S7 locality, Tulip Bed community, Fossil Gully, Mount Stephen: Delamaran, Canada
collected by Royal Ontario Museum 1983
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
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unclassified
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Marpolia spissa
Walcott 1919
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O’Brien and Caron 2016 | 693 specimens | ||||||||
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Florideophyceae
- Ceramiales
- Rhodomelaceae
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Wahpia insolens
Walcott 1919
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O’Brien and Caron 2016 | |||||||||
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unclassified
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Laenigma aff. striatum
Krassilov 2005
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O’Brien and Caron 2016 | |||||||||
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Siphusauctidae
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Siphusauctum gregarium n. gen., n. sp.
O’Brien and Caron 2012
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O’Brien and Caron 2012 | 1133 specimens | ||||||||
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Eocrinoidea
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Eocrinoidea indet.
Jaekel 1918
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O’Brien and Caron 2016 | |||||||||
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unclassified
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Priapulida indet.
Théel 1906
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O’Brien and Caron 2016 | |||||||||
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Ottoidae
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Ottoia prolifica
Walcott 1911
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O’Brien and Caron 2016 | |||||||||
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Selkirkiimorpha
- Selkirkiidae
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Selkirkia columbia
Conway Morris 1977
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O’Brien and Caron 2016 | 3985 specimens | ||||||||
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Luolishaniida
- Collinsovermidae
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Entothyreos synnaustrus n. gen., n. sp.
Aria and Caron 2024
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Aria and Caron 2024 | 51 specimens | ||||||||
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Radiodonta
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Caryosyntrips serratus
Daley and Budd 2010
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Daley and Budd 2010 | |||||||||
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Laggania nathorsti
|
Daley and Budd 2010 | |||||||||
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Radiodonta
- Amplectobeluidae
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Amplectobelua stephenensis
Daley and Budd 2010
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Daley and Budd 2010 | |||||||||
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Radiodonta
- Hurdiidae
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Laggania cambria
Walcott 1911
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O’Brien and Caron 2016 | |||||||||
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Hurdia victoria
Walcott 1912
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Daley and Budd 2010 | |||||||||
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Radiodonta
- Anomalocarididae
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Anomalocaris canadensis
Whiteaves 1892
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Daley and Budd 2010 | |||||||||
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Dinocarida
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Dinocarida indet.
Collins 1996
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O’Brien and Caron 2016 | |||||||||
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Agnostida
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Agnostida indet.
Salter 1864
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O’Brien and Caron 2016 | |||||||||
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Agnostida
- Peronopsidae
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Ptychagnostus praecurrens
(Westergård 1936)
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O’Brien and Caron 2016 | |||||||||
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unclassified
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Molaria sp.
|
O’Brien and Caron 2016 | |||||||||
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Sidneyia sp.
Walcott 1911
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O’Brien and Caron 2016 | |||||||||
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Trilobita
- Corynexochida
- Dorypygidae
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Olenoides serratus
(Rominger 1887)
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O’Brien and Caron 2016 | |||||||||
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Kootenia sp.
(Walcott 1899)
|
O’Brien and Caron 2016 | |||||||||
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Ogygopsis klotzi
(Rominger 1887)
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O’Brien and Caron 2016 | |||||||||
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Trilobita
- Corynexochida
- Oryctocephalidae
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Oryctocara geikiei
Walcott 1908
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O’Brien and Caron 2016 | |||||||||
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Oryctocephalus sp.
Walcott 1886
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O’Brien and Caron 2016 | |||||||||
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Trilobita
- Corynexochida
- Zacanthoididae
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Zacanthoides romingeri
|
O’Brien and Caron 2016 | |||||||||
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Trilobita
- Corynexochida
- Dolichometopidae
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Polypleuraspis sp.
Poulson 1927
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O’Brien and Caron 2016 | |||||||||
| as "Polyplearaspis", presumably a mis-spelling | ||||||||||
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cf. Anoria sp.
