Daohugou (CNU 2011 collection): Callovian - Oxfordian, China

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Arachnida - Araneae - Mongolarachnidae
Mongolarachne jurassica (Selden et al. 2011)
Selden et al. 2013 1 specimen
CNU-ARA-NN2011001
Insecta - Diptera - Ptychopteridae
Eoptychopterina postica n. sp. Liu et al. 2012
Liu et al. 2012 2 specimens
CNU-DIP-NN2011001, 002 (1 measurement)
Insecta - Diptera - Mesosciophilidae
Similsciophila singularis n. gen., n. sp. Shi et al. 2014
Shi et al. 2015 2 specimens
CNU-DIP-NN2011226, 147 (3 measurements)
Similsciophila sinuata Shi et al. 2014
Shi et al. 2015 1 specimen
CNU-DIP-NN2011511 (2 measurements)
Insecta - Odonata - Campterophlebiidae
Karatawia sinica n. sp. Li et al. 2012
Li et al. 2012 1 specimen
CNU-ODO-NN2011017 (2 measurements)
Angustiphlebia mirabilis n. gen., n. sp. Li et al. 2013
Li et al. 2013 1 specimen
CNU-ODO-NN2011016 (1 measurement)
Insecta - Hemiptera - Progonocimicidae
Cicadocoris varians n. sp. Dong et al. 2012
Dong et al. 2012 1 specimen
recombined as Mesocimex varians
CNU-PRO-NN2011079 (2 measurements)
Cicadocoris anisomeridis Dong et al. 2014
Dong et al. 2014 29 specimens
(2 measurements)
Cicadocoris assimilis n. sp. Dong et al. 2013
Dong et al. 2013 1 specimen
CNU-PRO-NN2011037 (2 measurements)
Insecta - Hemiptera - Protopsyllidiidae
Poljanka strigosa n. sp. Yang et al. 2013
Yang et al. 2013 2 specimens
CNU-PSY-NN2011021, 003 (2 measurements)
Poljanka hirsuta n. sp. Yang et al. 2012
Yang et al. 2012 2 specimens
CNU-PSY-NN2011008, 2011018 (4 measurements)
Insecta - Blattaria - Fuziidae
Colorifuzia agenora n. gen., n. sp. Wei et al. 2013
Wei et al. 2013 5 specimens
CNU-BLA-NN2011001, 003, 004, 015, 016 (2 measurements)
Parvifuzia peregrina n. sp. Wei et al. 2012
Wei et al. 2012 3 specimens
CNU-BLA-NN-2011055, 056, 057 (4 measurements)
Arcofuzia cana n. gen., n. sp. Wei et al. 2012
Wei et al. 2012 2 specimens
CNU-BLA-NN2011012, 058 (4 measurements)
Insecta - Grylloblattodea
Plesioblattogryllus minor Ren and Aristov 2011
Cui 2012 4 specimens
recombined as Aristoviblattogryllus minor
CNU-GRY-NN2011001, 002, 003, 004 (4 measurements)
Insecta - Grylloblattodea - Plesioblattogryllidae
Duoduo qianae n. gen., n. sp. Cui 2012
Cui 2012 1 specimen
recombined as Plesioblattogryllus qianae
CNU-GRY-NN2011005 (1 measurement)
Insecta - Orthoptera - Prophalangopsidae
Bacharaboilus curvus n. sp. Gu et al. 2022
Gu et al. 2022 2 specimens
CNU-ORT-NN2011037, 038 (1 measurement)
Insecta - Orthoptera - Locustopsidae
Locustopsis rhytofemoralis n. sp. Gu et al. 2016
Gu et al. 2016 1 specimen
CNU-ORT-NN2011010 (2 measurements)
Insecta - Neuroptera - Mesochrysopidae
Protoaristenymphes daohugouensis n. sp. Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011103 (2 measurements)
Insecta - Neuroptera - Osmylopsychopidae
Daopsychops bifasciatus n. sp. Peng et al. 2015
Peng et al. 2016 2 specimens
CNU-NEU-NN2011058, 217 (1 measurement)
Daopsychops dissectus n. gen., n. sp. Peng et al. 2015
Peng et al. 2016 1 specimen
CNU-NEU-NN2011059 (1 measurement)
Daopsychops cubitalis n. sp. Peng et al. 2015
Peng et al. 2016 1 specimen
CNU-NEU-NN2011432 (1 measurement)
Daopsychops clausus n. sp. Peng et al. 2015
Peng et al. 2016 1 specimen
CNU-NEU-NN2011061 (1 measurement)
Daopsychops inanis n. sp. Peng et al. 2015
Peng et al. 2016 3 specimens
CNU-NEU-NN2011435, 073, 064 (3 measurements)
