Aix-en-Provence (MNHN Paris Coll): Chattian, France

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Arachnida - Araneae - Salticidae
Attoides eresiformis n. gen., n. sp. Brongniart 1877
Brongniart 1877 1 specimen
In "Le Muséum", sent by M. Gaudry
Arachnida - Araneae - Gnaphosidae
Drassodes sextii n. sp. Berland 1939
Berland 1939 1 specimen
Insecta - Diptera - Mycetophilidae
Mycetophila meigeniana Heer 1856
Theobald 1937 1 specimen
recombined as Gnoriste meigeniana
Coll Museum Paris Am 32
Mycetophila confusa Theobald 1937
Theobald 1937 1 specimen
Coll Museum Paris Am 30
Mycetophila cf. morio Heer 1856
Theobald 1937 2 specimens
Coll Museum Paris Am 34-36
Mycomyia curvithoracis n. sp. Theobald 1937
Theobald 1937 1 specimen
recombined as Mycomya curvithoracis
Coll Museum Paris Am 27
Insecta - Diptera - Sciaridae
Sciara oblonga Theobald 1937
Theobald 1937 2 specimens
Coll Museum Paris Am 31, 33
Insecta - Diptera - Bibionidae
Plecia livida (Heer 1849)
Theobald 1937 8 specimens
Coll Museum Paris Am 51-56, 58, 60
Plecia brevipennis (Heer 1856)
Theobald 1937 1 specimen
synonym of Plecia livida
Coll Museum Paris Am 65
Plecia guinieri Theobald 1937
Theobald 1937 1 specimen
synonym of Plecia lygaeoides
Coll Museum Paris Am 64
Plecia splendida Theobald 1937
Theobald 1937 1 specimen
synonym of Plecia foersteri
Coll Museum Paris Am 63
Plecia bucklandi (Heer 1849)
Theobald 1937 2 specimens
Coll Museum Paris Am 50, 19
Plecia dilatativentris Theobald 1937
Theobald 1937 2 specimens
synonym of Plecia bucklandi
Coll Museum Paris Am 42, 48
Plecia elegans (Heer 1856)
Theobald 1937 5 specimens
synonym of Plecia bucklandi
Coll Museum Paris Am 43-46, 59
Plecia gracilis (Heer 1856)
Theobald 1937 1 specimen
synonym of Plecia bucklandi
Coll Museum Paris Am 57
Bibiodes provincialis n. sp. Skartveit and Nel 2017
Skartveit and Nel 2017 6 specimens
MNHN Aix 1079, 23, 62, 543, 676, 959
Bibio sp. Geoffroy 1762
Theobald 1937 1 specimen
Coll Museum Paris Am 21
Bibio aquaesextiae n. sp. Skartveit and Nel 2017
Skartveit and Nel 2017 11 specimens
MNHN Aix 152, 309, 416,421, 532, 610, 1010, 1014, 1294, 1436, 1452
Bibio spectabilis Theobald 1937
Theobald 1937 16 specimens
Coll Museum Paris Am 78; 15 in Skarveti & Nel 2017 (Aix 6, 10, 11, 52, 107, 161, 309, 845, 1090, 1299, 1307, 1377, 1446, 1447, P25)
Bibio curtisi (Heer 1856)
Theobald 1937 1 specimen
original and current combination Bibio curtisii
Coll Museum Paris Am 77
Bibio curtisii Heer 1856
Skartveit and Nel 2017 6 specimens
MNHN Aix 42, 54, 584, 873, 950, B.24431
Bibio serresi Theobald 1937
Theobald 1937 7 specimens
synonym of Bibio curtisii
Coll Museum Paris Am 67-73
Bibio romani Theobald 1937
Theobald 1937 1 specimen
synonym of Bibio curtisii
Coll Museum Paris Am 74
Bibio disjectus Theobald 1937
Theobald 1937 1 specimen
nomen dubium belonging to Bibio
Coll Museum Paris Am 75
Dilophus sp. Meigen 1803
Skartveit and Nel 2017 6 specimens
MNHN Aix 18, 31,34, 491, 797, 1079
Penthetria nervisinuata (Theobald 1937)
