Aix-en-Provence (Coll. Nel): Chattian, France
collected by Andre Nel

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Odonata - Calopterygidae
? Umma sp. Kirby 1896
Nel and Paicheler 1993 1 specimen
MNHN IPM-R.07914
Insecta - Odonata - Coenagrionidae
Coenagrionidae indet. Kirby 1890
Nel et al. 1997 1 specimen
MNHN-LP-R.55176
Insecta - Odonata - Lestidae
Lestes informal sp. 1 Leach 1815
Nel and Paicheler 1994 1 specimen
MNHN-LP-R 54241 Coll. Nel (1 measurement)
Lestes informal sp. 2 Leach 1815
Nel and Paicheler 1994 1 specimen
MNHN-LP-R 54371 Coll. Nel (1 measurement)
Lestes aquisextana n. sp. Nel 1985
Nel 1985 1 specimen
MNHN-LP-R 07447 (1 measurement)
Insecta - Odonata - Lindeniidae
Lindeniidae indet. Jacobson and Bianchi 1905
Nel et al. 1997 1 specimen
MNHN-LP-R.55186
Insecta - Odonata - Libellulidae
Sympetrum bigoti n. sp. Nel and Papazian 1985
Nel and Papazian 1985 15 specimens
Nel 1 A2, 183A, 186A, 188A (MNHN-LP-R.07452, 07455, 07454, 07453, 07915, 07916-07924) (9 measurements)
Palaeotramea aquisextana n. gen., n. sp. Nel and Papazian 1985
Nel and Papazian 1985 31 specimens
isolate fore and hindwing (31 specimens given by Nel & Paicheler, 1993) (19 measurements)
Caussanelia papaziani n. gen., n. sp. Nel et al. 1997
Nel et al. 1997 1 specimen
MNHN-LP-R.55179 (1 measurement)
Insecta - Odonata - Aeshnidae
Aeshna oligocenica n. sp. Nel 1994
1 specimen
MNHN-LP-R.07895 (1 measurement)
Insecta - Blattodea - Kalotermitidae
Oligokalotermes fischeri n. gen., n. sp. Nel 1987
Nel 1987 1 specimen
Coll Nel 202 (2 measurements)
Insecta - Coleoptera - Curculionidae
Hylesinus (Hylesinus) neli n. sp. Petrov and Zherikhin 2004
Petrov and Zherikhin 2004 1 specimen
recombined as Hylesinus neli
MNHN-LP-R.55233 (1 measurement)
Insecta - Coleoptera - Hydrophilidae
Limnoxenus olenus n. sp. Fikáček et al. 2010
Fikáček et al. 2010 2 specimens
MNHN A32835, A32836 (4 measurements)
Insecta - Coleoptera - Gyrinidae
Gyrinus aquisextanea n. sp. Nel 1989
Nel 1989 1 individual
MNHN-LP-R 07724 (2 measurements)
Insecta - Coleoptera - Cupedidae
Cainocups aixensis n. gen., n. sp. Kirejtshuk et al. 2016
Kirejtshuk et al. 2016 1 specimen
MNHN.F.A.51230 ("Aix 18")
see common names

Geography
Country:France State/province:Bouches-Du-Rhone
Coordinates: 43.5° North, 5.5° East (view map)
Paleocoordinates:43.3° North, 2.8° East
Basis of coordinate:estimated from map
Geographic resolution:local area
Time
Period:Paleogene Epoch:Oligocene
Stage:Chattian 10 m.y. bin:Cenozoic 4
Key time interval:Chattian
Age range of interval:27.82000 - 23.03000 m.y. ago
Stratigraphy
Formation:Niveau du gypse d'Aix
Stratigraphy comments: "The ‘insect formation’ is 2.5 m thick, divided in two parts that correspond to two very different environmental conditions. The first (lower) episode is remarkable by its lateral monotony of facies on a distance of 6 km at least, in the outcrops observed on the northwest of Aix-en-Provence. This lower episode is distinguished from the rest of the ‘insect formation’ and from the whole formation of Aix-en-Provence by the presence of a very numerous thin limestone laminae. The second (upper) episode is remarkable by the presence of many silici- fied laminae. The fauna in this level is also very well preserved"….This collection consists of unspecified layers
Lithology and environment
Primary lithology:planar lamination lithified "limestone"
Lithology description: The deposit presents an alternation of limestone and marls with gypsum.
Environment:lacustrine - large Tectonic setting:rift
Geology comments: This lacustrine formation was deposited in a very large but shallow-water lake, with variable salinity and near the marine coast (Nury 1990, Nury & Thomassin 1994). Irregular basins formed during extension of the European platform in the Oligocene resulting from the formation of the West-European rift system.
Taphonomy
Modes of preservation:mold/impression,adpression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Museum repositories:MNHN
Collectors:Andre Nel
Taxonomic list comments:"This formation has yielded a diverse flora and fauna, with finely preserved fossils. The flora is represented by numerous plant remains (pollen, roots, wood, stems, leaves, seeds and flowers). The fauna of vertebrates is represented by abundant fishes (GAUDANT 1978), and some frogs, birds, and bats with preserved skin outlines. Some invertebrates are perfectly preserved, including jellyfish and shells of aquatic molluscs with original colors and ornamentations (NEL et al. 1987). Aquatic and terrestrial arthropods (insects and arachnids) are very diverse and frequently well preserved….
In spite of this, aquatic insects remain comparatively rare among the 30,000 fossil insects one of us (A.N.) collected at the locality."
Metadata
Database number:127415
Authorizer:M. Clapham Enterer:J. Karr, M. Clapham, L. Edwards
Modifier:M. Clapham Research group:paleoentomology
Created:2012-05-13 08:30:12 Last modified:2014-08-24 11:08:36
Access level:the public Released:2013-05-13 08:30:12
Creative Commons license:CC BY
Reference information