Nematopsychops unicus n. gen., n. sp. Peng et al. 2015
Peng et al. 2016 1 specimen
CNU-NEU-NN2011086 (1 measurement)
Eupypsychops confinis n. gen., n. sp. Peng et al. 2015
Peng et al. 2016 1 specimen
CNU-NEU-NN2011063 (1 measurement)
Stenopteropsychops trifasciatus n. gen., n. sp. Peng et al. 2015
Peng et al. 2016 1 specimen
CNU-NEU-NN2011517 (1 measurement)
Ochropsychops multus n. gen., n. sp. Peng et al. 2015
Peng et al. 2016 1 specimen
CNU-NEU-NN2011055 (1 measurement)
Insecta - Neuroptera - Psychopsidae
Gigantopsychops reticulatus n. gen., n. sp. Peng et al. 2015
Peng et al. 2016 1 specimen
CNU-NEU-NN2011040 (1 measurement)
Insecta - Neuroptera - Nymphidae
Daonymphes bisulca n. gen., n. sp. Makarkin et al. 2013
Makarkin et al. 2013 1 specimen
CNU-NEU-NN2011119 (1 measurement)
Nymphites informal sp. A Haase 1890
Shi et al. 2013 1 specimen
CNU-NEU-NN2011118
Nymphites bimaculatus n. sp. Shi et al. 2013
Shi et al. 2013 2 specimens
CNU-NEU-NN2011116, 117 (2 measurements)
Insecta - Neuroptera - Berothidae
Sinosmylites sp. Hong 1983
1 specimen
(1 measurement)
Sinosmylites rasnitsyni n. sp. Makarkin et al. 2011
1 specimen
(2 measurements)
Sinosmylites fumosus n. sp. Makarkin et al. 2011
1 specimen
(1 measurement)
Insecta - Neuroptera - Mantispidae
Clavifemora rotundata n. gen., n. sp. Jepson et al. 2013
Jepson et al. 2013 1 specimen
CNU-NEU-NN2011001 (1 measurement)
Insecta - Neuroptera - Parakseneuridae
Pseudorapisma maculatum n. sp. Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011018 (1 measurement)
Pseudorapisma angustipenne n. sp. Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011014 (1 measurement)
Pseudorapisma jurassicum n. gen., n. sp. Yang et al. 2012
Yang et al. 2012 4 specimens
CNU-NEU-NN2011033, 006, 027, 013 (4 measurements)
Parakseneura informal sp. A Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011020
Parakseneura informal sp. B Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011012
Parakseneura informal sp. C Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011028
Parakseneura informal sp. D Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011025
Parakseneura curvivenis n. sp. Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011021
Parakseneura inflata n. sp. Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011010
Parakseneura nigrolinea n. sp. Yang et al. 2012
Yang et al. 2012 2 specimens
CNU-NEU-NN2011017, 011 (2 measurements)
Parakseneura albadelta n. sp. Yang et al. 2012
Yang et al. 2012 2 specimens
CNU-NEU-NN2011015, 022 (2 measurements)
Parakseneura cavomaculata n. sp. Yang et al. 2012
Yang et al. 2012 2 specimens
CNU-NEU-NN2011008, 007 (1 measurement)
Parakseneura albomacula n. sp. Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011029 (1 measurement)
Parakseneura undula n. sp. Yang et al. 2012
Yang et al. 2012 2 specimens
CNU-NEU-NN2011030, 031 (2 measurements)
Parakseneura directa n. sp. Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011016 (1 measurement)
Parakseneura metallica n. sp. Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011019 (1 measurement)
Parakseneura nigromacula n. gen., n. sp. Yang et al. 2012
Yang et al. 2012 2 specimens
CNU-NEU-NN2011009, 026 (2 measurements)
Parakseneura emarginata n. sp. Yang et al. 2012
Yang et al. 2012 1 specimen
CNU-NEU-NN2011024 (1 measurement)
Insecta - Megaloptera - Corydalidae
Eochauliodes striolatus n. gen., n. sp. Liu et al. 2012