Skartveit and Nel 2017 19 specimens
MNHN Aix 332, 428, 645, 752, 779, 799, 884, 890, 926, 965, 974, 1013, 1061,1085, 1332, 1337, 1347, 1404, P26
Plecia cf. lugens (Oustalet 1870)
Theobald 1937 2 specimens
recombined as Penthetria lugens
Coll Museum Paris Am 39, 41
Insecta - Diptera - Bombyliidae
Exoprosopa romani Theobald 1937
Theobald 1937 1 specimen
Insecta - Diptera - Sciomyzidae
Tetanocera alireticulata n. sp. Theobald 1937
Theobald 1937 1 specimen
Coll Museum Paris Am 25
Insecta - Diptera - Syrphidae
Chilosia oligocenica n. sp. Theobald 1937
Theobald 1937 1 specimen
recombined as Cheilosia oligocenica
Coll Museum Paris Am 24
Insecta - Diptera
? Anthomyia pulcherrima Meunier 1915
Theobald 1937 2 specimens
nomen dubium belonging to Brachycera
Coll Museum Paris Am 26, 37
Insecta - Diptera - Tabanidae
Chrysops sp. Meigen 1803
Theobald 1937 1 specimen
Coll Museum Paris Am 28(29)
Insecta - Odonata - Libellulidae
Libellulidae informal larva
Theobald 1937 7 specimens
Am 80-87
Insecta - Odonata - Platycnemididae
Platycnemididae indet. Jacobson and Bianchi 1905
Nel and Papazian 1990 1 specimen
MNHN IPM-R07704
Insecta - Odonata - Coenagrionidae
Coenagrionidae informal indet. C Kirby 1890
Nel and Papazian 1990 1 specimen
MNHN IPM-R07460
Coenagrionidae informal indet. D Kirby 1890
Nel and Papazian 1990 1 specimen
MNHN IPM-R07706
Argiinae informal indet. A Tillyard 1917
Nel and Papazian 1990 1 specimen
MNHN IPM-R07449
Argiinae informal indet. B Tillyard 1917
Nel and Papazian 1990 1 specimen
MNHN IPM-R07705
Insecta - Hemiptera - Rhyparochromidae
Aphanus dilatatus n. sp. Theobald 1937
Theobald 1937 10 specimens
Coll Museum Paris Am 132
Aphanus murchisonae (Heer 1853)
Theobald 1937 27 specimens
Coll Museum Paris Am 122-126, 100-121
Aphanus pulchellus (Heer 1853)
Theobald 1937 5 specimens
Coll Museum Paris Am 143-146, 138
Diniella fasciata (Heer 1853)
Theobald 1937 5 specimens
Coll Museum Paris Am 147-151
Insecta - Hemiptera - Berytidae
? Neides oligocenicus n. sp. Nel 1992
Nel 1992 4 specimens
MNHN IPM-R.10372, 10373, 10374, 10375 (4 measurements)
Insecta - Hemiptera - Heterogastridae
Heterogaster antiquus Heer 1853
Theobald 1937 1 specimen
Coll Museum Paris Am 127
Heterogaster pumilio Heer 1853
Theobald 1937 3 specimens
Coll Museum Paris Am 140-142
Insecta - Hemiptera - Cydnidae
Cydnus meunieri Theobald 1937
Theobald 1937 1 specimen
Coll Museum Paris Am 139
Insecta - Hemiptera - Hydrometridae
Protobacillometra oligocenica n. gen., n. sp. Nel and Paicheler 1993
Nel and Paicheler 1993 1 specimen
MNHN LP-R.07893 (1 measurement)
Insecta - Orthoptera - Gryllidae
Nemobius sp. Serville 1839
Scudder 1890 2 specimens
in the collection of Oustalet
    = Gryllidae indet. Laicharting 1781
Zeuner 1939
Grylloides
Gryllidae indet. 6 and 7 in Zeuner; form only mentioned by Scudder
Insecta - Dermaptera - Forficulidae
Forficula informal sp. A Linnaeus 1758
Nel et al. 1994 1 specimen
MNHN-LP-R.10401
Insecta - Blattodea
Paleotermopsis oligocenicus n. gen., n. sp. Nel and Paicheler 1993