Walcott 1924
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O’Brien and Caron 2016 | |||||||||
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Poliella prima
(Walcott 1908)
|
O’Brien and Caron 2016 | |||||||||
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Glossopleura sp.
Poulsen 1927
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O’Brien and Caron 2016 | |||||||||
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Trilobita
- Ptychopariida
- Ptychopariidae
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Elrathina sp.
Resser 1937
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O’Brien and Caron 2016 | |||||||||
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Trilobita
- Ptychopariida
- Alokistocaridae
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Ehmaniella sp.
Resser 1937
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O’Brien and Caron 2016 | |||||||||
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Alokistocare sp.
Lorenz 1906
|
O’Brien and Caron 2016 | |||||||||
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Nektaspidida
- Naraoiidae
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Naraoia spp.
Walcott 1912
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O’Brien and Caron 2016 | |||||||||
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unclassified
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Helmetia expansa
|
O’Brien and Caron 2016 | |||||||||
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Isoxyida
- Isoxyidae
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Isoxys sp.
Walcott 1890
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O’Brien and Caron 2016 | |||||||||
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Mollisoniida
- Mollisoniidae
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Mollisonia sp.
Walcott 1912
|
O’Brien and Caron 2016 | |||||||||
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Hymenocarina
- Perspicarididae
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Perspicaris sp.
Briggs 1977
|
O’Brien and Caron 2016 | |||||||||
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Canadaspidida
- Canadaspididae
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Canadaspis perfecta
(Walcott 1912)
|
O’Brien and Caron 2016 | 4525 specimens | ||||||||
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Waptiida
- Waptiidae
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Waptia sp.
Walcott 1912
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O’Brien and Caron 2016 | |||||||||
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unclassified
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Branchiocaris sp.
Briggs 1976
|
O’Brien and Caron 2016 | |||||||||
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Nereocaris exilis
Legg et al. 2012
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O’Brien and Caron 2016 | |||||||||
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Megacheira
- Yohoiida
- Yohoiidae
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Yohoia tenuis
|
O’Brien and Caron 2016 | |||||||||
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Megacheira
- Leanchoiliida
- Leanchoiliidae
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? Leanchoilia cf. protogonia
Simonetta 1970
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O’Brien and Caron 2016 | |||||||||
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Alalcomenaeus cambricus
Simonetta 1970
|
O’Brien and Caron 2016 | |||||||||
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unclassified
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? Habelia brevicauda
|
O’Brien and Caron 2016 | |||||||||
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Odaraiida
- Pectocariidae
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Odaraia alata
|
O’Brien and Caron 2016 | |||||||||
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unclassified
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aff. Plenocaris aff. plena
|
O’Brien and Caron 2016 | |||||||||
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Bradoriida
- Svealutidae
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Liangshanella spp.
Huo 1956
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O’Brien and Caron 2016 | 6263 specimens | ||||||||
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Tuzoida
- Tuzoiidae
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Tuzoia burgessensis
Resser 1929
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O’Brien and Caron 2016 | |||||||||
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unclassified
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Hanburia gloriosa
|
O’Brien and Caron 2016 | |||||||||
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Hallucigeniidae
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Hallucigenia sp.
Conway Morris 1977
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O’Brien and Caron 2016 | |||||||||
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Tergomya
- Scenellidae
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Scenella amii
(Matthew 1902)
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O’Brien and Caron 2016 | |||||||||
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unclassified
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Odontogriphus omalus
Conway-Morris 1976
|
O’Brien and Caron 2016 | |||||||||
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Wiwaxia sp.
Walcott 1911
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O’Brien and Caron 2016 | |||||||||
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Orthrozanclidae
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Oikozetetes seilacheri
Conway Morris 1995
|
O’Brien and Caron 2016 | |||||||||
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Lingulata
- Acrotretida
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Acrothyra gregaria
|
O’Brien and Caron 2016 | |||||||||
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Lingulata
- Lingulida
- Obolidae
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Lingulella waptaensis
|
O’Brien and Caron 2016 | |||||||||
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Paterinata
- Paterinida
- Paterinidae
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Micromitra burgessensis
|
O’Brien and Caron 2016 | |||||||||
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Paterina zenobia
|
O’Brien and Caron 2016 | |||||||||
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Dictyonina sp.