Liu et al. 2012 8 specimens
CNU-MEG-NN2011004-011 (3 measurements)
Jurochauliodes ponomarenkoi Wang and Zhang 2010
Liu et al. 2012 4 specimens
CNU-MEG-NN2001001, 002, 003, 012 (2 measurements)
Insecta - Coleoptera - Lophocateridae
Sinopeltis jurassica n. gen., n. sp. Yu et al. 2012
Yu et al. 2012 1 specimen
CNU-COL-NN2011102 (2 measurements)
Sinopeltis amoena n. sp. Yu et al. 2012
Yu et al. 2012 1 specimen
CNU-COL-NN2011103 (2 measurements)
Insecta - Coleoptera - Trogossitidae
Latitergum glabrum n. gen., n. sp. Yu et al. 2014
Yu et al. 2014 2 specimens
recombined as Neolatitergum glabrum
CNU-COL-NN2011107, 108 (4 measurements)
Marginulatus venustus n. gen., n. sp. Yu et al. 2014
Yu et al. 2014 2 specimens
CNU-COL-NN2011104, 105 (4 measurements)
Insecta - Coleoptera - Nemonychidae
Sinoeuglypheus daohugouensis n. gen., n. sp. Yu et al. 2019
Yu et al. 2019 1 specimen
recombined as Belonotaris daohugouensis
CNU-COL-NN2011122 (1 measurement)
Insecta - Trichoptera - Rhyacophilidae
Declinimodus setulosus n. gen., n. sp. Gao et al. 2013
Gao et al. 2013 2 specimens
CNU-TRI-NN2011006, 007 (2 measurements)
Insecta - Trichoptera - Philopotamidae
Pulchercylindratus punctatus n. gen., n. sp. Gao et al. 2013
Gao et al. 2013 3 specimens
CNU-Tri-NN2011003, 004, 005 (6 measurements)
Insecta - Mecoptera - Choristopsychidae
Choristopsyche perfecta n. sp. Qiao et al. 2013
Qiao et al. 2013 3 specimens
CNU-MEC-NN2011082, 079, 084 (2 measurements)
Choristopsyche tenuinervis Martynov 1937
Qiao et al. 2013 6 specimens
CNU-MEC-NN2011070, 071, 075, 080, 083, 085 (2 measurements)
Choristopsyche asticta Qiao et al. 2013
Qiao et al. 2013 2 specimens
CNU-MEC-NN2011081, 086 (2 measurements)
Paristopsyche angelineae n. gen., n. sp. Qiao et al. 2013
Qiao et al. 2013 5 specimens
CNU-NEU-NN2011076, 069, 074, 077, 078 (4 measurements)
Insecta - Mecoptera - Cimbrophlebiidae
Bellicimbrophlebia cruciata Yang et al. 2013
Yang et al. 2013 3 specimens
CNU-M-NN2011011, 010, 007 (3 measurements)
Bellicimbrophlebia disvena Yang et al. 2013
Yang et al. 2013 2 specimens
CNU-M-NN2011001, 005 (2 measurements)
Insecta - Mecoptera - Orthophlebiidae
Orthophlebia riccardii n. sp. Petrulevičius and Ren 2012
Petrulevičius and Ren 2012 1 specimen
recombined as Gigaphlebia riccardii
CNU-MEC-NN2011001 (2 measurements)
Orthophlebia stigmosa n. sp. Qiao et al. 2012
Qiao et al. 2012 1 specimen
recombined as Longiphlebia stigmosa
CNU-MEC-NN2011061 (1 measurement)
Orthophlebia nervulosa n. sp. Qiao et al. 2012
Qiao et al. 2012 1 specimen
CNU-MEC-NN2011060 (2 measurements)
Insecta - Mecoptera - Mesopsychidae
Epicharmesopsyche pentavenulosa n. gen., n. sp. Shih et al. 2013
Shih et al. 2013 1 specimen
CNU-MEC-NN2011095 (2 measurements)
see common names

Geography
Country:China State/province:Nei Mongol
Coordinates: 41.3° North, 119.2° East (view map)
Paleocoordinates:42.9° North, 122.8° East
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Jurassic
10 m.y. bin:Jurassic 5
Key time interval:Callovian - Oxfordian
Age range of interval:165.30000 - 154.80000 m.y. ago
Stratigraphy
Formation:Jiulongshan Member:Daohugou
Stratigraphic resolution:group of beds
Stratigraphy comments: The age of the Daohugou fauna has been controversial. Liu et al. (2012) obtained U-Pb shrimp ages of 160.5 and 161.0 Myr on ash beds associated with fossils of the Daohugou fauna, suggesting a Callovian-Oxfordian age.