Nel and Paicheler 1993 1 specimen
MNHN IPM-R.54190 (2 measurements)
Ulmeriella aquisextana n. sp. Nel 1986
Nel 1986 2 specimens
MNHN IPM-R.07443 (Nel no. 1358); IPM-R.54242 in Nel & Paicheler 1993 (3 measurements)
Insecta - Blattodea - Mastotermitidae
Blattotermes massiliensis n. sp. Nel 1986
Nel 1986 1 specimen
MNHN IPM-R.07442 (Nel no. 1380) (1 measurement)
Mastotermes sp. Froggatt 1897
Nel and Paicheler 1993 2 specimens
MNHN IPM-R.54195, 54324 (2 measurements)
Mastotermes gallica n. sp. Nel 1986
Nel 1986 1 specimen
MNHN IPM-R.07441 (Nel no. 1) (2 measurements)
Insecta - Hymenoptera - Braconidae
Diospilus verus n. sp. Theobald 1937
Theobald 1937 1 specimen
recombined as Taphaeus verus
Coll Museum Paris Am 23
Insecta - Hymenoptera - Ichneumonidae
Promethes tilloyi n. sp. Theobald 1937
Theobald 1937 1 specimen
recombined as Zagryphus tilloyi
Coll Museum Paris Am 8 (MNHN.F.24392) (2 measurements)
? Demophorus fumipennis n. sp. Theobald 1937
Theobald 1937 1 specimen
Coll Museum Paris Am 14
Ophion annulatus n. sp. Theobald 1937
Theobald 1937 1 specimen
Coll Museum Paris Am 13
? Nemeritis longicornis n. sp. Theobald 1937
Theobald 1937 1 specimen
recombined as Dimophora longicornis
Coll Museum Paris Am 12 (MNHN.F.B24398) (2 measurements)
Exetastes postornata n. sp. Theobald 1937
Theobald 1937 1 specimen
Coll Museum Paris Am 16
Pimpla antiqua n. sp. Saussure 1852
Saussure 1852 1 specimen
recombined as Lithoserix antiquus
Coll Museum Paris 21 (MNHN.F.B24391); Theobald said that this specimen was different from Saussure's holotype, but it is likely the same (Spasojevic et al., 2022) (2 measurements)
? Pimpla saussurei (Heer 1856)
Theobald 1937 1 specimen
Coll Museum Paris 17
Insecta - Hymenoptera - Pompilidae
Pompilus coquandi n. sp. Theobald 1937
Theobald 1937 1 specimen
recombined as Pompilinites coquandi
Coll Museum Paris Am 1
Pompilus fasciatus n. sp. Theobald 1937
Theobald 1937 1 specimen
recombined as Pompilites fasciatus
Coll Museum Paris Am 11
Insecta - Hymenoptera - Thynnidae
Methoca sp. (Latreille 1804)
Theobald 1937 1 specimen
original and current combination Methocha
Coll Museum Paris Am 3
Insecta - Hymenoptera - Formicidae
Lasius epicentrus n. sp. Theobald 1937
Theobald 1937 1 specimen
Coll Museum Paris Am 7
Formica minutula Heer 1849
Theobald 1937 1 specimen
synonym of Lasius occultatus
Coll Museum Paris Am 15
Formica sp. Linnaeus 1758
Theobald 1937 1 specimen
Coll Museum Paris Am 6
Formica latinodosa n. sp. Theobald 1937
Theobald 1937 1 specimen
Coll Museum Paris Am 9
Formica sepulta n. sp. Theobald 1937
Theobald 1937 1 specimen
Coll Museum Paris Am 5
Camponotus longiventris n. sp. Theobald 1937
Theobald 1937 2 specimens
Coll Museum Paris Am 2, 18
Camponotus saussurei n. sp. Theobald 1937
Theobald 1937 1 specimen
replaced by Camponotus theobaldi
Coll Museum Paris Am 4
Dolichoderus coquandi n. sp. Theobald 1937
Theobald 1937 1 specimen
Coll Museum Paris Am 22
Brachyponera dubia Theobald 1937
Theobald 1937 1 specimen
recombined as Pachycondyla dubia