Cooper 1942
|
O’Brien and Caron 2016 | |||||||||
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Kutorginata
- Kutorginida
- Nisusiidae
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Nisusia burgessensis
|
O’Brien and Caron 2016 | |||||||||
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Rhynchonellata
- Orthida
- Bohemiellidae
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Diraphora bellicostata
|
O’Brien and Caron 2016 | |||||||||
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unclassified
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Burgessochaeta sp.
|
O’Brien and Caron 2016 | |||||||||
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Hyolithelminthida
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Byronia annulata
|
O’Brien and Caron 2016 | |||||||||
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unclassified
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Tubulella flagellum
|
O’Brien and Caron 2016 | 299 specimens | ||||||||
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Coeloscleritophora
- Chancelloriida
- Chancelloriidae
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Chancelloria sp.
Walcott 1920
|
O’Brien and Caron 2016 | |||||||||
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Anthozoa
- Actiniaria
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Mackenzia costalis
|
O’Brien and Caron 2016 | |||||||||
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unclassified
|
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Cambrorhytium major
(Walcott 1908)
|
O’Brien and Caron 2016 | |||||||||
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Dinomischidae
|
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Dinomischus isolatus
Chen et al. 1989
|
O’Brien and Caron 2016 | |||||||||
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Hexactinellida
- Reticulosa
- Dictyospongiidae
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Protoprisma annulata n. gen., n. sp.
Rigby and Collins 2004
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Hexactinellida
- Reticulosa
- Hintzespongiidae
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Hintzespongia sp.
Rigby and Gutschick 1976
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O’Brien and Caron 2016 | |||||||||
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Hexactinellida
- Reticulosa
- Protospongiidae
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Protospongia hicksi
Hinde 1887
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Diagoniella cyathiformis
(Dawson and Hinde 1889)
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? Sanshapentella sp.
Mehl and Erdtmann 1994
|
O’Brien and Caron 2016 | |||||||||
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Heteractinida
- Octactinellida
- Eiffeliidae
|
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Eiffelia globosa
Walcott 1920
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Petaloptyon danei
Raymond 1931
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Demospongiae
- Vauxiidae
|
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Vauxia gracilenta
Walcott 1920
|
O’Brien and Caron 2016 | |||||||||
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Vauxia bellula
Walcott 1920
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Demospongiae
- Protomonaxonida
- Choiidae
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Choia carteri
Walcott 1920
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Demospongiae
- Protomonaxonida
- Wapkiidae
|
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Wapkia elongata n. sp.
Rigby and Collins 2004
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Wapkia grandis
Walcott 1920
|
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Demospongiae
- Protomonaxonida
- Hamptoniidae
|
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Hamptonia bowerbanki
Walcott 1920
|
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Demospongiae
- Protomonaxonida
- Piraniidae
|
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Pirania muricata
Walcott 1920
|
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Demospongiae
- Protomonaxonida
- Takakkawiidae
|
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Takakkawia lineata
Walcott 1920
|
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Demospongiae
- Protomonaxonida
- Leptomitidae
|
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Leptomitella incepta
Rigby and Collins 2004
|
O’Brien and Caron 2016 | |||||||||
|
Leptomitus lineatus
(Walcott 1920)
|
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Demospongiae
- Protomonaxonida
- Halichondritidae
|
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Halichondrites sp.
Walcott 1920
|
O’Brien and Caron 2016 | |||||||||
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Demospongiae
- Protomonaxonida
- Hazeliidae
|
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Crumillospongia sp.