Lithology and environment
Primary lithology:tuffaceous lithified "shale"
Includes fossils?Y
Lithology description: "tuffaceous shale"; "freshwater shales interbedded with tuffs"
Environment:lacustrine - large Tectonic setting:rift
Geology comments: Basins characterized by flexure and early rifting (Li et al., 2004)
Taphonomy
Modes of preservation:mold/impression,adpression
Lagerst�tten type:conservation
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:excellent
Collection methods and comments
Collection methods:bulk,selective quarrying,mechanical
Reason for describing collection:taxonomic analysis
Collection method comments: Repository: Capital Normal University, Beijing, China.
Metadata
Database number:117610
Authorizer:M. Clapham Enterer:M. Clapham, J. Karr
Modifier:M. Clapham Research group:paleoentomology
Created:2011-09-25 07:31:09 Last modified:2023-05-27 10:47:30
Access level:the public Released:2011-09-25 07:31:09
Creative Commons license:CC BY
Reference information

Primary reference:
37846. V. N. Makarkin, Q. Yang, and D. Ren. 2011. Two new species of Sinosmylites Hong (Neuroptera, Berothidae) from the Middle Jurassic of China, with notes on Mesoberothidae. ZooKeys 130:199-215 [M. Clapham/M. Clapham]

Secondary references:

50181 Y. Y. Cui. 2012. New data on the Blattogryllidae-Plesioblattogryllidae-Grylloblattidae complex (Insecta: Grylloblattida: Blattogryllopterida tax. n.). Arthropod Systematics & Phylogeny 70:167-180 [M. Clapham/M. Clapham]
54506 Q. P. Dong, Y. Z. Yao, and D. Ren. 2012. A new species of Progonocimicidae (Hemiptera: Coleorrhyncha) from northeastern China. Zootaxa 3495:73-78 [M. Clapham/M. Clapham]
46278 Q. P. Dong, Y. Z. Yao, and D. Ren. 2013. A new species of Progonocimicidae (Hemiptera, Coleorrhyncha) from the Middle Jurassic of China. Alcheringa 37:31-37 [M. Clapham/M. Clapham]
52743 Q. P. Dong, Y. Z. Yao, and D. Ren. 2014. New fossil Progonocimicidae (Hemiptera: Coleorrhyncha: Progonocimicoidea) from the Upper Mesozoic of northeastern China, with a phylogeny of Coleorrhyncha. Systematic Entomology 39:773-782 [M. Clapham/M. Clapham]
49485 Y. Gao, Y. Z. Yao, and D. Ren. 2013. New genus and species of Rhyacophilidae (Insecta: Trichoptera) from the Middle Jurassic of China. Acta Geologica Sinica 87:1495-1500 [M. Clapham/M. Clapham]
45183 Y. Gao, Y. Z. Yao, and D. Ren. 2013. A new Middle Jurassic caddisfly (Trichoptera, Hydrobiosidae) from China. Fossil Record 16:111-116 [M. Clapham/M. Clapham]
81881 J. J. Gu, Z. Q. Xu, R. Huang, H. J. Wang, Y. L. Yue and D. Ren. 2022. Systematic significance of wing morphology in extinct Prophalangopsidae (Insecta, Ensifera) revealed by geometric morphometrics and description of two new species. Journal of Systematic Palaeontology 19:1587-1599 [M. Clapham/M. Clapham]
60392 J. J. Gu, Y. L. Yue, F. M. Shi, H. Tian, and D. Ren. 2016. First Jurassic grasshopper (Insecta, Caelifera) from China. Zootaxa 4169:377-380 [M. Clapham/M. Clapham]