Coll Museum Paris Am 20
Aphaenogaster maculata Theobald 1937
Theobald 1937 1 specimen
recombined as Paraphaenogaster maculata
Coll Museum Paris Am 10
Formica oculata Heer 1849
Theobald 1937 1 specimen
nomen dubium belonging to Formicariae
Coll Museum Paris Am 19
Insecta - Hymenoptera - Apidae
Apis informal sp. B Linnaeus 1758
Nel et al. 1999 1 specimen
MNHN-LP-R.10429 (2 measurements)
Apis informal sp. C Linnaeus 1758
Nel et al. 1999 1 specimen
MNHN-LP-R.10426 (2 measurements)
Apis aquisextana n. sp. Nel et al. 1996
Arillo et al. 1996 3 specimens
MNHN-LP-R.08381, 08382, 10421 (6 measurements)
    = Apis aquisextusensis n. sp. Nel et al. 1999
Nel et al. 1999
synonym of Apis (Synapis) henshawi
Insecta - Neuroptera - Chrysopidae
Nothochrysa oligocenica n. sp. Ngô-Muller et al. 2019
Ngô-Muller et al. 2019 1 specimen
MNHN.F.A770996 (1 measurement)
Insecta - Coleoptera - Carabidae
Polystichus hopei n. sp. Oustalet 1874
Oustalet 1874 1 specimen
(1 measurement)
Insecta - Coleoptera - Geotrupidae
Geotrupes atavus n. sp. Oustalet 1874
Oustalet 1874 1 specimen
(2 measurements)
Insecta - Coleoptera - Staphylinidae
Erinnys deleta n. sp. Oustalet 1874
Oustalet 1874 1 specimen
recombined as Lithoplanes deleta
(2 measurements)
Staphylinus aquisextanus n. sp. Oustalet 1874
Oustalet 1874 1 specimen
(1 measurement)
Staphylinus prodromus Oustalet 1874
Oustalet 1874 2 specimens
Staphylinus calvus Oustalet 1874
Oustalet 1874 1 specimen
Staphylinus priscus Oustalet 1874
Oustalet 1874 1 specimen
Staphylinus (Ocypus) provincialis n. sp. Oustalet 1874
Oustalet 1874 1 specimen
recombined as Ocypus provincialis
(1 measurement)
Stenus gypsi n. sp. Oustalet 1874
Oustalet 1874 1 specimen
(2 measurements)
Insecta - Coleoptera - Curculionidae
Hipporhinus heeri (Germar 1849)
Theobald 1937 32 specimens
nomen dubium belonging to Entimini
Hipporhinus heerii Germar 1849
Oustalet 1874 16 specimens
nomen dubium belonging to Entimini
Hipporhinus reynesii Oustalet 1874
Oustalet 1874 5 specimens
synonym of Rhytidoderes sexsulcatus
(2 measurements)
Sitones margarum (Germar 1849)
Oustalet 1874 4 specimens
recombined as Protainophthalmus margarus
(2 measurements)
Brachyderes aquisextanus Oustalet 1874
Oustalet 1874 1 specimen
synonym of Protainophthalmus asperulus
Brachyderes longipes Oustalet 1874
Oustalet 1874 4 specimens
synonym of Protainophthalmus asperulus
Cleonus marcelli Oustalet 1874
Oustalet 1874 1 specimen
synonym of Protainophthalmus asperulus
Cleonus inflexus Oustalet 1874
Oustalet 1874 1 specimen
nomen dubium belonging to Curculionites
Insecta - Coleoptera - Chrysomelidae
Chrysomela matheroni Oustalet 1874
Oustalet 1874 1 specimen
Chrysomela matrona n. sp. Oustalet 1874
Oustalet 1874 2 specimens
synonym of Plagiodera lyelliana
(1 measurement)
Gonioctena curtisii n. sp. Oustalet 1874
Oustalet 1874 1 specimen
(1 measurement)
Cassida blancheti Heer 1856
Oustalet 1874 3 specimens
(2 measurements)
Insecta - Lepidoptera - Pterophoridae
Pterophorus oligocenicus n. sp. Bigot et al. 1986