Rigby 1986
|
O’Brien and Caron 2016 | |||||||||
|
Hazelia conferta
Walcott 1920
|
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Falospongia ramosa
Rigby and Collins 2004
|
O’Brien and Caron 2016 | |||||||||
|
Marrellomorpha
- Marrellida
|
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Marrella splendens
|
O’Brien and Caron 2016 | 5028 specimens | ||||||||
| see common names | ||||||||||
Geography
| Country: | Canada | State/province: | British Columbia |
| Coordinates: | 51.4° North, 116.4° West (view map) | ||
| Paleocoordinates: | 3.7° South, 80.0° West | ||
| Basis of coordinate: | based on nearby landmark | ||
| Geographic resolution: | small collection |
Time
| Period: | Cambrian | ||
| 10 m.y. bin: | Cambrian 3 | ||
| Key time interval: | Delamaran | Trilobite zone: | Bathyuriscus-Elrathina |
| Age range of interval: | 511 - 506.5 m.y. ago |
Stratigraphy
| Formation: | Burgess Shale | Member: | Campsite Cliff Shale | ||
| Stratigraphic resolution: | bed | ||||
| Stratigraphy comments: Pagetia bootes subzone, S7 community | |||||
Lithology and environment
| Primary lithology: | lithified "shale" |
| Environment: | deep-water indet. |
| Geology comments: Aria and Caron (2024) add "more distal palaeoenvironment from the edge of the Cathedral Escarpment compared to the nearby Walcott Quarry, but otherwise shares similar taphonomic pathways with fossils being buried by rapid mud-flow deposits." | |
Taphonomy
| Modes of preservation: | mold/impression |
| Lagerst�tten type: | conservation |
| Size of fossils: | macrofossils |
| Temporal resolution: | snapshot |
Collection methods and comments
| Collection methods: | field collection | ||
| Reason for describing collection: | taxonomic analysis | ||
| Collectors: | Royal Ontario Museum | Collection dates: | 1983 through present |
| Collection method comments: Caron and Aria (2024) note most samples were collected in 1983, 1996, 2000, 2008, and 2016 | |||
Metadata
| Also known as: | Tulip Beds | ||
| Database number: | 121959 | ||
| Authorizer: | W. Kiessling, P. Novack-Gottshall | Enterer: | U. Merkel, L. Na, P. Novack-Gottshall |
| Modifier: | P. Novack-Gottshall | Research group: | marine invertebrate |
| Created: | 2011-12-12 21:57:03 | Last modified: | 2024-07-26 16:44:23 |
| Access level: | authorizer only | Released: | 2012-12-12 21:57:03 |
| Creative Commons license: | CC BY | ||
Reference information
Primary reference:
| 38794. | J. K. Rigby and D. Collins. 2004. Sponges of the Middle Cambrian Burgess Shale and Stephen Formations, British Columbia. Royal Ontario Museum Contributions in Science 1:1-155 [W. Kiessling/U. Merkel/U. Merkel] |
Secondary references:
| 89049 | C. Aria and J. Caron. 2024. Deep origin of articulation strategies in panarthropods: evidence from a new luolishaniid lobopodian (Panarthropoda) from the Tulip Beds, Burgess Shale. Journal of Systematic Palaeontology 22(1):1-22 [P. Novack-Gottshall/P. Novack-Gottshall] | |
| 50587 | A. C. Daley and G. E. Budd. 2010. New anomalocaridid appendages from the Burgess Shale, Canada. Palaeontology 53(4):721-738 [W. Kiessling/L. Na] | |
| 63041 | L. J. O’Brien and J. B. Caron. 2012. A new stalked filter-feeder from the Middle Cambrian Burgess Shale, British Columbia, Canada. PLoS ONE 7(1):e29233-1-e29233-21 [L. Na/L. Na/P. Novack-Gottshall] | |
| 89050 | L. J. O’Brien and J. Caron. 2016. Paleocommunity analysis of the Burgess Shale Tulip Beds, Mount Stephen, British Columbia: Comparison with the Walcott Quarry and implications for community variation in the Burgess Shale. Paleobiology 42(01):27-53 [P. Novack-Gottshall/P. Novack-Gottshall] |