996	1 specimen
	LGA 1229
Insecta - Odonata - Liassogomphidae

	Liassogomphidae informal indet. 1 Tillyard 1935	Ansorge 1996	1 specimen
	LGA 922

	Liassogomphidae informal indet. 2 Tillyard 1935	Ansorge 1996	1 specimen
	LGA 349
Insecta - Odonata - Liassophlebiidae

	Grimmenopteron elegantulum n. gen., n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 345 (1 measurement)
Insecta - Odonata - Heterophlebiidae

	Trigonophlebia zessini n. gen., n. sp. Ansorge 1996	Ansorge 1996	1 specimen
	LGA 739 (1 measurement)

	Heterophlebia buckmani (Brodie 1845)	Ansorge 1996	1 specimen
	LGA 46 (1 measurement)
Insecta - Odonata - Asiopteridae

	Oreopteridae indet. Pritykina 1968	Ansorge 1996	3 specimens
	synonym of Asiopteridae
	LGA 250, 712, 1358 (1 measurement)

	Sphenophlebia pommerana Ansorge 1996	Ansorge 1996	3 specimens
	recombined as Turanopteron pommerana
	LGA 480, 510, 1197 (1 measurement)
Insecta - Protomyrmeleontidae

	Protomyrmeleon brunonis Geinitz 1888	Ansorge 1996	4 specimens
	LGA 621, 815, 988, 1000 (3 measurements)

	Zirzipanagrion quadriordinum n. gen., n. sp. Zessin 1991	Zessin 1991	1 specimen
	recombined as Protomyrmeleon quadriordinum
	LGA 162/1 (1 measurement)

	Obotritagrion tenuiformum Zessin 1991	Ansorge 1996	1 specimen
	LGA 876 (1 measurement)
Insecta - Ephemeroptera

	Ephemeroptera indet. Hyatt and Arms 1890	Ansorge 1996	1 specimen
	LGA 1502
Arachnida - Araneae

	Seppo koponeni n. gen., n. sp. Selden and Dunlop 2014	Selden and Dunlop 2014	1 specimen
	MB.A 2966
see common names

Geography

Country:	Germany	State/province:	Mecklenburg-Vorpommern	County:	Röstock
Coordinates:	54.1° North, 13.1° East (view map)
Paleocoordinates:	43.7° North, 18.9° East
Basis of coordinate:	based on political unit
Altitude:	8 meters

Time

Period:	Jurassic	Epoch:	Early/Lower Jurassic
Stage:	Toarcian	10 m.y. bin:	Jurassic 3
Key time interval:	Early/Lower Toarcian	Zone:	Dactylioceras tenuicostatum
Age range of interval:	184.20000 - 182.90000 m.y. ago

Stratigraphy

Stratigraphic resolution:	bed
Stratigraphy comments: unnamed "bituminous shale bed" equivalent to Posidonian Shale (Posidonienschiefer); Lias epsilon level = lowest Toarcian; also lower part of Harpoceras falcifer Zone

Lithology and environment

Primary lithology:	lithified carbonaceous "shale"
Lithology description: bituminous shale ("bituminösen Schiefer")
Environment:	offshore indet.	Tectonic setting:	passive margin

Taphonomy

Modes of preservation:	adpression,concretion
Size of fossils:	macrofossils,mesofossils
Preservation of anatomical detail:	good

Collection methods and comments

Collection methods:	field collection,survey of museum collection
Reason for describing collection:	taxonomic analysis
Museum repositories:	MfN
Collectors:	Ansorge

Metadata

Database number:	114993
Authorizer:	M. Clapham	Enterer:	J. Karr, M. Clapham
Modifier:	M. Clapham	Research group:	paleoentomology
Created:	2011-08-26 14:31:37	Last modified:	2023-05-27 16:04:08
Access level:	the public	Released:	2011-08-26 14:31:37
Creative Commons license:	CC BY

Reference information

Primary reference:

35779.