Bigot et al. 1986 1 specimen
recombined as Merrifieldia oligocenicus
Oustalet collection (2 measurements)
see common names

Geography
Country:France State/province:Bouches-Du-Rhone
Coordinates: 43.5° North, 5.5° East (view map)
Paleocoordinates:43.3° North, 2.8° East
Basis of coordinate:estimated from map
Geographic resolution:local area
Time
Period:Paleogene Epoch:Oligocene
Stage:Chattian 10 m.y. bin:Cenozoic 4
Key time interval:Chattian
Age range of interval:27.82000 - 23.03000 m.y. ago
Stratigraphy
Formation:Niveau du gypse d'Aix
Stratigraphy comments: "The ‘insect formation’ is 2.5 m thick, divided in two parts that correspond to two very different environmental conditions. The first (lower) episode is remarkable by its lateral monotony of facies on a distance of 6 km at least, in the outcrops observed on the northwest of Aix-en-Provence. This lower episode is distinguished from the rest of the ‘insect formation’ and from the whole formation of Aix-en-Provence by the presence of a very numerous thin limestone laminae. The second (upper) episode is remarkable by the presence of many silici- fied laminae. The fauna in this level is also very well preserved"….This collection consists of unspecified layers
Lithology and environment
Primary lithology:planar lamination lithified "limestone"
Lithology description: The deposit presents an alternation of limestone and marls with gypsum.
Environment:lacustrine - large Tectonic setting:rift
Geology comments: This lacustrine formation was deposited in a very large but shallow-water lake, with variable salinity and near the marine coast (Nury 1990, Nury & Thomassin 1994). Irregular basins formed during extension of the European platform in the Oligocene resulting from the formation of the West-European rift system.
Taphonomy
Modes of preservation:mold/impression,adpression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Museum repositories:MNHN
Metadata
Database number:122501
Authorizer:M. Clapham Enterer:J. Karr, M. Clapham
Modifier:M. Clapham Research group:paleoentomology
Created:2011-12-31 10:57:54 Last modified:2014-08-24 11:05:55
Access level:the public Released:2011-12-31 10:57:54
Creative Commons license:CC BY
Reference information

Primary reference:
39047. F. Meunier. 1915. Nouvelles recherches sur quelques insectes des Plâtrières d'Aix en Provence. Verhandelingen der Koninklijke Akademie van Wetenschappen te Amsterdam 18(5):1-17 [M. Clapham/J. Karr/M. Clapham]

Secondary references:

50848 A. Arillo, A. Nel, and V. M. Ortuño. 1996. Two fossil bees from Oligocene of Izarra (Alava, Spain) (Hymenoptera, Apoidea). Bulletin de la Société Entomologique de France 101:59-64 [M. Clapham/M. Clapham/M. Clapham]
47456 L. Berland. 1939. Description de quelques arignées fossils. Revue Française d’Entomologie 6:1-9 [M. Clapham/M. Clapham]
45423 L. Bigot, A. Nel, and J. Nel. 1986. Description de la première espèce fossile connue de Ptérophore (Lepidoptera Pterophoridae). Alexanor 14:283-288 [M. Clapham/M. Clapham/M. Clapham]