46738 J. E. Jepson, S. W. Heads, V. N. Makarkin and D. Ren. 2013. New fossil mantidflies (Insecta: Neuroptera: Mantispidae) from the Mesozoic of north-eastern China. Palaeontology 56:603-613 [M. Clapham/M. Clapham/M. Clapham]
53887 Y. J. Li, A. Nel, D. Ren, B. L. Zhang, and H. Pang. 2012. Reassessment of the Jurassic damsel-dragonfly genus Karatawia (Odonata: Campterophlebiidae). Zootaxa 3417:64-68 [M. Clapham/M. Clapham]
46087 Y. J. Li, A. Nel, D. Ren and H. Pang. 2013. A new damsel-dragonfly from the Mesozoic of China with a hook-like male anal angle (Odonata: Isophlebioptera: Campterophlebiidae). Journal of Natural History 1-6 [M. Clapham/J. Karr]
44269 L. X. Liu, C. K. Shih, and D. Ren. 2012. Two new species of Ptychopteridae and Trichoceridae from the Middle Jurassic of northeastern China (Insecta: Diptera: Nematocera). Zootaxa 3501:55-62 [M. Clapham/M. Clapham]
42265 X. Y. Liu, Y. J. Wang, C. K. Shih, D. Ren, and D. Yang. 2012. Early evolution and historical biogeography of fishflies (Megaloptera: Chauliodinae): implications from a phylogeny combining fossil and extant taxa. PLoS One 7(7):e40345 [M. Clapham/M. Clapham/M. Clapham]
47995 V. N. Makarkin, Q. Yang, C. F. Shi and D. Ren. 2013. The presence of the recurrent veinlet in the Middle Jurassic Nymphidae (Neuroptera): a unique character condition in Myrmeleontoidea. ZooKeys 325:1-20 [M. Clapham/M. Clapham]
58382 Y. Y. Peng, V. N. Makarkin, and D. Ren. 2016. Diverse new Middle Jurassic Osmylopsychopidae (Neuroptera) from China shed light on the classification of psychopsoids. Journal of Systematic Palaeontology 14:261-295 [M. Clapham/M. Clapham]
42709 J. F. Petrulevičius and D. Ren. 2012. A new species of “Orthophlebiidae” (Insecta: Mecoptera) from the Middle Jurassic of Inner Mongolia, China. Revue de Paléobiologie, Genève, Vol. spéc 11:311-315 [M. Clapham/M. Clapham/M. Clapham]
43750 X. Qiao, C. K. Shih, and D. Ren. 2012. Two new Middle Jurassic species of orthophlebiids (Insecta: Mecoptera) from Inner Mongolia, China. Alcheringa 36:467-472 [M. Clapham/M. Clapham]
50245 X. Qiao, C. K. Shih, J. F. Petrulevičius and D. Ren. 2013. Fossils from the Middle Jurassic of China shed light on morphology of Choristopsychidae (Insecta, Mecoptera). ZooKeys 318:91-111 [M. Clapham/M. Clapham/M. Clapham]
49560 P. A. Selden, C. K. Shih, and D. Ren. 2013. A giant spider from the Jurassic of China reveals greater diversity of the orbicularian stem group. Naturwissenschaften 100:1171-1181 [M. Clapham/M. Clapham]
50978 C. F. Shi, V. N. Makarkin, Q. Yang, S. B. Archibald, and D. Ren. 2013. New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700:393-410 [M. Clapham/M. Clapham]
55391 G. F. Shi, C. K. Shih, and D. Ren. 2015. A new genus with two new species of mesosciophilids from the Middle Jurassic of China (Diptera: Nematocera: Mesosciophilidae). Journal of Natural History 49:1147-1158 [M. Clapham/M. Clapham]