W. Zessin. 1987. Variabilität, Merkmalswandel und Phylogenie der Elcanidae im Jungpaläozoikum und Mesozoikum und die Phylogenie der Ensifera. Deutsche Entomologische Zeitschrift 34(1-3):1-76 [M. Clapham/J. Karr/J. Karr]

Secondary references:

39822		J. Ansorge. 1994. Tanyderidae and Psychodidae (Insecta: Diptera) from the Lower Jurassic of northeastern Germany. Paläontologische Zeitschrift 68:199-210 [M. Clapham/M. Clapham/M. Clapham]
40666		J. Ansorge. 1996. Insekten aus dem oberen Lias von Grimmen (Vorpommern, Norddeutschland). Neue Paläontologische Abhandlungen 2:1-132 [M. Clapham/J. Karr]
48324		J. Ansorge. 1999. Aenne liasina gen. et sp. n. - the most primitive non biting midge (Diptera: Chironomidae: Aenninae subfam. n.) - from the Lower Jurassic of Germany. Polskie Pismo Entomologiczne 68:431-443 [M. Clapham/M. Clapham]
48326		J. Ansorge. 2002. Revision of the "Trichoptera" described by Geinitz and Handlirsch from the Lower Toarcian of Dobbertin (Germany) based on new material. Proceedings of the 10th International Symposium on Trichoptera - Nova Supplementa Entomologica 15:55-74 [M. Clapham/M. Clapham]
48323		J. Ansorge. 2003. Insects from the Lower Toarcian of middle Europe and England. Acta Zoologica Cracoviensia 46(Suppl):291-310 [M. Clapham/M. Clapham/M. Clapham]
51695		J. Ansorge and W. Krzemiński. 2002. Lower Jurassic tanyderids (Diptera: Tanyderidae) from Germany. Studia Dipterologica 9:21-29 [M. Clapham/M. Clapham]
84824		A. G. Kirejtshuk and J. Ansorge. 2022. An extraordinarily preserved new genus and species of Trachypachidae (Coleoptera, Adephaga) from the Early Jurassic of Germany and a review of fossil trachypachid genera. Historical Biology 35:958-975 [M. Clapham/M. Clapham]
64881		K. Kopeć, A. Soszyńska-Maj, A. Gehler, J. Ansorge, and W. Krzemiński. 2018. Mecoptera and Diptera from the early Toarcian (Early Jurassic) deposits of Wolfsburg – Große Kley (Lower Saxony, Germany). Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107:163-171 [M. Clapham/M. Clapham/M. Clapham]
51817		E. Krzemińska, W. Krzemiński, and C. Dahl. 2009. Monograph of Fossil Trichoceridae (Diptera): Over 180 Million Years of Evolution 1-171 [M. Clapham/M. Clapham]
40045		A. P. Rasnitsyn, J. Ansorge, and H. C. Zhang. 2006. Ancestry of the orussoid wasps, with description of three new genera and species of Karatavitidae (Hymenoptera = Vespida: Karatavitoidea stat. nov.). Insect Systematics & Evolution 37:179-190 [M. Clapham/M. Clapham/M. Clapham]
53833		P. A. Selden and J. A. Dunlop. 2014. The first fossil spider (Araneae: Palpimanoidea) from the Lower Jurassic (Grimmen, Germany). Zootaxa 3894:161-168 [M. Clapham/M. Clapham]
37861		P. Vršanský and J. Ansorge. 2007. Lower Jurassic cockroaches (Insecta: Blattaria ) from Germany and England. African Invertebrates 48(1):103-126 [M. Clapham/J. Karr/M. Clapham]
52427		W. Zessin. 1988. Neue Saltatoria (Insecta) aus dem Oberlias Mitteleuropas. Freiberger Forschungshefte C 419:107-121 [M. Clapham/M. Clapham]
39401		W. Zessin. 1991. Die Phylogenie der Protomyrmeleontidae unter einbeziehung neuer Oberliassischer funde (Odonata: Archizygoptera sens. nov.). Odonatologica 20(1):97-126 [M. Clapham/J. Karr/M. Clapham]
65500		Q. Q. Zhang, W. Mey, J. Ansorge, T. A. Starkey, L. T. McDonald, M. E. McNamara, E. A. Jarzembowski, W. Wichard, R. Kelly, X. Y. Ren, J. Chen, H. C. Zhang, and B. Wang. 2018. Fossil scales illuminate the early evolution of lepidopterans and structural colors. Science Advances 4:e1700988 [M. Clapham/M. Clapham]