47457 C. Brongniart. 1877. Note sue une aranéide fossile des terrains tertiaires d'Aix (Provence). Annales de la Société Entomologique de France, Cinquième Série 7:221-224 [M. Clapham/M. Clapham/M. Clapham]
51478 A. Nel. 1986. Sur trois espèces nouvelles de Termites fossiles du stampien d'Aix-en-Provence (Bouches-du-Rhône) (Dictyoptera, Hodotermitidae, Mastotermitidae). L'Entomologiste 42:271-278 [M. Clapham/M. Clapham]
51672 A. Nel. 1992. Essai de révision des Berytidae fossiles (Heteroptera, Pentatomorpha). Bulletin du Muséum National d'Histoire Naturelle. Section C, Sciences de la Terre, Paléontologie, Géologie, Minérologie 14:275-287 [M. Clapham/M. Clapham]
51667 A. Nel, V. Albouy, C. Caussanel and C. Jamet. 1994. Réflexion paléo-entomologique sur la systématique des Dermaptères. Quatre nouveaux forficules fossiles de l'Oligocène de Provence (France). Bulletin de la Société Entomologique de France 99:253-266 [M. Clapham/M. Clapham]
44434 A. Nel, X. Martínez-Delclòs, A. Arillo and E. Peñalver. 1999. A review of the European fossil species of the bee Apis. Palaeontology 42:243-285 [M. Clapham/M. Clapham]
51528 A. Nel and J. C. Paicheler. 1993. Les Isoptera fossiles. État actuel des connaissances, implications paléoécologiques et paléoclimatologiques (Insecta, Dictyoptera). Cahiers de Paléontologie 1993:102-179 [M. Clapham/M. Clapham]
55973 A. Nel and J. C. Paicheler. 1993. Les Heteroptera aquatiques fossiles, état actuel des connaissances (Heteroptera: Nepomorpha et Gerromorpha) (suite et fin). Entomologica Gallica 4:79-89 [M. Clapham/M. Clapham]
68069 A. Nel and M. Papazian. 1990. Les Coenagrionoidea fossiles de l’Oligocène du sud-est de la France (Zygoptera: Platycnemididae, Coenagrionidae). Odonatologica 19:251-262 [M. Clapham/M. Clapham]
82517 V. Ngô-Muller, K. Blot, and A. Nel. 2019. A new species of Nothochrysa from the latest Oligocene of France (Insecta: Neuroptera: Chrysopidae). Palaeoentomology 2:111-114 [M. Clapham/M. Clapham]
49085 E. Oustalet. 1874. Recherches sur les insectes fossiles des terrains Tertiaires de la France, deuxième partie, insectes fossiles d'Aix en Provence. Annales des Sciences Géologiques, Paris 5(2):1-347 [M. Clapham/M. Clapham]
49945 H. Saussure. 1852. Note sur un nouvel insecte hyménoptère fossile. Revue et Magasin de Zoologie, Deuxième Série 4:579-581 [M. Clapham/M. Clapham]
5136ETE 848S. H. Scudder. 1890. The Tertiary Insects of North America. Report of the United States Geological Survey of the Territories XIII:1-734 [C. Labandeira/C. Labandeira/L. Edwards]
61455 J. Skartveit and A. Nel. 2017. Revision of fossil Bibionidae (Insecta: Diptera) from French Oligocene deposits. Zootaxa 4225:1-83 [M. Clapham/M. Clapham]
38999 N. Theobald. 1937. Les insectes fossiles des terrains oligocènes de France. Bulletin Mensuel (Mémoires) de la Société des Sciences de Nancy 1:1-473 [M. Clapham/J. Karr]
39932 F. E. Zeuner. 1939. Fossil Orthoptera Ensifera 1-321 [M. Clapham/M. Clapham/M. Clapham]