48500 C. K. Shih, X. Qiao, C. C. Labandeira and D. Ren. 2013. A new mesopsychid (Mecoptera) from the Middle Jurassic of northeastern China. Acta Geologica Sinica 87:1235-1241 [M. Clapham/M. Clapham]
65057 A. Soszyńska-Maj, W. Krzemiński, K. Kopeć, Y. Z. Cao, and D. Ren. 2018. Large Jurassic scorpionflies belonging to a new subfamily of the family Orthophlebiidae (Mecoptera). Annales Zoologici 68:85-92 [M. Clapham/M. Clapham/M. Clapham]
42781 D. D. Wei, J. H. Liang, and D. Ren. 2012. A new species of Fuziidae (Insecta, Blattida) from the Inner Mongolia, China. ZooKeys 217:53-61 [M. Clapham/M. Clapham]
47416 D. D. Wei, J. H. Liang, and D. Ren. 2013. A new fossil genus of Fuziidae (Insecta, Blattida) from the Middle Jurassic of Jiulongshan Formation, China. Geodiversitas 35:335-343 [M. Clapham/M. Clapham]
44267 D. D. Wei, C. K. Shih, and D. Ren. 2012. Arcofuzia cana gen. et sp. n. (Insecta, Blattaria, Fuziidae) from the Middle Jurassic sediments of Inner Mongolia, China. Zootaxa 3597:25-32 [M. Clapham/M. Clapham]
42226 G. Yang, Y. Yao, and D. Ren. 2012. A new species of Protopsyllidiidae (Hemiptera, Sternorrhyncha) from the Middle Jurassic of China. Zootaxa 3274:36-42 [M. Clapham/J. Karr]
46281 G. Yang, Y. Z. Yao, and D. Ren. 2013. Poljanka strigosa, a new species of Protopsyllidiidae (Hemiptera, Sternorrhyncha) from the Middle Jurassic of China. Alcheringa 37:125-130 [M. Clapham/M. Clapham]
44246 Q. Yang, V. N. Makarkin, and D. Ren. 2012. New fossil Mesochrysopidae (Neuroptera) from the Mesozoic of China. Zootaxa 3597:1-14 [M. Clapham/M. Clapham]
43113 Q. Yang, V. N. Makarkin, S. L. Winterton, A. V. Khramov, and D. Ren. 2012. A remarkable new family of Jurassic insects (Neuroptera) with primitive wing venation and its phylogenetic position in Neuropterida. PLoS One 7(9):e44762 [M. Clapham/M. Clapham/M. Clapham]
47581 X. G. Yang, C. K. Shih, and D. Ren. 2013. New fossil hangingflies (Insecta: Mecoptera: Raptipedia: Cimbrophlebiidae) from the Middle Jurassic of Inner Mongolia, China. Palaeontology 56:711-726 [M. Clapham/M. Clapham]
70681 Y. L. Yu, S. R. Davis, C. K. Shih, D. Ren, and H. Pang. 2019. The earliest fossil record of Belidae and its implications for the early evolution of Curculionoidea (Coleoptera). Journal of Systematic Palaeontology 17:2105-2117 [M. Clapham/M. Clapham/M. Clapham]
51072 Y. L. Yu, R. A. B. Leschen, A., D. Ren, and H. Pang. 2012. The first fossil bark-gnawing beetle from the Middle Jurassic of Inner Mongolia, China (Coleoptera: Trogossitidae). Annales Zoologici 62:245-252 [M. Clapham/M. Clapham/M. Clapham]
53916 Y. L. Yu, A. Ślipiński, R. A. B. Leschen, D. Ren, and H. Pang. 2014. Enigmatic Mesozoic bark-gnawing beetles (Coleoptera: Trogossitidae) from the Jiulongshan Formation in China. Annales Zoologici 64:667-676 [M. Clapham/M. Clapham/M. Clapham]