Primary reference:
38446. A. Nel, X. Martínez-Delclòs, F. Escuillé and P. Brisac. 1994. Les Aeshnidae fossiles: Etat actuel des connaissances (Odonata, Anisoptera). Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 194(2/3):143-186 [M. Clapham/J. Karr/M. Clapham]

Secondary references:

45304 M. Fikáček, J. Prokop, and A. Nel. 2010. Fossil water scavenger beetles of the subtribe Hydrobiusina (Coleoptera: Hydrophilidae) from the Late Oligocene locality of Aix-en-Provence (France). Acta Entomologica Musei Nationalis Pragae 50:445-458 [M. Clapham/M. Clapham/M. Clapham]
63371 A. G. Kirejtshuk, A. Nel, and P. A. Kirejtshuk. 2016. Taxonomy of the reticulate beetles of the subfamily Cupedinae (Coleoptera: Archostemata), with a review of the historical development. Invertebrate Zoology 13(2):61-190 [M. Clapham/M. Clapham]
51554 A. Nel. 1985. Sur la présence d’un Lestes Leach,1815 fossile de la lignée de Lestes regina Théobald dans les calcaires stampiens d’Aix-en-Provence (Odon. Lestidae). Entomologica Gallica 1:317-319 [M. Clapham/M. Clapham]
47079 A. Nel. 1987. Deux nouveaux termites du Tertiaire du sud-est de la France (Dictyoptera, Hodotermitidae, Kalotermitidae). Revue Française d'Entomologie (N.S.) 9(1):17-20 [M. Clapham/J. Karr]
50847 A. Nel, A. Arillo, and V. M. Ortuño. 1997. New western Palaearctic Cenozoic Odonata (Zygoptera and Anisoptera). Bulletin de la Société Entomologique de France 102:265-270 [M. Clapham/M. Clapham]
41732 A. Nel, X. Martínez-Delclòs, F. Papier and J. Oudard. 1997. New Tertiary Fossil Odonata from France (Sieblosiidae, Lestidae, Coenagrionidae, Megapodagrionidae, Libellulidae). Deutsche Entomologische Zeitschrift 44(2):231-258 [M. Clapham/L. Edwards/M. Clapham]
42270 A. Nel and J. C. Paicheler. 1993. Les Odonata fossiles: état actuel des connaissances. Huitième partie: Les Calopterygoidea fossiles (Odonata, Zygoptera). Bulletin de la Societe Entomologique de France 97(4):381-396 [M. Clapham/J. Karr]
51552 A. Nel and M. Papazian. 1985. Description d'une nouvelle espèce d'odonate fossile du Stampien d'Aix-en-Provence (Odon., Libellulidae). Entomologica Gallica 1:351-355 [M. Clapham/M. Clapham/M. Clapham]
46752 A. Nel and M. Papazian. 1985. Une nouvelle espèce d'Odonata fossile du stampien a'Aix-en Provence (B.d.R.) (Odonata Libellulidae). L'Entomologiste 41(2):55-59 [M. Clapham/J. Karr/J. Karr]
49497 A. V. Petrov and V. V. Zherikhin. 2004. A new species of bark beetles of the genus Hylesinus (Insecta: Coleoptera: Scolytidae) from the Oligocene of France. Paleontological Journal 38:187-189 [M. Clapham/M. Clapham]